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Pentatominae
Pentatoma rufipes
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hemiptera
Suborder:	Heteroptera
Family:	Pentatomidae
Subfamily:	Pentatominae Leach, 1815

Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera. Species in this subfamily are phytophages and several of them are considered agricultural pests. Some invasive pentatomines such as Halyomorpha halys and Bagrada hilaris have been considered household pests. Higher systematics of the group have been revised by Rider et al.

Tribes and genera

Tribes within this subfamily include:

• Aelini Douglas & Scott, 1865

1. Aelia Fabricius, 1803
2. Aeliopsis Bergevin, 1931
3. Neottiglossa Kirby, 1837

• Aeptini Stål, 1871

1. Aeliosoma Baehr, 1985
2. Aeptus Dallas, 1851
3. Caridillus Bergroth, 1909
4. Eribotes (bug) Stål, 1867
5. Hillieria Distant, 1910
6. Mariomella Faúndez & Rider, 2019
7. Menestheus (bug) Stål, 1867
8. Paramenestheus Breddin, 1900
9. Pseudaelia Distant, 1910

• Aeschrocorini Distant, 1902

1. Aeschrocoris Bergroth, 1887
2. Aeschrus Spinola, 1850
3. Chraesus Cachan, 1952
4. Geomorpha (bug) Bergroth, 1893
5. Risbecella Schouteden, 1942
6. Scylax (bug) Distant, 1887
7. Tyoma Spinola, 1850
8. Tyomana Miller, 1952

• Agaeini Cachan, 1952

1. Agaeus Dallas, 1851

• Agonoscelidini Atkinson, 1888

1. Agonoscelis Spinola, 1837

• Amyntorini Distant, 1902

1. Ambohicorypha Cachan, 1952
2. Belopis Distant, 1879
3. Bolaca Walker, 1867
4. Halyabbas Distant, 1900
5. Madecorypha Cachan, 1952
6. Sennertus Distant, 1900

• Antestiini Distant, 1902 - selected genera:
  • Antestia Stål, 1865
  • Antestiopsis Leston, 1952
  • Dryadocoris Kirkaldy, 1909
  • Plautia Stål, 1864
• Aulacetrini Mulsant & Rey, 1866

1. Holcogaster Fieber, 1860

• Axiagastini Atkinson, 1888

1. Acesines Stål, 1876
2. Axiagastus Dallas, 1851
3. Indrapura (bug) China, 1928
4. Oncotropis Stål, 1871

• Bathycoeliini Atkinson, 1888

1. Bathycoelia Amyot & Serville, 1843

• Cappaeini Atkinson, 1888

1. Adelolcus Bergroth, 1893
2. Benia (bug) Schouteden, 1916
3. Boerias Kirkaldy, 1909
4. Cappaea Ellenrieder, 1862
5. Caura (bug) Stål, 1865
6. Cauromorpha Jeannel, 1913
7. Erlangerella Schouteden, 1905
8. Halycorypha Jeannel, 1913
9. Halydicoris Jeannel, 1913
10. Halyoides Cachan, 1952
11. Halyomorpha Mayr, 1864
12. Homalogonia Jakovlev, 1876
13. Hymenomaga Karsch, 1892
14. Lerida (bug) Karsch, 1894
15. Leridella Jeannel, 1913
16. Lokaia Linnavuori, 1975
17. Mabusana Distant, 1912
18. Massocephalus Dallas, 1851
19. Paralerida Linnavuori, 1982
20. Prytanicoris Gross, 1978
21. Tolumnia Stål, 1867
22. Tripanda Berg, 1899
23. Tropicorypha Mayr, 1864
24. Veterna Stål, 1865

• Carpocorini Mulsant & Rey, 1866 selected genera:
  • Antheminia Mulsant & Rey, 1866
  • Carpocoris Kolenati, 1846
  • Chlorochroa Stål, 1872
  • Dolycoris Mulsant & Rey, 1866
  • Graphorn Faúndez, Rider, & Carvajal, 2017
  • Peribalus Mulsant & Rey, 1866
  • Rubiconia Dohrn, 1860
  • Thestral Faúndez & Rider, 2014
• Catacanthini Atkinson, 1888

1. Arocera Spinola, 1837
2. Boea (bug) Walker, 1867
3. Catacanthus Spinola, 1837
4. Chalcocoris Dallas, 1851
5. Rhyssocephala Rider, 1992
6. Runibia Stål, 1861
7. Vitruvius (bug) Distant, 1901
8. Vulsirea Spinola, 1837

• Caystrini Ahmad & Afzal, 1979

1. Caystrus Stål, 1861
2. Cephaloplatus White, 1842
3. Heissocoris Ahmad & Afzal, 1979
4. Hippotiscus Bergroth, 1906
5. Kyrtalus Van Duzee, 1929
6. Lyalocoris Zahid, 2006
7. Mcdonaldiana Zahid, 2006
8. Neodius (Stål, 1867)
9. Neoparodius Zahid, 2006
10. Neoriazocoris Zahid, 2006
11. Paramecocoris Stål, 1853
12. Parodius (bug) Distant, 1900
13. Praetextatus (bug) Distant, 1901
14. Riazocoris Ahmad & Afzal, 1979
15. Valescus Distant, 1901

• Chlorocorini Rider, Greve, Schwertner & Grazia, 2018

1. Arvelius Spinola, 1837
2. Chlorocoris Spinola, 1837
3. Chloropepla Stål, 1867
4. Eludocoris Thomas, 1992
5. Fecelia Stål, 1872
6. Mayrinia Horváth, 1925
7. Rhyncholepta Bergroth, 1911

• Coquereliini Cachan, 1952

1. Cleoqueria Cachan, 1952
2. Coquerelia Signoret, 1861
3. Coquerelidea Reuter, 1887
4. Neocoquerelidea Cachan, 1952

• Degonetini Azim & Shafee, 1984

1. Degonetus Distant, 1902

• Diemeniini Kirkaldy, 1909 - selected genera:
  • Commius (bug) Stål, 1876
  • Diemenia (bug) Spinola, 1850
• Diplostirini Distant, 1902

1. Ambiorix (bug) Stål, 1876
2. Diplostira Dallas, 1851

• Diploxyini Atkinson, 1888

1. Acoloba Spinola, 1850
2. Aeladria Cachan, 1952
3. Diploxys Amyot & Serville, 1843

• Eurysaspidini Atkinson, 1888

1. Eurysaspis Signoret, 1851
2. Flaminia (bug) Stål, 1865

• Eysarcorini Mulsant & Rey, 1866

1. Adria (bug) Stål, 1876
2. Aspavia Stål, 1865
3. Cachanocoris Rider, 1998
4. Carbula Stål, 1864
5. Corisseura Cachan, 1952
6. Cratonotus Distant, 1879
7. Durmia Stål, 1865
8. Dymantiscus Hsiao, 1981
9. Empiesta Linnavuori, 1982
10. Eysarcoris Hahn, 1834
11. Hermolaus Distant, 1902
12. Mulungua Linnavuori, 1982
13. Pseudolerida Schouteden, 1958
14. Sepontia Stål, 1865
15. Sepontiella Miyamoto, 1990
16. Spermatodes Bergroth, 1914
17. Stagonomus Gorski, 1852

• Halyini Amyot & Serville, 1843 - selected genera:
  • Brochymena Amyot & Serville, 1843
• Hoplistoderini Atkinson, 1888

1. Alcimocoris Bergroth, 1891
2. Glottaspis Bergroth, 1918
3. Hoplistodera Westwood, 1837
4. Paracritheus Bergroth, 1891
5. Paralcimocoris Zhang & Lin, 1988
6. Stachyomia Stål, 1871
7. Taurodes Dallas, 1851

• Lestonocorini Ahmad & Mohammad, 1980

1. Dardjilingia Yang, 1936
2. Gynenica Dallas, 1851
3. Lestonocoris Ahmad & Mohammad, 1980
4. Neogynenica Yang, 1935
5. Umgababa (bug) Leston, 1953

• Mecideini Distant, 1902

1. Mecidea Dallas, 1851

• Memmiini Cachan, 1952

1. Memmia Stål, 1865

• Menidini Atkinson, 1888

1. Actuarius (bug) Distant, 1901
2. Aegaleus (bug) Stål, 1876
3. Amphimachus (bug) Stål, 1867
4. Anchesmus Stål, 1876
5. Apines Dallas, 1851
6. Aspideurus Signoret, 1880
7. Banya (bug) Schouteden, 1916
8. Brachycoris Stål, 1871
9. Carvalhocoris Leston, 1953
10. Cresphontes (bug) Stål, 1867
11. Dabessus Distant, 1902
12. Decellella Schouteden, 1964
13. Desertomenida Kiritshenko, 1914
14. Dunnius Distant, 1902
15. Elanela Rolston, 1980
16. Eurymenida Bergroth, 1917
17. Gwea Schouteden, 1958
18. Keleacoris Rider & Rolston, 1995
19. Lathraedoeus Breddin, 1900
20. Menida Motschulsky, 1861
21. Neostrachia Saunders, 1877
22. Phavorinus Distant, 1902
23. Rio (bug) Kirkaldy, 1909
24. Saceseurus Breddin, 1900
25. Sciomenida Gross, 1976
26. Solenocoris Oshanin, 1922
27. Udonga Distant, 1921

• Myrocheini Stål, 1871

1. Aednulus Breddin, 1901
2. Aednus Dallas, 1851
3. Arniscus Distant, 1899
4. Delegorguella Spinola, 1850
5. Dictyotus Dallas, 1851
6. Dictytotus Dallas, 1851
7. Dissocolpus Bergroth, 1906
8. Dollingiana Ahmad & Kamaluddin, 1986
9. Dymantis Stål, 1861
10. Ennius (bug) Stål, 1861
11. Erachtheus Stål, 1861
12. Humria Linnavuori, 1975
13. Jitka (bug) Kment, 2015
14. Laprius Stål, 1861
15. Lodosia Ahmad & Önder, 1996
16. Munshiana Ahmad & Kamaluddin, 1987
17. Myrochea Amyot & Serville, 1843
18. Neococalus Bergroth, 1891
19. Neodorpius Ahmad & Afzal, 1989
20. Paradictyotus Gross, 1975
21. Pretorius (bug) Distant, 1898
22. Stysicoris Ahmad & Kamaluddin, 1985
23. Tholosanus Distant, 1899
24. Utheria Gross, 1975

• Nealeriini Cachan, 1952

1. Nealeria Bergroth, 1893
2. Paraleria Reuter, 1887

• Nezarini Atkinson, 1888 - selected genera:
  • Nezara Amyot et Serville, 1843
  • Palomena Mulsant et Rey, 1866
• Opsitomini Cachan, 1952

1. Opsitoma Cachan, 1952

• Pentamyrmexini Rider & Brailovsky, 2014 (monotypic, Thailand)
  • Pentamyrmex Rider & Brailovsky, 2014
• Pentatomini Leach, 1815 - selected genera:
  • Pentatoma Olivier, 1789
• Phricodini Cachan, 1952

1. Phricodus Spinola, 1839

• Piezodorini Atkinson, 1888

1. Anaximenes (bug) Stål, 1876
2. Chaubattiana Distant, 1912
3. Pausias (bug) Jakovlev, 1905
4. Piezodorus Fieber, 1860

• Procleticini Pennington, 1920

1. Aleixus McDonald, 1981
2. Brepholoxa Van Duzee, 1904
3. Dendrocoris Bergroth, 1891
4. Lobepomis Berg, 1891
5. Neoderoploa Pennington, 1922
6. Odmalea Bergroth, 1914
7. Parodmalea Rider, 1994
8. Procleticus Berg, 1891
9. Pseudobebaeus Distant, 1911
10. Terania Pirán, 1963
11. Thoreyella Spinola, 1850

• Rhynchocorini Stål, 1871

1. Araducta Walker, 1867
2. Avicenna (bug) Distant, 1901
3. Biprorulus Breddin, 1900
4. Cuspicona Dallas, 1851
5. Diaphyta Bergroth, 1891
6. Everardia (bug) Gross, 1975
7. Hoffmanseggiella Spinola, 1850
8. Koogobatha Distant, 1910
9. Morna (bug) Stål, 1867
10. Mygoodano Distant, 1910
11. Ocirrhoe Stål, 1876
12. Parocirrhoe Gross, 1975
13. Pegala Stål, 1867
14. Petalaspis Bergroth, 1916
15. Pugione Stål, 1867
16. Rhynchocoris Westwood, 1837
17. Utana Distant, 1900
18. Vitellus Stål, 1865

• Rolstoniellini Rider, 1997

1. Agathocles (bug) Stål, 1876
2. Amasenus Stål, 1863
3. Amatembuna Distant, 1910
4. Critheus Stål, 1867
5. Exithemus Distant, 1902
6. Rolstoniellus Rider, 1997

• Sciocorini Amyot & Serville, 1843 selected genera:
  • Dyroderes Spinola, 1837
  • Sciocoris Fallén, 1829
• Strachiini Mulsant & Rey, 1866 selected genera:
  • Afrania Stål, 1865
  • Bagrada Stål, 1862
  • Eurydema Laporte de Castelnau, 1833
  • Murgantia (bug) Stål, 1862
  • Stenozygum Fieber, 1861
  • Trochiscocoris Reuter, 1890
• Triplatyxini Cachan, 1952

1. Anoano Cachan, 1952
2. Nene (bug) Kment, 2015
3. Tricompastes Cachan, 1952
4. Triplatyx Horváth, 1904

Genera incertae sedis

1. Amphidexius Bergroth, 1918
2. Bachesua Gross, 1975
3. Birna McDonald, 2006
4. Brizocoris Gross, 1975
5. Buthumka Gross, 1975
6. Cooperocoris Gross, 1975
7. Dysnoetus Bergroth, 1918
8. Eremophilacoris Gross, 1975
9. Gadarscama Reuter, 1887
10. Ippatha Distant, 1910
11. Kitsonia Gross, 1975
12. Kitsoniocoris Rider, 1998
13. Kumbutha Distant, 1910
14. Lakhonia Yang, 1936
15. Macrocarenoides Gross, 1975
16. Macrocarenus Stål, 1873
17. Mcphersonarcys Thomas, 2012
18. Muscanda Walker, 1868
19. Ochisme Kirkaldy, 1904
20. Ooldeon Gross, 1975
21. Patanius Rolston, 1987
22. Pirricoris Gross, 1975
23. Poecilotoma Dallas, 1851
24. Riaziana Hasan, Afzal & Ahmad, 1989
25. Senectius Rolston, 1987
26. Tepperocoris Gross, 1975
27. Thryptomenecoris Gross, 1975
28. Trachyops Dallas, 1851
29. Turrubulana Distant, 1910
30. Xiengia Distant, 1921
31. Zouicoris Zheng, 1986

Notable species

• Acrostemum hilare – green stink bug
• Bagrada hilaris – bagrada bug
• Bathycoelia thalassina - African cocoa shield bug
• Halyomorpha halys – brown marmorated stink bug
• Nezara viridula - green vegetable bug
• Pentatoma rufipes – forest bug
• Piezodorus lituratus – gorse shield bug

References

1. Faúndez, E. I.; Rider, D. A.; Garcete-Barret, B. (2018). "New records of Alcaeorrhynchus grandis (Dallas, 1851)(Heteroptera: Pentatomidae) from Paraguay, with description of a teratological case". Boletín del Museo Nacional de Historia Natural del Paraguay. 22 (1): 27–29.
2. Faúndez, E. I.; Rider, D. A. (2017). "The brown marmorated stink bug Halyomorpha halys (Stål, 1855)(Heteroptera: Pentatomidae) in Chile". Arquivos Entomolóxicos. 17: 305–307.
3. Faúndez, E. I. (2018). "From agricultural to household pest: The case of the painted bug Bagrada hilaris (Burmeister)(Heteroptera: Pentatomidae) in Chile". Journal of Medical Entomology. 55 (5): 1365–1368. doi:10.1093/jme/tjy091. PMID 29931070.
4. Rider, D.A., Schwertner, C.F., Vilimová, J., Redei, D., Kment, P. & D.B. Thomas. 2018. Pp. 25-204. Higher systematics of Pentatomidea in McPherson, J. E. (Ed.). Invasive Stink Bugs and Related Species (Pentatomoidea): Biology, Higher Systematics, Semiochemistry, and Management. CRC Press, Florida.
5. Zicha, Ondrej. "BioLib: Biological library". www.biolib.cz.

External links

• Data related to Pentatominae at Wikispecies
• Media related to Pentatominae at Wikimedia Commons

• Pentatominae
• Hemiptera subfamilies