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Cladoxylopsida Temporal range: Middle Devonian–Early Carboniferous PreꞒ Ꞓ O S D C P T J K Pg N
Wattieza
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Pteridophyta (?)
Class:	†Cladoxylopsida Novák (1930)
Orders
†Cladoxylales †Hyeniales †Iridopteridales †Pseudosporochnales ?†Foozia

The cladoxylopsids are an extinct group of plants related to ferns and sphenopsids.

They had a central trunk, from the top of which several lateral branches were attached. Fossils of these plants originate in the Middle Devonian to Early Carboniferous periods (around 390 to 320 million years ago), mostly just as stems.

Cladoxylopsida contains two orders. The order Hyeniales is now included in Pseudosporochnales.

Intact fossils of the Middle Devonian cladoxylopsid Wattieza show it to have been a tree, the earliest identified in the fossil record as of 2007. In 2019, experts from Cardiff University, UK; Binghamton University and the New York State Museum discovered more fossils of Cladoxylopsida and Archaeopteris in a quarry in Cairo, New York.

A recent (2017) discovery in Xinjiang in China of early Late Devonian (Frasnian, ca. 374 Ma) silicified fossil cladoxylopsid tree trunks (diameter up to c.70 cm) with preserved cellular anatomy showed an internal arrangement with many xylem bundles in the outer part and none in the interior; each bundle was surrounded by its own cambium layer, by which the tree's trunk widened.

References

1. "Introduction to the Cladoxylopsida". University of California Museum of Paleontology. Archived from the original on November 5, 2021. Retrieved January 10, 2022.
2. Thomas N. Taylor, Edith L. Taylor, Michael Krings: Paleobotany. The Biology and Evolution of Fossil Plants. Second Edition, Academic Press 2009, ISBN 978-0-12-373972-8 , p. 387-401, 1028
3. Foster, Laura (Dec 19, 2019). "World's oldest fossil trees uncovered in New York". BBC News Online. Retrieved Dec 19, 2019.
4. Xu, Hong-He; Berry, Christopher M.; Stein, William E.; Wang, Yi; Tang, Peng; Fu, Qiang (2017). "Unique growth strategy in the Earth's first trees revealed in silicified fossil trunks from China". Proceedings of the National Academy of Sciences. 114 (45): 12009–12014. Bibcode:2017PNAS..11412009X. doi:10.1073/pnas.1708241114. PMC 5692553. PMID 29078324.
5. https://www.cardiff.ac.uk/news/view/981090-worlds-oldest-and-most-complex-trees (with image of trunk cross-section)
6. "Prehistoric tree mystery baffles scientists". Independent.co.uk. 2017-10-23.

• Stein, William E.; Mannolini, Frank; Hernick, Linda Vanaller; Landing, Ed; Berry, Christopher M. (2007). "Giant cladoxylopsid trees resolve the enigma of the Earth's earliest forest stumps at Gilboa". Nature. 446 (7138): 904–907. Bibcode:2007Natur.446..904S. doi:10.1038/nature05705. PMID 17443185. S2CID 2575688.

External links

Media related to Cladoxylopsida at Wikimedia Commons Links with images:

• UC Museum of Paleontology
• https://sites.google.com/site/paleoplant/classification/cladoxylopsid

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• Devonian plants
• Middle Devonian first appearances
• Mississippian extinctions
• Plant classes
• Fern stubs
• Devonian plant stubs
• Paleozoic plant stubs
• Carboniferous stubs