Episoriculus - Misplaced Pages

Genus of mammals

Episoriculus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Eulipotyphla
Family:	Soricidae
Tribe:	Nectogalini
Genus:	Episoriculus Ellerman & Morrison-Scott, 1966
Type species
Sorex caudatus
Species
See text

Episoriculus is a genus of shrew in the red-toothed shrew subfamily. Its common is brown-toothed shrew. It has been described as a subgenus to Soriculus in the past. The genus occurs at a number of locations in Asia, including Nepal and China.

Species

Currently, the four identified species of the genus Episoriculus include:

Hodgson's brown-toothed shrew (E. caudatus)
- E. c. sacratus
- E. c. umbrinus
Taiwanese brown-toothed shrew (E. fumidus)
Long-tailed brown-toothed shrew (E. leucops)
- E. l. leucops
- E. l. baileyi
Long-tailed mountain shrew (E. macrurus)

The monophyly of the genus has been questioned, as E. fumidus has been found to be only distantly related to other species in the genus.

Phylogeny of Nectogalini based on DNA and morphological characters after Bover et al. (2018).

Nectogalini

Episoriculus fumidus

	Chimarrogale himalayica (Himalayan water shrew)

	Nectogale elegans (elegant water shrew)

Neomys

	Neomys fodiens (water shrew)

	†Neomys newtoni

Soriculus nigrescens (Himalayan shrew)

	†Nesiotites hidalgo

	†Asoriculus gibberodon

Episoriculus

	Episoriculus leucops

	Episoriculus caudatus

Episoriculus macrurus

Chodsigoa

	Chodsigoa parca

	Chodsigoa sodalis

Chodsigoa hypsibia

References

Nomenclator Zoologicus Record Detail. Ubio.org. Retrieved on 2011-10-21.
Data Use Agreement - GBIF Portal. Data.gbif.org (2007-02-22). Retrieved on 2011-10-21.
Namebank Record Detail. Ubio.org (2003-04-14). Retrieved on 2011-10-21.
"ION: Index to Organism Names".
Namebank Record Detail. Ubio.org (2005-10-11). Retrieved on 2011-10-21.
"ION: Index to Organism Names".
ADW: Episoriculus: Pictures. Animaldiversity.ummz.umich.edu. Retrieved on 2011-10-21.
Mammal Species of the World - Browse: Episoriculus. Bucknell.edu. Retrieved on 2011-10-21.
ITIS Standard Report Page: Episoriculus. Itis.gov. Retrieved on 2011-10-21.
Namebank Record Detail. Ubio.org (2007-05-23). Retrieved on 2011-10-21.
Data Use Agreement - GBIF Portal. Data.gbif.org (2007-02-22). Retrieved on 2011-10-21.
Episoriculus - Encyclopedia of Life. EOL. Retrieved on 2011-10-21.
"Catalogue of Life - 2010 Annual Checklist :: Taxonomic tree". catalogueoflife.org. Retrieved 17 June 2015.
Bover, Pere; Mitchell, Kieren J.; Llamas, Bastien; Rofes, Juan; Thomson, Vicki A.; Cuenca-Bescós, Gloria; Alcover, Josep A.; Cooper, Alan; Pons, Joan (August 2018). "Molecular phylogenetics supports the origin of an endemic Balearic shrew lineage (Nesiotites) coincident with the Messinian Salinity Crisis". Molecular Phylogenetics and Evolution. 125: 188–195. Bibcode:2018MolPE.125..188B. doi:10.1016/j.ympev.2018.03.028. PMID 29608962. S2CID 5010906.

Atelerix	Four-toed hedgehog (A. albiventris) North African hedgehog (A. algirus) Southern African hedgehog (A. frontalis) Somali hedgehog (A. sclateri)
Erinaceus	Amur hedgehog (E. amurensis) Southern white-breasted hedgehog (E. concolor) West European hedgehog (E. europaeus) Northern white-breasted hedgehog (E. roumanicus)
Hemiechinus	Long-eared hedgehog (H. auritus) Indian long-eared hedgehog (H. collaris)
Mesechinus	Daurian hedgehog (M. dauuricus) Hugh's hedgehog (M. hughi) Gaoligong forest hedgehog (M. wangi) Small-toothed forest hedgehog (M. miodon) Eastern forest hedgehog (M. orientalis)
Paraechinus	Desert hedgehog (P. aethiopicus) Brandt's hedgehog (P. hypomelas) Indian hedgehog (P. micropus) Bare-bellied hedgehog (P. nudiventris)

Family: Erinaceidae
subfamily: Galericinae

Echinosorex	Moonrat (E. gymnura)
Hylomys	Dwarf gymnure (H. parvus) Javan short-tailed gymnure (H. suillus) Bornean short-tailed gymnure (H. dorsalis) Max's short-tailed gymnure (H. maxi) Dalat gymnure (H. macarong) Northern short-tailed gymnure (H. peguensis) Leuser gymnure (H. vorax)
Neohylomys	Hainan gymnure (N. hainanensis)
Neotetracus	Shrew gymnure (N. sinensis)
Otohylomys	Long-eared gymnure (O. megalotis)
Podogymnura	Dinagat gymnure (P. aureospinula) Mindanao gymnure (P. truei)

Family: Soricidae
subfamily: Crocidurinae

Crocidura
(White-toothed
shrews)

Family: Soricidae
subfamily: Crocidurinae (continued)

Diplomesodon	Piebald shrew (D. pulchellus)
Feroculus	Kelaart's long-clawed shrew (F. feroculus)
Palawanosorex	Palawan moss shrew (P. muscorum)
Paracrocidura (Large-headed shrews)	Grauer's large-headed shrew (P. graueri) Greater large-headed shrew (P. maxima) Lesser large-headed shrew (P. schoutedeni)
Ruwenzorisorex	Ruwenzori shrew (R. suncoides)
Scutisorex	Hero shrew (S. somereni) Thor's hero shrew (S. thori)
Solisorex	Pearson's long-clawed shrew (S. pearsoni)
Suncus	Taita shrew (S. aequatorius) Black shrew (S. ater) Day's shrew (S. dayi) Etruscan shrew (S. etruscus) Sri Lankan shrew (S. fellowesgordoni) Bornean pygmy shrew (S. hosei) Least dwarf shrew (S. infinitesimus) Greater dwarf shrew (S. lixus) Madagascan pygmy shrew (S. madagascariensis) Malayan pygmy shrew (S. malayanus) Climbing shrew (S. megalura) Flores shrew (S. mertensi) Asian highland shrew (S. montanus) Asian house shrew (S. murinus) Remy's pygmy shrew (S. remyi) Anderson's shrew (S. stoliczkanus) Lesser dwarf shrew (S. varilla) Jungle shrew (S. zeylanicus)
Sylvisorex (Forest shrews)	Dudu Akaibe's pygmy shrew (S. akaibei) Cameroonian forest shrew (S. cameruniensis) Corbet's forest shrew (S. corbeti) Grant's forest shrew (S. granti) Howell's forest shrew (S. howelli) Bioko forest shrew (S. isabellae) Johnston's forest shrew (S. johnstoni) Kongana shrew (S. konganensis) Moon forest shrew (S. lunaris) Mount Cameroon forest shrew (S. morio) Greater forest shrew (S. ollula) Lesser forest shrew (S. oriundus) Rain forest shrew (S. pluvialis) Volcano shrew (S. vulcanorum)

Family: Soricidae
subfamily: Soricinae

Anourosoricini

Anourosorex (Asian mole shrews)	Assam mole shrew (A. assamensis) Giant mole shrew (A. schmidi) Chinese mole shrew (A. squamipes) Taiwanese mole shrew (A. yamashinai)

Blarinellini

Blarinella (some Asiatic short-tailed shrews)	Asiatic short-tailed shrew (B. quadraticauda) Burmese short-tailed shrew (B. wardi)
Parablarinella (some Asiatic short-tailed shrews)	Indochinese short-tailed shrew (P. griselda) Anhui short-tailed shrew (P. latimaxillata)

Blarinini

Blarina
(American short-tailed
shrews)

Cryptotis
(Small-eared
shrews)

C. mexicana group: Mexican small-eared shrew (C. mexicana); Nelson's small-eared shrew (C. nelsoni); Grizzled Mexican small-eared shrew (C. obscura); Phillips' small-eared shrew (C. phillipsii)

C. goldmani set: Central Mexican broad-clawed shrew (C. alticola); Goldman's broad-clawed shrew (C. goldmani); Goodwin's broad-clawed shrew (C. goodwini); Guatemalan broad-clawed shrew (C. griseoventris); C. lacertosus; C. mam; Oaxacan broad-clawed shrew (C. peregrina)

C. nigrescens group: Eastern Cordillera small-footed shrew (C. brachyonyx); Colombian small-eared shrew (C. colombiana); Honduran small-eared shrew (C. hondurensis); Yucatan small-eared shrew (C. mayensis); Darién small-eared shrew (C. mera); Merriam's small-eared shrew (C. merriami); Blackish small-eared shrew (C. nigrescens)

C. thomasi group: Southern Colombian small-eared shrew (C. andinus); Ecuadorian small-eared shrew (C. equatoris); Rainer's small-eared shrew (C. huttereri); Medellín small-eared shrew (C. medellinia); Merida small-eared shrew (C. meridensis); Wandering small-eared shrew (C. montivaga); Peruvian small-eared shrew (C. peruviensis); Scaly-footed small-eared shrew (C. squaipes); Tamá small-eared shrew (C. tamensis); Thomas's small-eared shrew (C. thomasi)

C. parva group: Central American least shrew (C. orophila); North American least shrew (C. parva); Tropical small-eared shrew (C. tropicalis)

Ungrouped / relict: Enders's small-eared shrew (C. endersi); Talamancan small-eared shrew (C. gracilis); Big Mexican small-eared shrew (C. magna)

Nectogalini

Chimarrogale (Asiatic water shrews)	Malayan water shrew (C. hantu) Himalayan water shrew (C. himalayica) Bornean water shrew (C. phaeura) Japanese water shrew (C. platycephalus) Chinese water shrew (C. styani) Sumatran water shrew (C. sumatrana)
Chodsigoa	Van Sung's shrew (C. caovansunga) De Winton's shrew (C. hypsibia) Lamulate shrew (C. lamula) Lowe's shrew (C. parca) Pygmy brown-toothed shrew (C. parva) Salenski's shrew (C. salenskii) Smith's shrew (C. smithii) Lesser Taiwanese shrew (C. sodalis)
Episoriculus	Hodgsons's brown-toothed shrew (E. caudatus) Taiwanese brown-toothed shrew (E. fumidus) Long-tailed brown-toothed shrew (E. leucops) Long-tailed mountain shrew (E. macrurus)
Nectogale	Elegant water shrew (N. elegans)
Neomys	Mediterranean water shrew (N. anomalus) Eurasian water shrew (N. fodiens) Transcaucasian water shrew (N. teres)
Soriculus	Himalayan shrew (S. nigrescens)

Notiosoricini

Megasorex	Mexican shrew (M. gigas)
Notiosorex	Cockrum's gray shrew (N. cockrumi) Crawford's gray shrew (N. crawfordi) Large-eared gray shrew (N. evotis) Villa's gray shrew (N. villai)

Family: Soricidae
subfamily: Soricinae (tribe: Soricini)

Sorex
(Long-tailed
shrews)

Subgenus
Otisorex

S. vagrans complex: Glacier Bay water shrew (S. alaskanus); Baird's shrew (S. bairdii); Marsh shrew (S. bendirii); Montane shrew (S. monticolus); New Mexico shrew (S. neomexicanus); Pacific shrew (S. pacificus); American water shrew (S. palustris); Fog shrew (S. sonomae); Vagrant shrew (S. vagrans)

S. cinereus group: Kamchatka shrew (S. camtschatica); Cinereus shrew (S. cinereus); Prairie shrew (S. haydeni); Saint Lawrence Island shrew (S. jacksoni); Paramushir shrew (S. leucogaster); Southeastern shrew (S. longirostris); Mount Lyell shrew (S. lyelli); Portenko's shrew (S. portenkoi); Preble's shrew (S. preblei); Pribilof Island shrew (S. pribilofensis); Olympic shrew (S. rohweri); Barren ground shrew (S. ugyunak)

Subgenus
Sorex

S. alpinus group: Alpine shrew (S. alpinus); Ussuri shrew (S. mirabilis)

S. araneus group: Valais shrew (S. antinorii); Common shrew (S. araneus); Udine shrew (S. arunchi); Crowned shrew (S. coronatus); Siberian large-toothed shrew (S. daphaenodon); Iberian shrew (S. granarius); Caucasian shrew (S. satunini)

S. arcticus group: Arctic shrew (S. arcticus); Maritime shrew (S. maritimensis)

S. tundrensis group: Tien Shan shrew (S. asper); Gansu shrew (S. cansulus); Tundra shrew (S. tundrensis)

S. minutus group: Buchara shrew (S. buchariensis); Kozlov's shrew (S. kozlovi); Caucasian pygmy shrew (S. volnuchini)

S. caecutiens group: Laxmann's shrew (S. caecutiens); Taiga shrew (S. isodon); Eurasian least shrew (S. minutissimus); Eurasian pygmy shrew (S. minutus); Flat-skulled shrew (S. roboratus); Shinto shrew (S. shinto); Long-clawed shrew (S. unguiculatus)

S. gracillimus group: Slender shrew (S. gracillimus)

S. raddei group: Radde's shrew (S. raddei)

S. samniticus group: Apennine shrew (S. samniticus)

incertae sedis

Family: Soricidae
subfamily: Myosoricinae

Congosorex (Congo shrews)	Phillips's shrew (C. phillipsorum) Greater Congo shrew (C. polli) Lesser Congo shrew (C. verheyeni)
Myosorex (Mouse shrews)	Babault's mouse shrew (M. babaulti) Montane mouse shrew (M. blarina) Bururi forest shrew (M. bururiensis) Dark-footed mouse shrew (M. cafer) Eisentraut's mouse shrew (M. eisentrauti) Geata mouse shrew (M. geata) Nyika mouse shrew (M. gnoskei) Kihaule's mouse shrew (M. kihaulei) Long-tailed forest shrew (M. longicaudatus) Oku mouse shrew (M. okuensis) Rumpi mouse shrew (M. rumpii) Schaller's mouse shrew (M. schalleri) Sclater's mouse shrew (M. sclateri) Thin mouse shrew (M. tenuis) Forest shrew (M. varius) Kilimanjaro mouse shrew (M. zinki)
Surdisorex (African mole shrews)	Aberdare mole shrew (S. norae) Mount Kenya mole shrew (S. polulus)

Family: Talpidae

Scalopinae
(New World moles
and relatives)

Condylura	Star-nosed mole (C. cristata)
Parascalops	Hairy-tailed mole (P. breweri)
Scalopus	Eastern mole (S. aquaticus)
Scapanulus	Gansu mole (S. oweni)
Scapanus (Western North American moles)	Broad-footed mole (S. latimanus) Coast mole (S. orarius) Townsend's mole (S. townsendii)

Talpinae
(Old World moles
and relatives)

Desmana	Russian desman (D. moschata)
Dymecodon	True's shrew mole (D. pilirostris)
Euroscaptor	Greater Chinese mole (E. grandis) Kloss's mole (E. klossi) Long-nosed mole (E. longirostris) Himalayan mole (E. micrura) Small-toothed mole (E. parvidens) Vietnamese mole (E. subanura)
Galemys	Pyrenean desman (G. pyrenaicus)
Mogera	Echigo mole (M. etigo) Insular mole (M. insularis) Kano mole (M. kanoana) Kobe mole (M. kobeae) Small Japanese mole (M. imaizumii) Large mole (M. robusta) Sado mole (M. tokudae) Japanese mole (M. wogura) Senkaku mole (M. uchidai)
Neurotrichus	American shrew mole (N. gibbsii)
Oreoscaptor	Japanese mountain mole (O. mizura)
Parascaptor	White-tailed mole (P. leucura)
Scaptochirus	Short-faced mole (S. moschatus)
Scaptonyx	Long-tailed mole (S. fusicaudus)
Talpa	Altai mole (T. altaica) Aquitanian mole (T. aquitania) Blind mole (T. caeca) Caucasian mole (T. caucasica) European mole (T. europaea) Père David's mole (T. davidiana) Levant mole (T. levantis) Martino's mole (T. martinorum) Spanish mole (T. occidentalis) Ognev's mole (T. ognevi) Roman mole (T. romana) Balkan mole (T. stankovici) Talysch mole (T. talyschensis)
Urotrichus	Japanese shrew mole (U. talpoides)

Uropsilinae
(Chinese shrew-like
moles)

Uropsilus	Equivalent-teeth shrew mole (U. aequodonenia) Anderson's shrew mole (U. andersoni) Black-backed shrew mole (U. atronates) Dabie Mountains shrew mole (U. dabieshanensis) Gracile shrew mole (U. gracilis) Inquisitive shrew mole (U. investigator) Snow Mountain shrew mole (U. nivatus) Chinese shrew mole (U. soricipes)

Family: Solenodontidae

Atopogale	Cuban solenodon (A. cubana)
Solenodon	Hispaniolan solenodon (S. paradoxus)

Taxon identifiers
Episoriculus	Wikidata: Q2894104 Wikispecies: Episoriculus BOLD: 6855 CoL: 62RXN EoL: 34570 GBIF: 2435880 iNaturalist: 46901 IRMNG: 1035443 ITIS: 709694 MSW: 13700420 NCBI: 269272 Open Tree of Life: 1096818 Paleobiology Database: 40468

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