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Rhinoceros Temporal range: Late Miocene-Recent 9–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Indian rhinoceros (R. unicornis)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Perissodactyla
Family:	Rhinocerotidae
Tribe:	Rhinocerotini
Genus:	Rhinoceros Linnaeus, 1758
Type species
Rhinoceros unicornis Linnaeus, 1758
Species
R. unicornis R. sondaicus

Rhinoceros is a genus comprising one-horned rhinoceroses. This scientific name was proposed by Swedish taxonomist Carl Linnaeus in 1758. The genus contains two species, the Indian rhinoceros (Rhinoceros unicornis) and the Javan rhinoceros (Rhinoceros sondaicus). Although both members are threatened, the Javan rhinoceros is one of the most endangered large mammals in the world with only 60 individuals surviving in Java (Indonesia). The word 'rhinoceros' is of Greek origin meaning "nose-horn".

Etymology

The genus name Rhinoceros is a combination of the ancient Greek words ῥίς (rhis) meaning 'nose' and κέρας (keras) meaning 'horn of an animal'.

Classification

The genus Rhinoceros comprises:

• Indian rhinoceros (R. unicornis) Linnaeus, 1758 Indian subcontinent
• Javan rhinoceros (R. sondaicus) Desmarest, 1822 Southeast Asia
• †R. sivalensis Falconer and Cautley, 1846 northern Indian subcontinent (Siwalik Hills) Pliocene-Early Pleistocene
• †R. platyrhinus Falconer and Cautley 1847 syn Punjabitherium Khan (1971) Upper Siwaliks, Indian subcontinent, Early Pleistocene to early Middle Pleistocene, India. Largest species in the genus.
• †R. sinensis Owen, 1870 Has been used as a wastebasket taxon used to refer to rhinoceros material from the Pleistocene of China, with various specimens belong to other Rhinoceros species, Dicerorhinus and Stephanorhinus, though it is possible that some remains attributed to Rhinoceros sinensis represents a valid and distinct species of Rhinoceros.

The earliest remains of the genus are known from the Late Miocene, represented by remains such as an indeterminate species found in deposits in Myanmar dating to around 8-9 million years ago.

The species "Rhinoceros" philippinensis from the early Middle Pleistocene of the Philippines and "Rhinoceros" sinensis hayasakai from the Early-Middle Pleistocene of Taiwan have been transferred to Nesorhinus, which appears to be closely related to Rhinoceros. While Rhinoceros fusuiensis Yan et al. 2014 from the Early Pleistocene of South China has been transferred to Dicerorhinus.

 Relationships among Late Pleistocene and modern rhinoceros genera, based on nuclear DNA, after Liu et al, 2021:

Bayesian morphological phylogeny after Pandolfi, 2023 Note: This excludes living African rhinoceros species.

References

1. ^ Linnæus, C. (1758). "Rhinoceros unicornis". Caroli Linnæi Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Holmiae: Salvius. p. 56.
2. Liddell, H. G. & Scott, R. (1940). "ῥίς". A Greek-English Lexicon (Revised and augmented ed.). Oxford: Clarendon Press.
3. Liddell, H. G. & Scott, R. (1940). "κέρᾳ". A Greek-English Lexicon (Revised and augmented ed.). Oxford: Clarendon Press.
4. Desmarest, A. G. (1822). "Rhinocéros des Îles de La Sonde". Mammalogie, ou, Description des espèces de mammifères. Vol. 2. Paris: Mme Agasse. pp. 399–400.
5. ^ Pandolfi, Luca; Maiorino, Leonardo (2016-02-06). "Reassessment of the largest Pleistocene rhinocerotine Rhinoceros platyrhinus (Mammalia, Rhinocerotidae) from the Upper Siwaliks (Siwalik Hills, India)" (PDF). Journal of Vertebrate Paleontology. 36 (2): e1071266. Bibcode:2016JVPal..36E1266P. doi:10.1080/02724634.2015.1071266. ISSN 0272-4634. S2CID 86893957.
6. Schepartz, L. A.; Miller-Antonio, S. (2010). "Taphonomy, life history, and human exploitation of Rhinoceros sinensis at the Middle Pleistocene site of Panxian Dadong, Guizhou, China". International Journal of Osteoarchaeology. 20 (3): 253–268. doi:10.1002/oa.1025.
7. Antoine, Pierre-Olivier (March 2012). "Pleistocene and Holocene rhinocerotids (Mammalia, Perissodactyla) from the Indochinese Peninsula". Comptes Rendus Palevol. 11 (2–3): 159–168. Bibcode:2012CRPal..11..159A. doi:10.1016/j.crpv.2011.03.002.
8. Tong, Hao-wen (November 2012). "Evolution of the non-Coelodonta dicerorhine lineage in China". Comptes Rendus Palevol. 11 (8): 555–562. Bibcode:2012CRPal..11..555T. doi:10.1016/j.crpv.2012.06.002.
9. Longuet, Morgane; Zin-Maung-Maung-Thein; Thaung-Htike; Man-Thit-Nyein; Takai, Masanaru (2024-08-02). "New fossil remains of Rhinocerotidae (Perissodactyla) from the early Late Miocene Tebingan area, central Myanmar". Historical Biology. 36 (8): 1468–1481. doi:10.1080/08912963.2023.2218873. ISSN 0891-2963.
10. ^ Antoine, P.-O.; Reyes, M. C.; Amano, N.; Bautista, A. P.; Chang, C.-H.; Claude, J.; De Vos, J.; Ingicco, T. (2021). "A new rhinoceros clade from the Pleistocene of Asia sheds light on mammal dispersals to the Philippines". Zoological Journal of the Linnean Society. 194 (2): 416–430. doi:10.1093/zoolinnean/zlab009.
11. Yan, Yaling; Wang, Yuan; Jin, Changzhu; Mead, Jim I. (December 2014). "New remains of Rhinoceros (Rhinocerotidae, Perissodactyla, Mammalia) associated with Gigantopithecus blacki from the Early Pleistocene Yanliang Cave, Fusui, South China". Quaternary International. 354: 110–121. Bibcode:2014QuInt.354..110Y. doi:10.1016/j.quaint.2014.01.004.
12. Pandolfi, Luca (2023-01-19). "Reassessing the phylogeny of Quaternary Eurasian Rhinocerotidae". Journal of Quaternary Science. 38 (3): 291–294. Bibcode:2023JQS....38..291P. doi:10.1002/jqs.3496. hdl:11563/163194. ISSN 0267-8179. S2CID 256167036.
13. Liu, Shanlin; Westbury, Michael V.; Dussex, Nicolas; Mitchell, Kieren J.; Sinding, Mikkel-Holger S.; Heintzman, Peter D.; Duchêne, David A.; Kapp, Joshua D.; von Seth, Johanna; Heiniger, Holly; Sánchez-Barreiro, Fátima (16 September 2021). "Ancient and modern genomes unravel the evolutionary history of the rhinoceros family". Cell. 184 (19): 4874–4885.e16. doi:10.1016/j.cell.2021.07.032. hdl:10230/48693. PMID 34433011. S2CID 237273079.
14. Pandolfi, Luca (April 2023). "Reassessing the phylogeny of Quaternary Eurasian Rhinocerotidae". Journal of Quaternary Science. 38 (3): 291–294. Bibcode:2023JQS....38..291P. doi:10.1002/jqs.3496. hdl:11563/163194. ISSN 0267-8179.

External links

v t e Perissodactyla
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Perissodactyla
Perissodactyla Incertae sedis †Ghazijhippus †Mesolambdolophus †Olbitherium †Anthracobunia Radinskya †Anthracobunidae Anthracobune Hsanotherium Jozaria Obergfellia †Cambaytheriidae Cambaytherium Kalitherium Nakusia Perissobune †Brontotheriidae See Brontotheriidae Equoidea †Cymbalophus †Hallensia †Pliolophus †Systemodon Equidae See Equidae †Palaeotheriidae †Pachynolophinae Anchilophus Bepitherium Eurohippus Hyracotherium Lijiangia Lophiohippus Lophiotherium Metanchilophus Orolophus Pachynolophus Paranchilophus Propalaeotherium Qianohippus †Palaeotheriinae Cantabrotherium Iberolophus Idiodontherium Leptolophus Mekodontherium Metaplagiolophus Palaeotherium Paraplagiolophus Plagiolophus Pseudopalaeotherium †Hyopsodontidae? Aletodon Chacomylus Dorraletes Haplaletes Haplomylus Hyopsodus Litomylus Louisina Monshyus Paratricuspiodon Paschatherium Phenacodaptes Tricuspiodon †Phenacodontidae? Tetraclaenodon †Meniscotheriinae Ectocion Meniscotherium †Phenacodontinae Almogaver Copecion Eodesmatodon Lophocion Phenacodus Tapiromorpha See below↓
Tapiromorpha Incertae sedis †Cambaylophus †Skopaiolophus †Isectolophidae Chowliia Gandheralophus Karagalax Meridiolophus Orientolophus Sastrilophus †Homogalaxinae Homogalax †Isectolophinae Isectolophus Schizolophodon †Ancylopoda See Ancylopoda Ceratomorpha Tapiroidea †Deperetellidae Bahinolophus Deperetella Irenolophus Teleolophus †Helaletidae Colodon Dilophodon Helaletes Heptodon Heteraletes Paracolodon Plesiocolopirus Thuliadanta? Vastanolophus Tapiridae †Eotapirus †Miotapirus †Nexuotapirus †Paratapirus †Plesiotapirus †Protapirus †Tapiravus Tapirus Rhinocerotoidea See below↓
Rhinocerotoidea Rhinocerotoidea sensu lato †Breviodon †Fouchia †Minchenoletes †Siamolophus †Triplopus †Yimengia †Indolophidae Indolophus Rhinocerotoidea sensu stricto †Selenaletes †Hyrachyidae Hyrachyus †Eggysodontidae Allacerops Eggysodon Guangnanodon Proeggysodon †Hyracodontidae Ardynia Ephyrachyus Epitriplophus Hyracodon Prohyracodon Rhodopagus Triplopides †Amynodontidae Amynodontopsis Armania Cadurcotherium Caenolophus Huananodon Hypsamynodon Procadurcodon Rostriamynodon Teilhardia †Amynodontinae Amynodon †Cadurcodontini Cadurcodon Lushiamynodon Sharamynodon Sianodon †Metamynodontini Gigantamynodon Megalamynodon Metamynodon Paramynodon Zaisanamynodon †Paraceratheriidae †Forstercooperiinae Forstercooperia Gobicerops Pappaceras †Paraceratheriinae Aralotherium Dzungarotherium Juxia Paraceratherium Urtinotherium Rhinocerotidae See below↓
Rhinocerotidae Rhinocerotidae †Amphicaenopus †Epiaceratherium †Guixia †Teletaceras †Uintaceras †Woodoceras †Aceratheriinae Aceratherium Acerorhinus Alicornops Aphelops Chilotheridium Chilotherium Floridaceras Galushaceras Hoploaceratherium Mesaceratherium Peraceras Persiatherium Plesiaceratherium Ronzotherium Shansirhinus Sinorhinus Subchilotherium †Elasmotheriinae Gulfoceras Victoriaceros †Diceratheriini Diceratherium Penetrigonias Skinneroceras Subhyracodon Trigonias †Elasmotheriini Bugtirhinus Caementodon Elasmotherium Hispanotherium Iranotherium Kenyatherium Meninatherium Menoceras Ninxiatherium Ningxiatherium Ougandatherium Parelasmotherium Procoelodonta Sinotherium Tongxinotherium Rhinocerotinae †Dromoceratherium †Lartetotherium †Molassitherium †Protaceratherium †Teleoceratini Aprotodon Brachydiceratherium Brachypotherium Diaceratherium Prosantorhinus Shennongtherium Teleoceras Rhinocerotina Dicerorhinini †Coelodonta Dicerorhinus †Dihoplus †Pliorhinus †Stephanorhinus Dicerotini Ceratotherium Diceros †Miodiceros †Paradiceros Rhinocerotini †Gaindatherium †Nesorhinus Rhinoceros †Rusingaceros

v t e Extant Perissodactyla (Odd-toed ungulates) species by suborder
Kingdom Animalia Phylum Chordata Class Mammalia Infraclass Eutheria Superorder Laurasiatheria
Hippomorpha Equidae (Horse family) Equus (including Zebras) Subgenus Equus: Wild horse (E. ferus) Przewalski's horse (E. f. przewalskii) Domestic horse (E. caballus) Subgenus Asinus: African wild ass (E. africanus) Donkey (E. asinus) Onager (E. hemionus) Kiang (E. kiang) Subgenus Hippotigris: Plains zebra (E. quagga) Mountain zebra (E. zebra) Grévy's zebra (E. grevyi)
Ceratomorpha Rhinocerotidae (Rhinoceroses) Rhinoceros Indian rhinoceros (R. unicornis) Javan rhinoceros (R. sondaicus) Dicerorhinus Sumatran rhinoceros (D. sumatrensis) Ceratotherium White rhinoceros (C. simum) Diceros Black rhinoceros (D. bicornis) Tapiridae (Tapirs) Tapirus Baird's tapir (T. bairdii) Mountain tapir (T. pinchaque) South American tapir (T. terrestris) Malayan tapir (T. indicus)
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