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Vulpini
Clockwise from top: red fox, bat-eared fox, tanuki
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Subfamily:	Caninae
Tribe:	Vulpini Hemprich and Ehrenberg, 1832
Genera
†Ferrucyon †Metalopex Tedford, Wang, & Taylor 2008 Nyctereutes Otocyon †Prototocyon Vulpes

Vulpini is a taxonomic rank which represents the fox-like tribe of the subfamily Caninae (the canines), and is sister to the dog-like tribe Canini.

Genera

Image	Genus	Species
	Nyctereutes Temminck, 1838	Nyctereutes procyonoides Nyctereutes viverrinus †Nyctereutes abdeslami †Nyctereutes donnezani †Nyctereutes lockwoodi †Nyctereutes megamastoides †Nyctereutes sinensis †Nyctereutes tingi †Nyctereutes vinetorum
	Otocyon S. Müller, 1835	Otocyon megalotis
	Vulpes Garsault, 1764	Vulpes bengalensis Vulpes cana Vulpes chama Vulpes corsac Vulpes ferrilata Vulpes lagopus Vulpes macrotis Vulpes pallida Vulpes rueppellii Vulpes velox Vulpes vulpes Vulpes zerda †Vulpes hassani †Vulpes praeglacialis †Vulpes qiuzhudingi †Vulpes riffautae †Vulpes rooki †Vulpes skinneri †Vulpes stenognathus †Vulpes gigas
	†Ferrucyon Ruiz-Ramoni et al., 2020	†Ferrucyon avius
	†Metalopex S. Müller, 1835	†Metalopex bakeri Tedford et al. 2009 †Metalopex macconnelli Tedford et al. 2009 †Metalopex merriami Tedford and Wang, 2008
	†Prototocyon Pohle, 1928	†Prototocyon curvipalatus †Prototocyon recki

Taxonomy

The taxonomy of Carnivora in general and Canidae in particular correlates with various diagnostic features of the dentition and basicranium. Regarding Vulpini, Tedford has remarked:

These small canids are distinguished from all other Caninae in possessing a wide paroccipital process that is broadly sutured to the posterior surface of the bulla with a short and laterally turned free tip that barely extends below the body of the process. The presence of a metaconule and postprotocrista on M2 of vulpines represents the culmination of a reversal that began with late Leptocyon species to resume the form of the primitive canine M2.

— Richard H. Tedford

The cladogram below is based on the phylogeny of Lindblad-Toh (2005) modified to incorporate recent findings on Vulpes.

References

1. ^ Damián Ruiz-Ramoni; Francisco Juan Prevosti; Saverio Bartolini Lucenti; Marisol Montellano-Ballesteros; Ana Luisa Carreño (2020). "The Pliocene canid Cerdocyon avius was not the type of fox that we thought". Journal of Vertebrate Paleontology. 40 (2): e1774889. doi:10.1080/02724634.2020.1774889. S2CID 222214868.
2. ^ Tedford, Richard H.; Wang, Xiaoming; Taylor, Beryl E. (2009). "Phylogenetic Systematics of the North American Fossil Caninae (Carnivora: Canidae)" (PDF). Bulletin of the American Museum of Natural History. 325: 1–218. doi:10.1206/574.1. hdl:2246/5999. S2CID 83594819.
3. Lindblad-Toh, Kerstin; Wade, Claire M.; Mikkelsen, Tarjei S.; Karlsson, Elinor K.; Jaffe, David B.; Kamal, Michael; et al. (2005). "Genome sequence, comparative analysis and haplotype structure of the domestic dog". Nature. 438 (7069): 803–819. Bibcode:2005Natur.438..803L. doi:10.1038/nature04338. PMID 16341006.
4. Zhao, Chao; Zhang, Honghai; Liu, Guangshuai; Yang, Xiufeng; Zhang, Jin (2016). "The complete mitochondrial genome of the Tibetan fox (Vulpes ferrilata) and implications for the phylogeny of Canidae". Comptes Rendus Biologies. 339 (2): 68–77. doi:10.1016/j.crvi.2015.11.005. ISSN 1631-0691. PMID 26868757.

v t e Extinct Canidae
Kingdom Animalia Phylum Chordata Class Mammalia Order: Carnivora Suborder: Caniformia
Canidae Canidae †Prohesperocyon †Hesperocyoninae Hesperocyon Mesocyon Cynodesmus Sunkahetanka Philotrox Enhydrocyon Paraenhydrocyon Caedocyon Ectopocynus Osbornodon O. brachypus O. fricki O. iamonensis O. renjiei O. sesnoni O. scitulus O. wangi †Borophaginae Archaeocyon Oxetocyon Otarocyon Rhizocyon Phlaocyonini Cynarctoides Phlaocyon P. achoros P. annectens P. latidens P. leucosteus P. mariae P. marslandensis P. minor P. multicuspus P. taylori P. yatkolai Borophagini Cormocyon Desmocyon Metatomarctus Euoplocyon Psalidocyon Microtomarctus Protomarctus Tephrocyon Cynarctina Paracynarctus Cynarctus Aelurodontina Tomarctus Aelurodon Borophagina Paratomarctus Carpocyon Protepicyon Epicyon Borophagus B. diversidens B. dudleyi B. hilli B. littoralis B. orc B. parvus B. pugnator B. secundus Caninae see below↓
Caninae Caninae †Leptocyon †Metalopex Urocyon †U. citrinus †U. galushai †U. minicephalus †U. progressus †U. webbi Vulpini †Ferrucyon †Prototocyon Nyctereutes (raccoon dogs) †N. abdeslami †N. donnezani †N. lockwoodi †N. megamastoides †N. sinensis †N. tingi †N. vinetorum Vulpes (true foxes) †V. hassani †V. praeglacialis †V. qiuzhudingi †V. riffautae †V. rooki †V. skinneri †V. stenognathus †V. gigas Canini (true dogs) †Canis gezi †Nurocyon Cerdocyonina (zorro) †Protocyon †Theriodictis Speothos †Pleistocene bush dog (S. pacivorus) †Dusicyon Falkland Islands wolf (D. australis) D. avus D. cultridens Lycalopex (South American foxes) †L. cultridens (?) †L. ensenadensis Canina (wolf-like canids) see below↓
Canina (wolf-like canids) Canina †Dire wolf (Aenocyon) †Sardinian dhole (Cynotherium) †Trinil dog (Mececyon) †Merriam's dog (Megacyon) †Xenocyon †Eucyon E. davisi E. debonisi E. ferox E. khoikhoi Lycaon †L. magnus †L. sekowei Dhole (Cuon alpinus) †C. a. antiquus †C. a. caucasicus †European dhole (C. a. europaeus) †Tien Shan dhole (C. a. hesperius) Canis †C. antonii †C. apolloniensis †Armbruster's wolf (C. armbrusteri) †C. arnensis †C. borjgali †C. cedazoensis †C. chihliensis †Edward's wolf (C. edwardii) †Etruscan wolf (C. etruscus) †C. lepophagus †Mosbach wolf (C. mosbachensis) †C. nehringi †C. palmidens †C. variabilis Coyote (C. latrans) †Pleistocene coyote (C. l. orcutti) Red wolf (C. rufus) †Florida black wolf (C. r. floridanus) †Gregory's wolf (C. r. gregoryi) Wolf (C. lupus) †Prehistoric C. l. bohemica C. l. lunellensis C. l. maximus C. l. santenaisiensis Cave wolf (C. l. spelaeus) Beringian wolf †Recently extinct Sicilian wolf (C. l. cristaldii) Hokkaido wolf (C. l. hattai) Japanese wolf (C. l. hodophilax) Austro-Hungarian wolf Synonymized under C. l. arctos: Bernard's wolf (C. l. bernardi) Synonymized under C. l. occidentalis: Kenai Peninsula wolf (C. l. alces) Manitoba wolf (C. l. griseoalbus) Synonymized under C. l. nubilus: Newfoundland wolf (C. l. beothucus) Cascade mountain wolf (C. l. fuscus) Mogollon mountain wolf (C. l. mogollonensis) Texas wolf (C. l. monstrabilis) Southern Rocky Mountain wolf (C. l. youngi)
See also: Evolution of the wolf Category:Extinct canids

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