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{{good article}} | |||
{{Citations missing|date=September 2007}} | |||
{{Short description|Domesticated omnivorous even-toed ungulate}} | |||
{{AnimalsCleanup}} | |||
{{About|the domestic farm animal|other animals known as "swine"|Suina|other uses|Swine (disambiguation)|and|Pig (disambiguation)}} | |||
{{Expand|date=April 2008}} | |||
{{pp-protected|reason=Restoring per ], which used to reside here.|small=yes}} | |||
{{Taxobox | |||
{{Use dmy dates|date=February 2022}} | |||
| name = Domestic Pig | |||
{{Speciesbox | |||
| status = {{StatusDomesticated}} | |||
| name = Pig | |||
| image = Sus scrofa scrofa.jpg | |||
| image = Pig farm Vampula 1.jpg | |||
| image_width = 250px | |||
| image_caption = |
| image_caption = Domestic pigs | ||
| status = DOM | |||
| regnum = ]ia | |||
| genus = Sus | |||
| phylum = ] | |||
| species = domesticus | |||
| classis = ]ia | |||
| authority = ], 1777 | |||
| ordo = ] | |||
| synonyms_ref = <ref name="Groves 1995">{{cite journal |last=Groves |first=Colin P. |year=1995 |title=On the nomenclature of domestic animals |journal=Bulletin of Zoological Nomenclature |volume=52 |issue=2 |pages=137–141 |doi=10.5962/bhl.part.6749 |doi-access=free}} </ref> | |||
| familia = ] | |||
| |
| synonyms = *''Sus domestica'' | ||
*''Sus scrofa domesticus'' <small>], ]</small> | |||
| species = '']'' | |||
| subspecies = '''''S. s. domestica''''' | |||
| trinomial = ''Sus scrofa domestica'' | |||
| trinomial_authority = ], 1758 | |||
| synonyms =<center>'''''Sus domestica''''' | |||
}} | }} | ||
The '''pig''' ('''''Sus domesticus'''''), also called '''swine''' ({{plural form}}: swine) or '''hog''', is an ], ], ]. It is named the '''domestic pig''' when distinguishing it from other members of the genus '']''. It is considered a ] of ''Sus scrofa'' (the ] or Eurasian boar) by some authorities, but as a distinct ] by others. Pigs were domesticated in the ], both in East Asia and in the Near East. When domesticated pigs arrived in Europe, they extensively interbred with wild boar but retained their domesticated features. | |||
The '''] ]''' (or in some areas '''hog''') is normally given the scientific name ''Sus scrofa scrofa'', though some taxonomists use the term ''S. domestica'', reserving ''S. scrofa'' for the ].{{Fact|date=September 2007}} | |||
] primarily for meat, called ]. The animal's skin or ] is used for ]. China is the world's largest pork producer, followed by the European Union and then the United States. Around 1.5 billion pigs are raised each year, producing some 120 million tonnes of meat, often cured as ]. Some are kept as ]s. | |||
The Domestic Pigs are believed to have been ] from wild boar as early as 7000 BC in the ] and, separately, in China<ref> </ref>. DNA evidence from sub-fossil remains of teeth and jawbones of Neolithic pigs in Europe shows that the first domestic pigs there had been brought from the Near East. It appears that this stimulated the domestication of European wild boar, effectively forming a third domestication event – the Near Eastern genes later died out in European pigs, and domesticated European pigs were then exported in turn to the ancient Near East<ref> 4 September 2007. The report concerns an article in the journal ''PNAS''</ref>. | |||
Pigs have featured ] since Neolithic times, appearing in art and literature for children and adults, and celebrated in cities such as ] for their meat products. | |||
The adaptable nature and ] diet of the wild boar allowed early humans to domesticate it much earlier than many other forms of livestock, such as ]. {{Fact|date=September 2007}}. ] were mostly used for food, but early civilizations also used the pigs' hides for shields, bones for tools and weapons, and bristles for brushes.<ref>http://www.flossybrush.ca/oral.html Oral Care. </ref> Pigs were brought to southeastern ] from ] by ] and other early ] ]. Escaped pigs became ] and were used by ] as food.<ref>http://www.cambridge.org/us/books/kiple/hogs.htm II.G.13. - Hogs.</ref> | |||
== Description == | |||
The distinction between wild and domestic animals is slight. Most domestic pigs usually have rather sparse hair covering on their skin, but the ] breeds are known (] pig), and some were popular in the past. <ref>http://www.malverngazette.co.uk/mostpopular.var.1476623.mostviewed.royal_visit_delights_at_the_three_counties_show.php Royal visit delights at the Three Counties Show.</ref> Escaped domestic pigs have become ] in many parts of the world (for example, ]) and have caused substantial environmental damage. <ref>http://query.nytimes.com/gst/fullpage.html?res=9E0CEED6103CF93AA15751C1A964958260&sec=&spon=&pagewanted=all Alien Species Threaten Hawaii's Environment. </ref> <ref>http://www.nzetc.org/tm/scholarly/tei-Bio27Tuat02-t1-body-d1.html Introduced Birds and Mammals in New Zealand and Their Effect on the Environment.</ref> | |||
The pig has a large head, with a long snout strengthened by a special prenasal bone and a disk of ] at the tip.<ref name="ADW Sus scrofa">{{cite web |title=Sus scrofa (wild boar) |url=http://animaldiversity.ummz.umich.edu/site/accounts/information/Sus_scrofa.html |website=Animal Diversity Web}}</ref> The snout is used to dig into the soil to find food and is an acute sense organ. The ] of adult pigs is {{DentalFormula |upper=3.1.4.3 |lower=3.1.4.3}}, giving a total of 44 ]. The rear teeth are adapted for crushing. In males, the canine teeth can form ]s, which grow continuously and are sharpened by grinding against each other.<ref name="ADW Sus scrofa"/> There are four ]ed toes on each foot; the two larger central toes bear most of the weight, while the outer two are also used in soft ground.<ref>{{cite web |last=Lockhart |first=Kim |title=American Wild Game / Feral Pigs / Hogs / Pigs / Wild Boar |url=http://www.gunnersden.com/index.htm.shooting-hunting-hogs.html |url-status=dead |archive-url=https://web.archive.org/web/20180823025758/http://gunnersden.com/index.htm.shooting-hunting-hogs.html |archive-date=23 August 2018 |access-date=15 August 2012 |website=gunnersden.com}}</ref> Most pigs have rather sparsely ]d hair on their skin, though there are some ]ly-coated breeds such as the ].<ref>{{cite web |title=Royal visit delights at the Three Counties Show |url=http://www.malverngazette.co.uk/mostpopular.var.1476623.mostviewed.royal_visit_delights_at_the_three_counties_show.php |website=] |date=15 June 2007 }}</ref> Adult pigs generally weigh between {{cvt|140|and|300|kg}}, though some breeds can exceed this range. Exceptionally, a pig called Big Bill weighed {{cvt|1157|kg}} and had a shoulder height of {{cvt|1.5|m}}.<ref name="Bradford Dutfield 2018">{{cite web |last1=Bradford |first1=Alina |last2=Dutfield |first2=Scott |title=Pigs, Hogs & Boars: Facts About Swine |url=https://www.livescience.com/50623-pigs-facts.html |website=LiveScience |access-date=4 March 2024 |date=5 October 2018}}</ref> | |||
==Farming== | |||
].]] | |||
The domestic pig is farmed for its meat called ], which is obtained by ]. Products made of pork include ], ] and ]. The head of a pig can be used to make ]. ], ], and other ] from pigs are also widely used for food. In some ]s, such as ] and ], there are ]. | |||
Pigs possess both ] and ]s, although the latter are limited to the snout.<ref name="Sumena 2010">{{cite journal |last1=Sumena |first1=K. B. |last2=Lucy |first2=K. M. |last3=Chungath |first3=J.J. |last4=Ashok |first4=N. |last5=Harshan |first5=K. R. |year=2010 |title=Regional histology of the subcutaneous tissue and the sweat glands of large white Yorkshire pigs |journal=Tamil Nadu Journal of Veterinary and Animal Sciences |volume=6 |issue=3 |pages=128–135}}</ref> Pigs, like other "hairless" mammals such as elephants, do not use thermal sweat glands in cooling.<ref name="Folk and Semken, (1991)">{{cite journal |last1=Folk |first1=G.E. |last2=Semken |first2=H.A. |year=1991 |title=The evolution of sweat glands |journal=International Journal of Biometeorology |volume=35 |issue=3 |pages=180–186 |bibcode=1991IJBm...35..180F |doi=10.1007/bf01049065 |pmid=1778649 |s2cid=28234765}}</ref> Pigs are less able than many other mammals to dissipate heat from wet ]s in the mouth by panting. Their ] zone is {{cvt|16-22|C|F}}.<ref>{{cite web |date=22 April 2008 |title=Sweat like a pig? |url=http://www.abc.net.au/science/articles/2008/04/22/2223974.htm |publisher=Australian Broadcasting Corporation}}</ref> At higher temperatures, pigs lose heat by ] in mud or water via evaporative cooling, although it has been suggested that wallowing may serve other functions, such as protection from sunburn, ] control, and scent-marking.<ref name="Bracke 2011">{{cite journal |last=Bracke |first=M. B. M. |year=2011 |title=Review of wallowing in pigs: Description of the behaviour and its motivational basis |journal=Applied Animal Behaviour Science |volume=132 |issue=1 |pages=1–13 |doi=10.1016/j.applanim.2011.01.002}}</ref> Pigs are among four mammalian species with mutations in the ] that protect against ]. ]s, ]s, ]s, and pigs all have different modifications to the receptor pocket which prevents ] from binding.<ref>{{cite journal |last1=Drabeck |first1=D. H. |last2=Dean |first2=A. M. |last3=Jansa |first3=S.A. |date=1 June 2015 |title=Why the honey badger don't care: Convergent evolution of venom-targeted nicotinic acetylcholine receptors in mammals that survive venomous snake bites |journal=Toxicon |volume=99 |pages=68–72 |doi=10.1016/j.toxicon.2015.03.007 |pmid=25796346|bibcode=2015Txcn...99...68D }}</ref> Pigs have small lungs for their body size, and are thus more susceptible than other domesticated animals to fatal ] and ].<ref>{{cite web |title=Pros and Cons of Potbellied Pigs |url=http://www.pigs.org/article.asp?article_id=3 |url-status=dead |archive-url=https://web.archive.org/web/20140317221208/http://www.pigs.org/article.asp?article_id=3 |archive-date=17 March 2014 |access-date=25 November 2017}}</ref> The ] of the pig has been sequenced; it contains about 22,342 ]-coding ]s.<ref>{{cite journal |last1=Li |first1=Mingzhou |last2=Chen |first2=Lei |last3=Tian |first3=Shilin |last4=Lin |first4=Yu |last5=Tang |first5=Qianzi |last6=Zhou |first6=Xuming |last7=Li |first7=Diyan |last8=Yeung |first8=Carol K. L. |last9=Che |first9=Tiandong |last10=Jin |first10=Long |last11=Fu |first11=Yuhua |date=2017-05-01 |title=Comprehensive variation discovery and recovery of missing sequence in the pig genome using multiple de novo assemblies |url=https://genome.cshlp.org/content/27/5/865 |journal=Genome Research |volume=27 |issue=5 |pages=865–874 |doi=10.1101/gr.207456.116 |pmc=5411780 |pmid=27646534}}</ref><ref>{{cite journal |year=2020 |doi=10.1093/gigascience/giaa051 |pmc=7448572 |pmid=32543654 |last1=Warr |first1=A. |last2=Affara |first2=N. |last3=Aken |first3=B. |last4=Beiki |first4=H. |last5=Bickhart |first5=D. M. |last6=Billis |first6=K. |last7=Chow |first7=W. |last8=Eory |first8=L. |last9=Finlayson |first9=H. A. |last10=Flicek |first10=P. |last11=Girón |first11=C. G. |last12=Griffin |first12=D. K. |display-authors=6 |title=An improved pig reference genome sequence to enable pig genetics and genomics research |journal=GigaScience |volume=9 |issue=6 |pages=giaa051}}</ref><ref>{{cite journal |last1=Karlsson |first1=Max |last2=Sjöstedt |first2=Evelina |last3=Oksvold |first3=Per |last4=Sivertsson |first4=Åsa |last5=Huang |first5=Jinrong |last6=Álvez |first6=María Bueno |last7=Arif |first7=Muhammad |last8=Li |first8=Xiangyu |last9=Lin |first9=Lin |last10=Yu |first10=Jiaying |last11=Ma |first11=Tao |display-authors=6 |date=25 January 2022 |title=Genome-wide annotation of protein-coding genes in pig |journal=BMC Biology |volume=20 |issue=1 |page=25 |doi=10.1186/s12915-022-01229-y |pmid=35073880 |pmc=8788080 |doi-access=free }}</ref> | |||
In ] nations, domestic pigs farming has shifted away from the ] to large-scale ] where meat can be mass-produced. This has resulted in lower production costs, but more significant ] concerns. | |||
<gallery class=center mode=nolines widths=220 heights=180> | |||
] | |||
File:Suíno alta (cropped).jpg|Skeleton | |||
A young female pig who has borne fewer than two litters is known as a ''gilt'' (In some usages, a gilt must never have been pregnant<ref name='ucdavis_swineguide'> from ]</ref>), a female who has borne more than two litters is a ''sow'', an intact male is a ''boar'', a ] male is a ''barrow'', and young pigs of both sexes are ''piglets'' or ''shoats'' (or in areas where the species is called ''hog'', just ''pigs''), or in Ireland ''banbhs'' (pronounced {{IPA|}}; from the ] ''banbh''). Pigs of various ages and types sometimes also have particular names, such as ''porker'' or ''baconer'', and the word ''swine'' (plural also ''swine'') can be used to refer to all pigs. | |||
File:Domestic pig skull (cropped).jpeg|Skull | |||
Female pigs used for reproduction are called ''breeding sow'', while the males are called ''breeding boar''. | |||
File:Pig hand skeleton (cropped).jpg|Bones of the foot | |||
</gallery> | |||
== Evolution == | |||
Giving birth in pigs is called ''farrowing'', and a pregnant gilt or sow is said to be ''in pig''. A small building in which a pig is kept is called a ''sty'', and a larger one a ''pig-shed'', and a low outdoor shelter for a pig (also other smaller animals such as chickens or rabbits) is an ''ark''. The feet of pigs are called ''trotters'' (they have four hoofed toes, walking mainly on the larger central two). A group of pigs is a ''herd'', or if a smaller group foraging in woodland, a ''sounder''. | |||
=== Phylogeny === | |||
In ]s, and often in ]s, the domestic pig is raised outdoors in yards or fields. In some cases pigs are allowed to ] in woods, where they are watched by ]s, the equivalent of ]s for pigs. | |||
Domestic pigs are related to other pig species as shown in the ], based on phylogenetic analysis using ].<ref>{{cite journal |last1=Wu |first1=Gui-Sheng |last2=Pang |first2=Jun-feng |last3=Zhang |first3=Ya-Ping |title=Molecular phylogeny and phylogeography of Suidae |journal=Zoological Research |volume=27 |issue=2 |year=2006 |pages=197–201 |url=https://www.researchgate.net/publication/43563862}}</ref> | |||
==As pets== | |||
]]] | |||
Pigs are known to be ] animals and have been found to be more trainable than ]s or ]s. <ref>http://www.rps.psu.edu/probing/pigs.html Are pigs smarter than dogs?</ref> Asian ]s, a small type of domestic pig, have made popular house ]s in the ] beginning in the latter half of the 20th century. Regular domestic farmyard pigs have also been known to be kept indoors, but due to their large size and destructive tendencies, they typically need to be moved into an outdoor pen as they grow older. Most pigs have a fear of being picked up, but will usually calm down once placed back on the floor. Pigs are rarely used as ]. An exception is the use of ] – ordinary pigs trained to find ]. | |||
{{clade | |||
==Breeds of pigs== | |||
|label1=] | |||
{{For|a list of breeds of domestic swine|List of domestic pig breeds}} | |||
|1={{clade | |||
] | |||
|sublabel1=''Indonesia'' | |||
Pigs are exhibited at ]s, judged either as ] compared to the standard features of each breed, or in commercial classes where the animals are judged primarily on their suitability for slaughter to provide premium meat. | |||
|1=] ] | |||
|2={{clade | |||
|sublabel1=''Africa'' | |||
|1={{clade | |||
|1=] ] | |||
|2=]s ] | |||
}} | |||
|sublabel2=''Southeast Asia'' | |||
|2={{clade | |||
|1=] ] | |||
|2={{clade | |||
|1=] ] | |||
|2={{clade | |||
|1={{clade | |||
|1=] ] | |||
|2=] ] | |||
}} | |||
|sublabel2=''Eurasia'' | |||
|2={{clade | |||
|label1=] | |||
|1=] | |||
|label2= '''Domestic pig''' | |||
|2={{clade | |||
|1=East Asian pigs ] | |||
|2=European pigs<!--inc wild boar--> ] | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
}} | |||
=== Taxonomy === | |||
{{Agricultural production box | |||
|year=2005 | |||
|animal=pig | |||
|country1={{CHN}} | |||
|amount1=488.8 | |||
|country2={{USA}} | |||
|amount2=60.4 | |||
|country3={{BRA}} | |||
|amount3=33.2 | |||
|country4={{VNM}} | |||
|amount4=27.0 | |||
|country5= {{GER}} | |||
|amount5=26.9 | |||
|country6= {{ESP}} | |||
|amount6=25.3 | |||
|country7= {{POL}} | |||
|amount7=18.0 | |||
|country8= {{FRA}} | |||
|amount8=15.0 | |||
|country9= {{CAN}} | |||
|amount9=14.7 | |||
|country10= {{MEX}} | |||
|amount10=14.6 | |||
|world=960.8 }} | |||
The pig is most often considered to be a ] of the ], which was given the name ''Sus scrofa'' by ] in 1758; following from this, the formal name of the pig is ''Sus scrofa domesticus''.<ref>{{cite web |title=Taxonomy Browser |url=https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?name=Sus+scrofa+domestica |website=ncbi.nlm.nih.gov}}</ref><ref name="gentry">{{cite journal |last1=Gentry |first1=Anthea |last2=Clutton-Brock |first2=Juliet |last3=Colin P. Groves |year=2004 |title=The naming of wild animal species and their domestic derivatives |url=http://arts.anu.edu.au/grovco/J%20Arch%20Sci.pdf |url-status=dead |journal=] |volume=31 |issue=5 |pages=645–651 |doi=10.1016/j.jas.2003.10.006 |bibcode=2004JArSc..31..645G |archive-url=https://web.archive.org/web/20110408044739/http://arts.anu.edu.au/grovco/J%20Arch%20Sci.pdf |archive-date=8 April 2011}}</ref> However, in 1777, ] classified the pig as a separate ] from the wild boar. He gave it the name ''Sus domesticus'', still used by some taxonomists.<!--<ref>Corbet, G. B.; Hill, J. E., 1992. The mammals of the Indomalayan | |||
===Rare pig breeds=== | |||
Region: a systematic review. Natural History Museum Publications, Oxford University Press. Pp. i-viii, 1-488., cited in {{MSW3|id=14200054}}</ref>--><ref name="gentry2">{{cite journal |last1=Gentry |first1=Anthea |last2=Clutton-Brock |first2=Juliet |last3=Groves |first3=Colin P. |year=1996 |title=Proposed conservation of usage of 15 mammal specific names based on wild species which are antedated by or contemporary with those based on domestic animals |journal=] |volume=53 |pages=28–37 |doi=10.5962/bhl.part.14102 |doi-access=free}}</ref> The ] considers it a separate species.<ref>{{cite web |title=Explore the Database |url=https://www.mammaldiversity.org/explore.html#species-id=1006374 |access-date=21 August 2021 |website=www.mammaldiversity.org}}</ref> | |||
According to the ] <ref>. Retrieved 18 April 2008.</ref>, seven breeds of swine in the U.S. are critically rare (having a global population of fewer than 2000). Outside the U.S., the ] (New Zealand) and ] (Danube area) are known to be critically rare. Rare U.S. breeds include: | |||
* ] | |||
* ] | |||
* ] | |||
=== Domestication in the Neolithic === | |||
==See also== | |||
{{Commons|Sus domesticus}} | |||
*] | |||
*] | |||
*] | |||
*] | |||
*] | |||
*] | |||
{{further|Domestication}} | |||
==Notes== | |||
{{reflist}} | |||
] | |||
==References== | |||
* {{IUCN2006|assessors=Pigs & Peccaries Specialist Group|year=1996|id=41775|title=Sus scrofa|downloaded=12 May 2006}} | |||
Archaeological evidence shows that pigs were ] from wild boar in the ] in or around the ] Basin,<ref>{{cite journal |last1=Ottoni |first1=C |last2=Flink |first2=LG. |last3=Evin |first3=A. |last4=Geörg |first4=C. |last5=De Cupere |first5=B. |last6=Van Neer |first6=W. |last7=Bartosiewicz |first7=L. |last8=Linderholm |first8=A. |last9=Barnett |first9=R. |last10=Peters |first10=J |last11=Decorte |first11=R. |display-authors=6 |year=2013 |title=Pig Domestication and Human-Mediated Dispersal in Western Eurasia Revealed through Ancient DNA and Geometric Morphometrics |journal=Molecular Biology and Evolution |volume=30 |issue=4 |pages=824–32 |doi=10.1093/molbev/mss261 |pmc=3603306 |pmid=23180578 |quote=our data suggest a narrative that begins with the domestication of pigs in Southwest Asia, at Upper Tigris sites including Çayönü Tepesi (Ervynck et al. 2001) and possibly Upper Euphrates sites including Cafer Höyük (Helmer 2008) and Nevalı Çori (Peters et al. 2005) }}</ref> being managed in a semi-wild state much as they are managed by some modern New Guineans.<ref>{{cite journal |last1=Rosenberg |first1=M. |last2=Nesbitt |first2=R. |last3=Redding |first3=R. W. |last4=Peasnall |first4=BL |year=1998 |title=Hallan Çemi, pig husbandry, and post-Pleistocene adaptations along the Taurus-Zagros Arc (Turkey) |journal=Paléorient |volume=24 |issue=1 |pages=25–41 |doi=10.3406/paleo.1998.4667 |s2cid=85302206 |url=http://www.persee.fr/web/revues/home/prescript/article/paleo_0153-9345_1998_num_24_1_4667}}</ref> There were pigs in ] more than 11,400 years ago, introduced from the mainland, implying domestication in the adjacent mainland by then.<ref>{{cite journal |last1=Vigne |first1=J. D. |last2=Zazzo |first2=A. |last3=Saliège |first3=J.F. |last4=Poplin |first4=F. |last5=Guilaine |first5=J. |last6=Simmons |first6=A. |year=2009 |title=Pre-Neolithic wild boar management and introduction to Cyprus more than 11,400 years ago |journal=] |volume=106 |issue=38 |pages=16135–16138 |bibcode=2009PNAS..10616135V |doi=10.1073/pnas.0905015106 |pmc=2752532 |pmid=19706455 |doi-access=free}}</ref> Pigs were separately domesticated in China, starting some 8,000 years ago.<ref>{{cite journal |last1=Lander |first1=Brian |last2=Schneider |first2=Mindi |last3=Brunson |first3=Katherine |title=A History of Pigs in China: From Curious Omnivores to Industrial Pork |journal=Journal of Asian Studies |date=2019 |volume=79 |issue=4 |pages=865–889 |doi=10.1017/S0021911820000054 |doi-access=|s2cid=225700922 }}</ref><ref name="Giuffra">{{cite journal |last1=Giuffra |first1=E. |last2=Kijas |first2=J. M. |last3=Amarger |first3=V. |last4=Carlborg |first4=O. |last5=Jeon |first5=J. T. |last6=Andersson |first6=L. |year=2000 |title=The origin of the domestic pig: independent domestication and subsequent introgression |journal=Genetics |volume=154 |issue=4 |pages=1785–1791 |doi=10.1093/genetics/154.4.1785 |pmc=1461048 |pmid=10747069}}</ref><ref>{{cite speech |date=3 July 2012 |title=History of domestication |author1=Jean-Denis Vigne |author2=Anne Tresset |author3=Jean-Pierre Digard |url=http://www2.mnhn.fr/archeozoo-archeobota/IMG/pdf/Cucchi_ChinaPig_JAS_Online.pdf}}</ref> In the Near East, pig husbandry spread for the next few millennia. It reduced gradually during the ], as rural populations instead focused on commodity-producing livestock, but it was sustained in cities.<ref>{{cite web |last=Price |first=Max |date=March 2020 |title=The Genesis of the Near Eastern Pig |url=https://www.asor.org/anetoday/2020/03/near-eastern-pig/ |access-date=8 August 2021 |website=American Society of Overseas Research |archive-date=10 February 2022 |archive-url=https://web.archive.org/web/20220210052924/https://www.asor.org/anetoday/2020/03/near-eastern-pig/ |url-status=dead }}</ref> | |||
* | |||
* | |||
Domestication did not involve reproductive isolation with population bottlenecks. Western Asian pigs were introduced into Europe, where they crossed with wild boar. There appears to have been interbreeding with a now extinct ] of wild pigs during the ]. The genomes of domestic pigs show strong selection for genes affecting behavior and morphology. Human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars and created ] in the genome.<ref name="frantz2015">{{cite journal |last=Frantz |first=L |year=2015 |title=Evidence of long-term gene flow and selection during domestication from analyses of Eurasian wild and domestic pig genomes |journal=] |volume=47 |issue=10 |pages=1141–1148 |doi=10.1038/ng.3394 |pmid=26323058 |s2cid=205350534}}</ref><ref name="pennisi2015">{{cite journal |last=Pennisi |first=E. |author-link=Elizabeth Pennisi |year=2015 |title=The taming of the pig took some wild turns |journal=Science |doi=10.1126/science.aad1692}}</ref> Pigs arrived in Europe from the Near East at least 8,500 years ago. Over the next 3,000 years they interbred with European wild boar until their genome showed less than 5% Near Eastern ancestry, yet retained their domesticated features.<ref>{{cite journal |last1=Frantz |first1=Laurent A. F. |last2=Haile |first2=James |last3=Lin |first3=Audrey T. |last4=Scheu |first4=Amelie |last5=Geörg |first5=Christina |last6=Benecke |first6=Norbert |last7=Alexander |first7=Michelle |last8=Linderholm |first8=Anna |last9=Mullin |first9=Victoria E. |last10=Daly |first10=Kevin G. |last11=Battista |first11=Vincent M. |last12=Price |first12=Max |display-authors=6 |year=2019 |title=Ancient pigs reveal a near-complete genomic turnover following their introduction to Europe |journal=] |volume=116 |issue=35 |pages=17231–17238 |doi=10.1073/pnas.1901169116 |pmc=6717267 |pmid=31405970 |doi-access=free |bibcode=2019PNAS..11617231F }}</ref> | |||
* | |||
] evidence from subfossil remains of teeth and jawbones of Neolithic pigs shows that the first domestic pigs in Europe were brought from the Near East. This stimulated the domestication of local European wild boar, resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. More recently there have been complex exchanges, with European domesticated lines being exported, in turn, to the ancient Near East.<ref> 4 September 2007. The report concerns an article in the journal ''PNAS''</ref><ref>{{cite journal |last1=Larson |first1=G. |last2=Albarella |first2=U. |last3=Dobney |first3=K. |last4=Rowley-Conwy |first4=P. |last5=Schibler |first5=J. |last6=Tresset |first6=A. |last7=Vigne |first7=J.D. |last8=Edwards |first8=C.J. |last9=Schlumbaum |first9=A. |last10=Dinu |first10=A. |last11=Balacsescu |first11=A. |last12=Dolman |first12=G. |display-authors=6 |year=2007 |title=Ancient DNA, pig domestication, and the spread of the Neolithic into Europe |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=104 |issue=39 |pages=15276–15281 |bibcode=2007PNAS..10415276L |doi=10.1073/pnas.0703411104 |pmc=1976408 |pmid=17855556 |doi-access=free |url=https://pure.uva.nl/ws/files/4125719/55285_284894.pdf}}</ref> Historical records indicate that Asian pigs were again introduced into Europe during the 18th and early 19th centuries.<ref name="Giuffra"/> | |||
== History == | |||
=== Columbian Exchange === | |||
Among the animals that the Spanish introduced to the ] in the 16th century ], pigs were the most successful in adapting to local conditions. The pigs benefited from abundant ] and ] exposed by the large ]s of the archipelago.<ref name="Torrejonetal2004">{{cite journal |last1=Torrejón |first1=Fernando |last2=Cisternas |first2=Marco |last3=Araneda |first3=Alberto |year=2004 |title=Efectos ambientales de la colonización española desde el río Maullín al archipiélago de Chiloé, sur de Chile |trans-title=Environmental effects of the spanish colonization from de Maullín river to the Chiloé archipelago, southern Chile |journal=] |language=es |volume=77 |issue=4 |pages=661–677 |doi=10.4067/s0716-078x2004000400009 |doi-access=free}}</ref> Pigs were brought to southeastern North America from Europe by ] and other early Spanish ]s. Escaped pigs became ].<ref> {{webarchive|url=https://web.archive.org/web/20071220183225/http://www.cambridge.org/us/books/kiple/hogs.htm |date=20 December 2007 }}</ref> | |||
=== Feral pigs === | |||
{{main|Feral pig}} | |||
] | |||
Pigs have escaped from farms and gone ] in many parts of the world. Feral pigs in the southeastern United States have migrated north to the ], where many state agencies have programs to remove them.<ref>{{cite web |title=Feral Hogs in Missouri |publisher=Missouri Department of Conservation |url=https://mdc.mo.gov/wildlife/nuisance-problem-species/invasive-species/feral-hogs-missouri |access-date=7 March 2017 |archive-date=8 March 2017 |archive-url=https://web.archive.org/web/20170308143218/https://mdc.mo.gov/wildlife/nuisance-problem-species/invasive-species/feral-hogs-missouri |url-status=dead}}</ref><ref>{{cite web |title=Feral Hog Hunting Regulations |url=http://www.agfc.com/hunting/Pages/HuntingRegulationsFeralHog.aspx |archive-url=https://web.archive.org/web/20170222171722/http://www.agfc.com/hunting/Pages/HuntingRegulationsFeralHog.aspx |archive-date=22 February 2017 |url-status=dead |access-date=7 March 2017 |website=agfc.com}}</ref><ref>{{cite web |title=Feral Hog Management |website=Georgia DNR – Wildlife Resources Division |url=http://georgiawildlife.com/node/3002 |url-status=dead |archive-url=https://web.archive.org/web/20170308133934/http://georgiawildlife.com/node/3002 |archive-date=8 March 2017 |access-date=8 March 2017}}</ref> Feral pigs in New Zealand and northern ] have caused substantial environmental damage.<ref>{{cite news |last=Yoon |first=Carol Kaesuk |date=2 December 1992 |title=Alien Species Threaten Hawaii's Environment |work=The New York Times |url=https://query.nytimes.com/gst/fullpage.html?res=9E0CEED6103CF93AA15751C1A964958260&sec=&spon=&pagewanted=all}}</ref><ref>{{cite web |title=Introduced Birds and Mammals in New Zealand and Their Effect on the Environment – NZETC |url=https://nzetc.victoria.ac.nz/tm/scholarly/tei-Bio27Tuat02-t1-body-d1.html |website=nzetc.org}}</ref> Feral hybrids of the European wild boar with the domestic pig are disruptive to both environment and agriculture, as they destroy crops, spread animal diseases including ], and consume wildlife such as juvenile seabirds and young ]s.<ref>{{cite news |date=21 November 2004 |title=World's 100 most destructive species named |work=] |url=https://www.independent.co.uk/environment/worlds-100-most-destructive-species-named-534039.html |access-date=7 March 2017}}</ref> Feral pig damage is especially an issue in southeastern South America.<ref>{{cite web |url=http://g1.globo.com/sp/bauru-marilia/noticia/2013/04/autorizacao-para-abate-do-javaporco-tranquiliza-produtores-em-assis-sp.html |title=Autorização para abate do javaporco tranquiliza produtores em Assis, SP |author1=Bauru |author2=Marília |date=12 April 2013 |website=Bauru e Marília}}</ref><ref>{{cite web |title=IBAMA authorizes capture and slaughter of 'javaporcos' – Folha do Sul Gaúcho |url=http://www.jornalfolhadosul.com.br/noticia/2013/02/07/ibama-autoriza-captura-e-abate-de-javaporcos |archiveurl=https://web.archive.org/web/20170703010021/http://www.jornalfolhadosul.com.br/noticia/2013/02/07/ibama-autoriza-captura-e-abate-de-javaporcos|url-status=dead |archivedate=3 July 2017}}</ref> | |||
== Reproduction == | |||
=== Physiology === | |||
{{multiple image|perrow=1|image1=The_breeding_of_animals_(1917)_(20385751936).jpg|caption1=] of the boar|image2=The_breeding_of_animals_(1917)_(20412003625).jpg|caption2=Reproductive system of the sow}} | |||
Female pigs reach sexual maturity at 3–12 months of age and come into ] every 18–24 days if they are not successfully bred. The variation in ovulation rate can be attributed to intrinsic factors such as age and genotype, as well as extrinsic factors like nutrition, environment, and the supplementation of exogenous hormones. The gestation period averages 112–120 days.<ref>{{cite web |date=16 May 2012 |title=Feral Hog Reproductive Biology |url=http://articles.extension.org/pages/63668/feral-hog-reproductive-biology |url-status=dead |archive-url=https://web.archive.org/web/20151122022154/http://articles.extension.org/pages/63668/feral-hog-reproductive-biology#.VlEm2HnP1qY |archive-date=22 November 2015}}</ref> | |||
] | |||
] lasts two to three days, and the female's displayed receptiveness to mate is known as standing heat. Standing heat is a reflexive response that is stimulated when the female is in contact with the saliva of a sexually mature boar. ] is one of the pheromones produced in the submaxillary salivary glands of boars that trigger the female's response.<ref>{{cite web |title=G2312 Artificial Insemination in Swine: Breeding the Female |publisher=University of Missouri Extension |url=http://extension.missouri.edu/publications/DisplayPub.aspx?P=G2312 |access-date=7 March 2017 |archive-date=8 March 2017 |archive-url=https://web.archive.org/web/20170308145924/http://extension.missouri.edu/publications/DisplayPub.aspx?P=G2312 |url-status=dead }}</ref> The female cervix contains a series of five interdigitating pads, or folds, that hold the boar's corkscrew-shaped penis during copulation.<ref>{{cite web |title=The Female – Swine Reproduction |url=http://livestocktrail.illinois.edu/swinerepronet/paperDisplay.cfm?ContentID=6274 |access-date=7 March 2017 |website=livestocktrail.illinois.edu |archive-date=10 February 2022 |archive-url=https://web.archive.org/web/20220210052921/http://livestocktrail.illinois.edu/swinerepronet/paperDisplay.cfm?ContentID=6274 |url-status=dead }}</ref> Females have ]es and two ]es must be present in both uterine horns to enable pregnancy to proceed.<ref>{{cite journal |last1=Bazer |first1=F. W. |last2=Vallet |first2=J. L. |last3=Roberts |first3=R. M. |last4=Sharp |first4=D. D. |last5=Thatcher |first5=W. W. |year=1986 |title=Role of conceptus secretory products in establishment of pregnancy |journal=J. Reprod. Fertil. |volume=76 |issue=2 |pages=841–850 |doi=10.1530/jrf.0.0760841 |pmid=3517318 |doi-access=free}}</ref> The mother's body recognises that it is pregnant on days 11 to 12 of pregnancy, and is marked by the ]'s producing the sex hormone ].<ref>{{cite journal |last1=Bazer |first1=Fuller W. |last2=Song |first2=Gwonhwa |last3=Kim |first3=Jinyoung |last4=Dunlap |first4=Kathrin A. |last5=Satterfield |first5=Michael Carey |last6=Johnson |first6=Gregory A. |last7=Burghardt |first7=Robert C. |last8=Wu |first8=Guoyao |author-link8=Guoyao Wu |date=1 January 2012 |title=Uterine biology in pigs and sheep |journal=Journal of Animal Science and Biotechnology |volume=3 |issue=1 |page=23 |doi=10.1186/2049-1891-3-23 |pmc=3436697 |pmid=22958877 |doi-access=free }}</ref> To sustain the pregnancy, the embryo signals to the corpus luteum with the hormones ] and ].<ref name="Ziecik, A. J. 2018">{{cite journal |last=Ziecik |first=A. J. |display-authors=etal |year=2018 |title=Regulation of the porcine corpus luteum during pregnancy |journal=Reproduction |volume=156 |issue=3 |pages=R57–R67 |doi=10.1530/rep-17-0662 |pmid=29794023 |doi-access=free}}</ref> This signaling acts on both the endometrium and luteal tissue to prevent the regression of the corpus luteum by activation of genes that are responsible for corpus luteum maintenance.<ref>{{cite journal |last=Waclawik |first=A. |display-authors=etal |year=2017 |title=Embryo-maternal dialogue during pregnancy establishment and implantation in the pig |journal=Molecular Reproduction and Development |volume=84 |issue=9 |pages=842–855 |doi=10.1002/mrd.22835 |pmid=28628266 |doi-access=free}}</ref> During mid to late pregnancy, the corpus luteum relies primarily on ] for maintenance until birth.<ref name="Ziecik, A. J. 2018"/><!--<ref>{{cite book |last=Farmer |first=Chantal |title=The gestating and lactating sow |publisher=Wageningen Academic Publishers |year=2015 |isbn=9789086868032 |location=The Netherlands |oclc=899008362}}</ref>--> | |||
Archeological evidence indicates that medieval European pigs farrowed, or bore a litter of piglets, once per year.<ref>{{cite journal |last1=Ervynck |first1=Anton |last2=Dobney |first2=Keith |year=2002 |title=A Pig for all Seasons? Approaches to the Assessment of Second Farrowing in Archaeological Pig Populations |journal=Archaeofauna |issue=11 |pages=7–22 |doi=10.15366/archaeofauna2002.11.001 |url=https://revistas.uam.es/archaeofauna/article/download/8489/8868}}</ref> By the nineteenth century, European piglets routinely double-farrowed, or bore two litters of piglets per year. It is unclear when this shift occurred.<ref name="Bintliff Earle Peebles 2008 p. 305">{{cite book |last1=Bintliff |first1=J. |last2=Earle |first2=T. |last3=Peebles |first3=C. |title=A Companion to Archaeology |publisher=Wiley |year=2008 |isbn=978-0-470-99860-1 |url=https://books.google.com/books?id=7w1kGdMiwekC&pg=PA305 |page=305}}</ref> Pigs have a ] of about 27 years.<ref name="pmid32361879">{{cite journal |last1=Hoffman |first1=J. |last2=Valencak |first2=T. G. |title=A short life on the farm: aging and longevity in agricultural, large-bodied mammals |journal=] |volume=42 |issue=3 |pages=909–922 |date=2020 |doi=10.1007/s11357-020-00190-4 |pmc=7286991 |pmid=32361879}}</ref> | |||
=== Nest-building === | |||
A characteristic of pigs which they share with carnivores is ]. Sows root in the ground to create depressions the size of their body, and then build nest mounds, using twigs and leaves, softer in the middle, in which to give birth. When the mound reaches the desired height, she places large branches, up to 2 metres in length, on the surface. She enters the mound and roots around to create a depression within the gathered material. She then gives birth in a lying position, unlike other artiodactyls which usually stand while birthing.<ref name="Clutton-Brock 1987"/> | |||
Nest-building occurs during the last 24 hours before the onset of farrowing, and becomes most intense 12 to 6 hours before farrowing.<ref name="Algers 2007"/> The sow separates from the group and seeks a suitable nest site with well-drained soil and shelter from rain and wind. This provides the offspring with shelter, comfort, and thermoregulation. The nest provides protection against weather and predators, while keeping the piglets close to the sow and away from the rest of the herd. This ensures they do not get trampled on, and prevents other piglets from stealing milk from the sow.<ref name="Wischner 2009">{{cite journal |last1=Wischner |first1=D. |last2=Kemper |first2=N. |last3=Krieter |first3=J. |year=2009 |title=Nest-building behaviour in sows and consequences for pig husbandry |journal=Livestock Science |volume=124 |issue=1 |pages=1–8 |doi=10.1016/j.livsci.2009.01.015}}</ref> The onset of nest-building is triggered by a rise in prolactin level, caused by a decrease in progesterone and an increase in prostaglandin; the gathering of nest material seems to be regulated more by external stimuli such as temperature.<ref name="Algers 2007"/> | |||
=== Nursing and suckling === | |||
Pigs have complex nursing and suckling behaviour.<ref name="Fraser, (1980)">{{cite journal |last=Fraser |first=D. |year=1980 |title=A review of the behavioural mechanisms of milk ejection of the domestic pig |url=https://animalstudiesrepository.org/cgi/viewcontent.cgi?article=1001&context=feebeh |journal=Applied Animal Ethology |volume=6 |issue=3 |pages=247–256 |doi=10.1016/0304-3762(80)90026-7}}</ref> Nursing occurs every 50–60 minutes, and the sow requires stimulation from piglets before milk let-down. Sensory inputs (vocalisation, odours from mammary and birth fluids, and hair patterns of the sow) are particularly important immediately post-birth to facilitate teat location by the piglets.<ref name="Rohde Parfet and Gonyou, (1991)">{{cite journal |last1=Rohde Parfet |first1=K.A. |last2=Gonyou |first2=H.W. |year=1991 |title=Attraction of newborn piglets to auditory, visual, olfactory and tactile stimuli |journal=] |volume=69 |issue=1 |pages=125–133 |doi=10.2527/1991.691125x |pmid=2005005 |s2cid=31788525}}</ref> Initially, the piglets compete for position at the udder; then the piglets massage around their respective teats with their snouts, during which time the sow grunts at slow, regular intervals. Each series of grunts varies in frequency, tone and magnitude, indicating the stages of nursing to the piglets.<ref name="Algers, (1993)">{{cite journal |last=Algers |first=B |year=1993 |title=Nursing in pigs: communicating needs and distributing resources |url=https://www.researchgate.net/publication/14973674 |journal=] |volume=71 |issue=10 |pages=2826–2831 |doi=10.2527/1993.71102826x |pmid=8226386}}</ref> | |||
The phase of competition for teats and of nosing the udder lasts for about a minute, ending when milk begins to flow. The piglets then hold the teats in their mouths and suck with slow mouth movements (one per second), and the rate of the sow's grunting increases for approximately 20 seconds. The grunt peak in the third phase of suckling does not coincide with milk ejection, but rather the release of oxytocin from the pituitary into the bloodstream.<ref name="Castren et al., (1989)">{{cite journal |last1=Castren |first1=H. |last2=Algers |first2=B. |last3=Jensen |first3=P. |last4=Saloniemi |first4=H. |year=1989 |title=Suckling behaviour and milk consumption in newborn piglets as a response to sow grunting |journal=Applied Animal Behaviour Science |volume=24 |issue=3 |pages=227–238 |doi=10.1016/0168-1591(89)90069-5}}</ref> Phase four coincides with the period of main milk flow (10–20 seconds) when the piglets suddenly withdraw slightly from the udder and start sucking with rapid mouth movements of about three per second. The sow grunts rapidly, lower in tone and often in quick runs of three or four, during this phase. Finally, the flow stops and so does the grunting of the sow. The piglets may dart from teat to teat and recommence suckling with slow movements, or nosing the udder. Piglets massage and suckle the sow's teats after milk flow ceases as a way of letting the sow know their nutritional status. This helps her to regulate the amount of milk released from that teat in future sucklings. The more intense the post-feed massaging of a teat, the more milk that teat later releases.<ref name="Jensen et al., (1998)">{{cite journal |last1=Jensen |first1=P. |last2=Gustafsson |first2=G. |last3=Augustsson |first3=H. |year=1998 |title=Massaging after milk ejection in domestic pigs – an example of honest begging? |journal=] |volume=55 |issue=4 |pages=779–786 |doi=10.1006/anbe.1997.0651 |pmid=9632466 |s2cid=12493158}}</ref> | |||
<gallery class=center mode=nolines widths=220 heights=220> | |||
File:Pig, Kilcullen.jpg|Sows typically have 12–14 nipples. | |||
File:PigCochon allaitant.jpg|A sow with suckling piglets | |||
</gallery> | |||
=== Teat order === | |||
In pigs, ] hierarchies are formed at an early age. Piglets are precocious, and attempt to suckle soon after being born. The piglets are born with sharp teeth and fight for the ] teats, as these produce more milk. Once established, this teat order remains stable; each piglet tends to feed on a particular teat or group of teats.<ref name="Clutton-Brock 1987"/> Stimulation of the anterior teats appears to be important in causing milk letdown,<ref name="Fraser, (1973)">{{cite journal |last=Fraser |first=D. |year=1973 |title=The nursing and suckling behaviour in pigs. I. The importance of stimulation of the anterior teats |journal=British Veterinary Journal |volume=129 |issue=4 |pages=324–336 |doi=10.1016/s0007-1935(17)36434-5 |pmid=4733757}}</ref> so it might be advantageous to the entire litter to have these teats occupied by healthy piglets. Piglets locate teats by sight and then by olfaction.<ref name="Jeppesen, (1982)">{{cite journal |last=Jeppesen |first=L.E. |year=1982 |title=Teat-order in groups of piglets reared on an artificial sow. II. Maintenance of teat order with some evidence for the use of odour cues |journal=Applied Animal Ethology |volume=8 |issue=4 |pages=347–355 |doi=10.1016/0304-3762(82)90067-0}}</ref> | |||
== Behaviour == | |||
=== Social === | |||
] pigs in a wallow]] | |||
Pig behaviour is intermediate between that of other ] and of ].<ref name="Clutton-Brock 1987">{{cite book |last=Clutton-Brock |first=Juliet |year=1987 |title=A Natural History of Domesticated Mammals |publisher=] |location=Cambridge |pages=73–74}}</ref> Pigs seek out the company of other pigs and often huddle to maintain physical contact, but they do not naturally form large herds. They live in groups of about 8–10 adult sows, some young individuals, and some single males.<ref name="Algers 2007">{{cite journal |last1=Algers |first1=Bo |last2=Uvnäs-Moberg |first2=Kerstin |date=1 June 2007 |title=Maternal behavior in pigs |journal=Hormones and Behavior |series=Reproductive Behavior in Farm and Laboratory Animals: 11th Annual Meeting of the Society for Behavioral Neuroendocrinology |volume=52 |issue=1 |pages=78–85 |doi=10.1016/j.yhbeh.2007.03.022 |pmid=17482189 |s2cid=9742677}}</ref> Pigs confined in a simplified, crowded, or uncomfortable environment may resort to ]; farmers sometimes ] of pigs to prevent the problem, or may enrich the environment with toys or other objects to reduce the risk.<ref name="AHDB">{{cite web |title=Tail docking and tail biting in pigs |url=https://ahdb.org.uk/knowledge-library/tail-docking-and-tail-biting-in-pigs |publisher=Animal Husbandry Development Board |access-date=4 March 2024}}</ref><ref name="NADIS">{{Cite web |url=http://www.nadis.org.uk/bulletins/tail-biting.aspx |title=Pig Health- Tail Biting |website=] |access-date=4 March 2024 |url-status=dead |archive-url=https://web.archive.org/web/20130113093850/http://www.nadis.org.uk/bulletins/tail-biting.aspx |archive-date=13 January 2013}}</ref> | |||
=== Temperature control === | |||
Because of their relative lack of sweat glands, pigs often control their body temperature using behavioural thermoregulation. ], coating the body with mud, is a common behaviour.<ref name="Bracke 2011" /> They do not submerge completely under the mud, but vary the depth and duration of wallowing depending on environmental conditions.<ref name="Bracke 2011"/> Adult pigs start wallowing once the ambient temperature is around {{cvt|17–21|C}}. They cover themselves in mud from head to tail.<ref name="Bracke 2011"/> They may use mud as a sunscreen, or to keep parasites away.<ref name="Bracke 2011"/> Most bristled pigs "blow their coat", meaning that they shed most of the longer, coarser stiff hair once a year, usually in spring or early summer, to prepare for the warmer months ahead.<ref>{{cite web |date=2 April 2016 |title=Blowing Coat – Mini Pig Shedding FAQ |url=http://americanminipigassociation.com/educational/blowing-coat-mini-pig-shedding |website=americanminipigassociation.com}}</ref> | |||
=== Eating, feeding, sleeping === | |||
] | |||
Where pigs are allowed to roam freely, they walk roughly 4 km daily, ] within a home range of around a hectare. Farmers in Africa often choose such a low-input, free-range production system.<ref name="Thomas de Glanville Cook Fèvre 2013">{{cite journal |last1=Thomas |first1=Lian F |last2=de Glanville |first2=William A |last3=Cook |first3=Elizabeth A |last4=Fèvre |first4=Eric M |title=The spatial ecology of free-ranging domestic pigs (Sus scrofa) in western Kenya |journal=BMC Veterinary Research |volume=9 |issue=1 |date=2013 |page=46 |issn=1746-6148 |pmid=23497587 |pmc=3637381 |doi=10.1186/1746-6148-9-46 |doi-access=free}}</ref> | |||
If conditions permit, pigs feed continuously for many hours and then sleep for many hours, in contrast to ], which tend to feed for a short time and then sleep for a short time. Pigs are ] and versatile in their feeding behaviour. They primarily eat leaves, stems, roots, fruits, and flowers.<ref>{{cite journal |last1=Kongsted |first1=A. G. |last2=Horsted |first2=K. |last3=Hermansen |first3=J. E. |year=2013 |title=Free-range pigs foraging on Jerusalem artichokes (Helianthus tuberosus L.) – Effect of feeding strategy on growth, feed conversion and animal behaviour |journal=Acta Agriculturae Scandinavica, Section A |volume=63 |issue=2 |pages=76–83 |doi=10.1080/09064702.2013.787116 |s2cid=84886946}}</ref> | |||
{{Anchor|Rooting}}Rooting is an instinctual comforting behaviour in pigs characterized by nudging the snout into something. It first happens when piglets are born to obtain their mother's milk, and can become a habitual, obsessive behaviour, most prominent in animals ] too early. Pigs root and dig into the ground to forage for food. Rooting is also a means of communication.<ref name="Minipigassoc 2016">{{cite web |title=Rooting & Nudging Behaviors in Mini Pigs |date=8 June 2016 |url=http://americanminipigassociation.com/educational/rooting-behaviors-in-mini-pigs |website=americanminipigassociation.com}}</ref> | |||
=== Intelligence === | |||
] | |||
Pigs are relatively intelligent animals, roughly on par with ]s. They distinguish each other as individuals, spend time in play, and form structured communities. They have good long-term memory and they experience emotions, changing their behaviour in response to the emotional states of other pigs. In terms of experimental tasks, pigs can perform tasks that require them to identify the locations of objects; they can solve mazes; and they can work with a simple language of symbols. They display ]. Pigs have been trained to associate different sorts of music (Bach and a military march) with food and social isolation respectively, and could communicate the resulting positive or negative emotion to untrained pigs.<ref name="Colvin Marino NHM">{{cite web |last1=Colvin |first1=Christina M. |last2=Marino |first2=Lori |title=Signs of Intelligent Life |website=Natural History Magazine |url=http://www.naturalhistorymag.com/features/122899/signs-of-intelligent-life |access-date=3 June 2019 }}</ref><ref name="nytimes2009-11-9">{{cite news |last=Angier |first=Natalie |date=9 November 2009 |title=Pigs Prove to Be Smart, if Not Vain |newspaper=] |url=https://www.nytimes.com/2009/11/10/science/10angier.html |access-date=28 July 2010}}</ref> Pigs can be trained to use a ] with their snout to select a target on screen.<ref name="Croney2021">{{cite journal |last1=Croney |first1=Candace C. |last2=Boysen |first2=Sarah T. |title=Acquisition of a Joystick-Operated Video Task by Pigs (Sus scrofa) |journal=] |volume=12 |date=11 February 2021 |doi=10.3389/fpsyg.2021.631755 |doi-access=free |pmid=33679560 |pmc=7928376 }}</ref> | |||
=== Senses === | |||
] using its sensitive nose to assist the search for wild ]s in France]] | |||
Pigs have ] vision of approximately 310° and ] of 35° to 50°. It is thought they have no ].<ref>{{cite web |title=Animalbehaviour.net (Pigs) |url=http://animalbehaviour.net/JudithKBlackshaw/Chapter3e.htm |url-status=dead |archive-url=https://web.archive.org/web/20120317194223/http://animalbehaviour.net/JudithKBlackshaw/Chapter3e.htm |archive-date=17 March 2012 |access-date=9 December 2012}}</ref> Other animals that have no accommodation, e.g. sheep, lift their heads to see distant objects.<ref>{{cite web |title=Animalbehaviour.net (Sheep) |url=http://animalbehaviour.net/JudithKBlackshaw/Chapter3b.htm |url-status=dead |archive-url=https://web.archive.org/web/20121226191014/http://animalbehaviour.net/JudithKBlackshaw/Chapter3b.htm |archive-date=26 December 2012 |access-date=9 December 2012}}</ref> The extent to which pigs have colour vision is still a source of some debate; however, the presence of ]s in the retina with two distinct wavelength sensitivities (blue and green) suggests that at least some colour vision is present.<ref name="Lomas et al., 1998)">{{cite journal |last1=Lomas |first1=C.A. |last2=Piggins |first2=D. |last3=Phillips |first3=C.J.C. |year=1998 |title=Visual awareness |journal=Applied Animal Behaviour Science |volume=57 |issue=3–4 |pages=247–257 |doi=10.1016/s0168-1591(98)00100-2}}</ref> | |||
Pigs have a well-developed sense of smell; this is exploited in Europe where ] find underground ].<ref>{{cite news |last=Sullivan |first=Walter |title=Truffles: Why Pigs Can Sniff Them Out |date=24 March 1982 |url=https://www.nytimes.com/1982/03/24/garden/truffles-why-pigs-can-sniff-them-out.html |work=]}}</ref> Pigs have 1,113 genes for smell receptors, compared to 1,094 in dogs; this may indicate an acute sense of smell, but against this, insects have only around 50 to 100 such genes but make extensive use of olfaction.<ref name="McGlone Archer Henderson 2022">{{cite journal |last1=McGlone |first1=John J. |last2=Archer |first2=Courtney |last3=Henderson |first3=Madelyn |title=Interpretive review: Semiochemicals in domestic pigs and dogs |journal=Frontiers in Veterinary Science |volume=9 |date=2022-10-25 |issn=2297-1769 |pmid=36387395 |pmc=9640746 |doi=10.3389/fvets.2022.967980 |doi-access=free}}</ref> Olfactory rather than visual stimuli are used in the identification of other pigs.<ref name="Houpt 2018">{{cite book |last=Houpt |first=Katherine A. |chapter=2. Aggression and Social Structure |title=Domestic Animal Behavior for Veterinarians and Animal Scientists |publisher=John Wiley & Sons |date=2018-03-27 |isbn=978-1-119-23276-6}}</ref> Hearing is well developed; sounds are localised by moving the head. Pigs use auditory stimuli extensively for communication in all social activities.<ref name="Gonyou 2001">{{cite book |last=Gonyou |first=H. W. |chapter=The Social Behaviour of Pigs |pages=147–176 |editor1=Keeling, L. J. |editor2=Gonyou, H. W. |title=Social behaviour in farm animals |publisher=CABI Publishing |year=2001 |isbn=978-0-85199-397-3 |doi=10.1079/9780851993973.0000}}</ref> Alarm or aversive stimuli are transmitted to other pigs not only by auditory cues but also by ]s.<ref name="Vieuille-Thomas and Signoret, (1992)">{{cite journal |last1=Vieuille-Thomas |first1=C. |last2=Signoret |first2=J. P. |year=1992 |title=Pheromonal transmission of an aversive experience in domestic pigs |journal=Journal of Chemical Ecology |volume=18 |issue=9 |pages=1551–1557 |doi=10.1007/bf00993228 |pmid=24254286 |bibcode=1992JCEco..18.1551V |s2cid=4386919}}</ref> Similarly, recognition between the sow and her piglets is by olfactory and vocal cues.<ref name="Jensen and Redbo, (1987)">{{cite journal |last1=Jensen |first1=P. |last2=Redbo |first2=I. |year=1987 |title=Behaviour during nest leaving in free-ranging domestic pigs |journal=] |volume=18 |issue=3–4 |pages=355–362 |doi=10.1016/0168-1591(87)90229-2}}</ref> | |||
== Pests and diseases == | |||
]'' larvae in uncooked pig meat]] | |||
Pigs are subject to many pests and diseases which can seriously affect productivity and cause death. These include ]s such as '']'' roundworms, ] diseases such as the ]-borne ], bacterial infections such as '']'', ] caused by '']'', and stillbirths caused by ].<ref>{{cite web |title=Disease A-Z for Pigs |url=https://www.nadis.org.uk/disease-a-z/pigs/ |website=NADIS Animal Health Skills |access-date=15 February 2024}}</ref> | |||
Some parasites of pigs are a ] risk as they can be transmitted to humans in undercooked pork. These are the pork tapeworm '']''; a protozoan, '']''; and a nematode, '']''. Transmission can be prevented by thorough sanitation on the farm; by meat inspection and careful commercial processing; and by thorough cooking, or alternatively by sufficient freezing and curing.<ref name="Gamble 1997">{{cite journal |last=Gamble |first=H. R. |title=Parasites associated with pork and pork products |journal=Revue Scientifique et Technique de l'OIE |volume=16 |issue=2 |date=1997-08-01 |doi=10.20506/rst.16.2.1032 |pages=496–506|pmid=9501363 }}</ref> | |||
== In agriculture == | |||
{{see|Pig farming#Terminology}} | |||
=== Production === | |||
{{Main|Pig farming}} | |||
Pigs have been raised outdoors, and sometimes allowed to ] in woods or pastures. In industrialized nations, pig production has largely switched to large-scale ]. This has lowered production costs but has caused concern about possible ]. As consumers have become concerned with the humane treatment of livestock, demand for pasture-raised pork in these nations has increased.<ref>{{cite news |last=Strom |first=Stephanie |date=2 January 2014 |title=Demand Grows for Hogs That Are Raised Humanely Outdoors |work=] |url=https://www.nytimes.com/2014/01/21/business/demand-grows-for-hogs-that-are-raised-humanely.html |access-date=15 April 2015}}</ref> Most pigs in the US receive ], a ], which promotes muscle instead of fat and quicker weight gain, requiring less feed to reach finishing weight, and producing less ]. China has requested that pork exports be ractopamine-free.<ref>{{cite web |last=Charles |first=Dan |date=14 August 2015 |title=A Muscle Drug For Pigs Comes Out Of The Shadows |publisher=] |url=https://www.npr.org/sections/thesalt/2015/08/14/432102733/a-muscle-drug-for-pigs-comes-out-of-the-shadows}}</ref> With a population of around 1 billion individuals, the domesticated pig is one of the most numerous large mammals on the planet.<ref>{{cite web |title=PSD Online |url=http://www.fas.usda.gov/psdonline/psdQuery.aspx |website=fas.usda.gov}}</ref><ref> {{webarchive |url=https://web.archive.org/web/20120329181120/http://www.fas.usda.gov/psdonline/psdReport.aspx?hidReportRetrievalName=Swine+Summary+Selected+Countries&hidReportRetrievalID=1649&hidReportRetrievalTemplateID=7 |date=29 March 2012}}, United States Department of Agriculture, Foreign Agricultural Service, (total number is Production (Pig Crop) plus Total Beginning Stocks</ref> | |||
Like all animals, pigs are susceptible to adverse impacts from ], such as ] from increased annual temperatures and more intense ]s. Heat stress has increased rapidly between 1981 and 2017 on pig farms in Europe. Installing a ] is an effective intervention.<ref name="Mikovits2019">{{cite journal |last1=Mikovits |first1=Christian |last2=Zollitsch |first2=Werner |last3=Hörtenhuber |first3=Stefan J. |last4=Baumgartner |first4=Johannes |last5=Niebuhr |first5=Knut |last6=Piringer |first6=Martin |last7=Anders |first7=Ivonne |last8=Andre |first8=Konrad |last9=Hennig-Pauka |first9=Isabel |last10=Schönhart |first10=Martin |last11=Schauberger |first11=Günther |display-authors=6 |date=22 January 2019 |title=Impacts of global warming on confined livestock systems for growing-fattening pigs: simulation of heat stress for 1981 to 2017 in Central Europe |journal=International Journal of Biometeorology |volume=63 |issue=2 |pages=221–230 |doi=10.1007/s00484-018-01655-0 |pmid=30671619 |bibcode=2019IJBm...63..221M |s2cid=58951606|doi-access=free }}</ref> | |||
<gallery mode=packed heights=150> | |||
File:NMA.0063325 Interiör från svinhus, Bjärka-Säby, Östergötland.jpg|Indoor pig farm, Sweden, 1911 | |||
File:Bundesarchiv Bild 183-69156-0002, Rhin-Havelluch, Sauenstall mit Ferkelbox.jpg|Sow in stall with separate piglet balcony to prevent crushing, Germany, 1959 | |||
Happy Hogs - geograph.org.uk - 279094.jpg|] pigs with field shelters, England, 2006 | |||
</gallery> | |||
<gallery class=center mode=nolines widths=220 heights=220 caption="] data for 2021"> | |||
File:World Production Of Meat, Main Items.svg|Pork is tied with chicken as the most commonly consumed meat worldwide. | |||
File:Pork_production_1961_2021.png|Pork production has grown substantially over the recent 60 years. | |||
File:Pigmeat-production-tonnes.png|Production of pork worldwide, by country in 2021. | |||
</gallery> | |||
=== Breeds === | |||
{{Main|List of pig breeds}} | |||
Around 600 ]s of pig have been created by farmers around the world, mainly in Europe and Asia, differing in coloration, shape, and size.<ref name="Miao Chen Zhang Wang 2023">{{cite journal |last1=Miao |first1=Jian |last2=Chen |first2=Zitao |last3=Zhang |first3=Zhenyang |last4=Wang |first4=Zhen |last5=Wang |first5=Qishan |last6=Zhang |first6=Zhe |last7=Pan |first7=Yuchun |title=A web tool for the global identification of pig breeds |journal=Genetics Selection Evolution |volume=55 |issue=1 |date=2023-03-21 |page=18 |issn=1297-9686 |pmid=36944938 |pmc=10029154 |doi=10.1186/s12711-023-00788-0 |doi-access=free}}</ref> According to ], as of 2016, three breeds of pig are critically rare (having a global population of fewer than 2000). They are the ], the ], and the ].<ref>{{cite web |title=The Livestock Conservancy |url=http://livestockconservancy.org/index.php/heritage/internal/conservation-priority-list#Pigs |access-date=7 March 2017 |publisher=livestock Conservancy}}</ref> The smallest known pig breed in the world is the ], typically weighing about {{convert|26|kg|lb}} as a healthy, full-grown adult.<ref>{{cite web |title=Taking good care of Ellegaard Göttingen Minipigs® |url=https://minipigs.dk/fileadmin/filer/pdf/Taking_good_care_of_Ellegaard_Goettingen_Minipigs_13.03.13.pdf |access-date=2 July 2018 |website=Ellegaard Göttingen Minipigs |archive-date=19 April 2016 |archive-url=https://web.archive.org/web/20160419112736/http://minipigs.dk/fileadmin/filer/pdf/Taking_good_care_of_Ellegaard_Goettingen_Minipigs_13.03.13.pdf |url-status=dead }}</ref> | |||
== As pets == | |||
] | |||
] pigs, a ], have been kept as pets in the United States, beginning in the latter half of the 20th century. | |||
Pigs are intelligent, social creatures. They are considered ] and are known to do quite well with people who have the usual animal allergies. Since these animals are known to have a life expectancy of 15 to 20 years, they require a long-term commitment. | |||
Given pigs are bred primarily as livestock and have not been bred as companion animals for very long, selective breeding for a placid or biddable temperament is not well established. Pigs have radically different ] and behaviours compared to dogs, and exhibit ] instincts, an independent nature, and natural ].<ref>{{Cite web |title=Info/Resource - Pigs 4 Ever - Gifts, supplies and resources for Pot-Bellied Pigs |url=https://pigs4ever.com/resource.php?recordID=6#:~:text=Pot%20belly%20pigs%20and%20children%20sometimes%20do%20not,see%20small%20children%20as%20something%20they%20can%20dominate. |access-date=11 October 2020 |website=pigs4ever.com}}</ref> Male and female swine that have not been de-sexed may express unwanted aggressive behavior, and are prone to developing serious health issues.<ref>{{Cite web |title=Spay and Neuter – American Mini Pig Association |url=http://americanminipigassociation.com/owners/helpful-owner-articles/spay-and-neuter |website=americanminipigassociation.com|date=18 September 2014 }}</ref> As ] is found to be comforting, pigs kept in the house may root household objects, furniture or surfaces. Pet pigs should be let outside to allow them to fulfill their natural desire of rooting around. | |||
== Economy == | |||
{{Infobox agricultural production|year=2019|amount1=310.4|amount10=12.7|amount9=14.1|amount8=18.4|amount7=19.6|amount6=21.6|amount5=23.7|amount4=40.6|amount3=78.7|amount2=143.1|animal=pig|country1=China (Mainland)|country10=Philippines|country9=Canada|country8=Mexico|country7=Vietnam|country6=Myanmar|country5=Russia|country4=Brazil|country3=United States|country2=European Union|world=850.3}} | |||
Approximately 1.5 billion pigs are slaughtered each year for meat.<ref>{{cite web |title=FAOSTAT |url=http://www.fao.org/faostat/en/#data/QL |access-date=25 October 2019 |website=fao.org}}</ref> | |||
The pork belly ] became an icon of ]. It appears in depictions of the arena in popular entertainment, such as the 1983 film '']''.<ref name="Davey 2011">{{cite news |last=Davey |first=Monica |title=Trade in Pork Bellies Comes to an End, but the Lore Lives |newspaper=] |date=30 July 2011 |url=https://www.nytimes.com/2011/07/31/us/31porkbelly.html?_r=0}}</ref> Trade in pork bellies declined, and they were delisted from the ] in 2011.<ref name="Davey 2011"/><ref>{{cite web |url=http://www.ftpress.com/articles/article.aspx?p=1436919&seqNum=8 |title=A Crash Course in Commodities |first=Carley |last=Garner |publisher=] |date=13 January 2010 |access-date=6 December 2011 |archive-date=21 October 2013 |url-status=dead |archive-url=https://web.archive.org/web/20131021185542/http://www.ftpress.com/articles/article.aspx?p=1436919&seqNum=8}}</ref> | |||
In 2023, China produced more pork than any other country, 55 million tonnes, followed by the European Union with 22.8 million tonnes and the United States with 12.5 million tonnes. Global production in 2023 was 120 million tonnes.<ref>{{cite web |title=Global pork production in 2022 and 2023, by country |url=https://www.statista.com/statistics/273232/net-pork-production-worldwide-by-country/ |website=Statista |access-date=15 February 2024}}</ref> India, despite its large population, consumed under 0.3 million tonnes of pork in 2023.<ref>{{cite web |title=Consumption volume of pork in India from 2013 to 2023 |url=https://www.statista.com/statistics/826720/india-pig-meat-consumption/ |publisher=Statista |access-date=15 February 2024}}</ref> International trade in pork (meat not consumed in the producing country) reached 13 million tonnes in 2020.<ref>{{cite web |last1=Ter Beek |first1=Vincent |title=The remarkable dynamics of global pig meat trade |url=https://www.pigprogress.net/the-industrymarkets/market-trends-analysis-the-industrymarkets-2/the-remarkable-dynamics-of-global-pig-meat-trade/ |website=Pig Progress |access-date=15 February 2024 |date=13 December 2023}}</ref> | |||
== Uses == | |||
=== Products === | |||
] primarily for meat, called ]. Pork is eaten in the form of pork chops, loin or rib roasts, shoulder joints, steaks, and loin (also called fillet). The many meat products made from pork include ], ] (mainly from the back and belly), and ]s.<ref>{{cite web |title=Pork Cuts |url=https://www.pork.org/cuts/ |publisher=National Pork Board |access-date=15 February 2024}}</ref> Pork is further made into ] products such as ]s, ]s, ]s and ]s.<ref>{{cite book |last=Davidson |first=Alan |author-link=Alan Davidson (food writer) |chapter=Charcuterie (with links to other chapters: Hams, Sausages, Terrines, Pâtés, Galantines, Crepinettes, Andouille and Andouillette, Blood Sausages, White Pudding, Tripe) |editor-last=Jaine |editor-first=Tom |title=] |edition=3rd |location=Oxford |publisher=] |year=2014 |isbn=978-0-19-967733-7 |page=161}}</ref> Some sausages such as ] are fermented and air-dried, to be eaten raw. There are many types, the original Italian varieties including Genovese, Milanese, and Cacciatorino, with spicier kinds from the South of Italy including Calabrese, Napoletano, and Peperone.<ref>{{cite book |last=Davidson |first=Alan |author-link=Alan Davidson (food writer) |chapter=Sausages of Italy |editor-last=Jaine |editor-first=Tom |title=The Oxford Companion to Food |edition=3rd |location=Oxford |publisher=] |year=2014 |isbn=978-0-19-967733-7 |page=719}}</ref> | |||
The hide is made into pigskin ], which is soft and durable; it can be brushed to form ] leather. These are used for products such as ]s, ]s, suede shoes, and ]s.<ref>{{cite web |title=Pig leather |url=https://www.leather-dictionary.com/index.php/Pig_leather#Pig_leather_clothing |website=Leather Dictionary |access-date=15 February 2024}}</ref> In the ], pig skin was the most popular book-binding material in ], though ] skin was more common elsewhere.<ref>{{cite web |title=Cover to Cover: Exposing the Bookbinder's Ancient Craft |url=https://www.adelaide.edu.au/library/special/exhibitions/cover-to-cover/leather/ |publisher=The ] |access-date=6 March 2024}}</ref> | |||
<gallery class=center mode=nolines widths=220 heights=220> | |||
File:Pork chops 167541218.jpg|Pork chops | |||
File:RawBacon.JPG|Streaky or side ] | |||
File:Salami aka.jpg|], a fermented and air-dried sausage, originally made in Italy | |||
File:Manuale._Aus_der_Kirchen_Agenda_-_Upper_cover_(c29h6).jpg|A 16th century book bound in pig skin | |||
File:Woman's Gloves (England), ca. 1820 (CH 18564653).jpg|A woman's suede gloves, England, c. 1820 | |||
</gallery> | |||
=== In medicine === | |||
] |volume=6 |date=16 September 2015 |page=293 |doi=10.3389/fgene.2015.00293 |doi-access=free |pmid=26442109 |pmc=4584988 }}</ref>]] | |||
Pigs, both as live animals and as a source of post-mortem tissues, are valuable animal models because of their biological, physiological, and anatomical similarities to human beings. For instance, human skin is very similar to the pigskin, therefore pigskin has been used in many preclinical studies.<ref name="porcine2018Janu">{{cite web |last=Herron |first=Alan J. |date=5 December 2009 |title=Pigs as Dermatologic Models of Human Skin Disease |url=http://www.ivis.org/proceedings/acvp/2009/Herron.pdf?LA=1 |access-date=27 January 2018 |website=ivis.org |publisher=DVM Center for Comparative Medicine and Department of Pathology Baylor College of Medicine Houston, Texas}}</ref><ref name="porcine2018Janu2">{{cite web |last1=Liu |first1=J. |last2=Kim |first2=L. |last3=Madsen |first3=T. |last4=Bouchard |first4=G. F. |title=Comparison of Human, Porcine and Rodent Wound Healing With New Miniature Swine Study Data |url=http://www.sinclairresearch.com/assets/25_Wound-Healing-AALAS-2009.pdf |access-date=27 January 2018 |website=sinclairresearch.com |publisher=Sinclair Research Centre, Auxvasse, MO, USA; Veterinary Medical Diagnostic Laboratory, Columbia, Missouri |archive-date=27 January 2018 |archive-url=https://web.archive.org/web/20180127084036/http://www.sinclairresearch.com/assets/25_Wound-Healing-AALAS-2009.pdf |url-status=dead }}</ref> | |||
Pigs are good non-human candidates for ] to humans, and in 2021 became the first animal to successfully ] body.<ref>{{cite web |title=Successful pig-to-human kidney transplant a "transformative moment" |url=https://www.yahoo.com/entertainment/successful-pig-human-kidney-transplant-130401356.html |access-date=2 November 2021 |website=www.yahoo.com |date=20 October 2021}}</ref><ref>{{cite news |last=Lapid |first=Nancy |date=20 October 2021 |title=U.S. surgeons successfully test pig kidney transplant in human patient |work=] |url=https://www.reuters.com/business/healthcare-pharmaceuticals/us-surgeons-successfully-test-pig-kidney-transplant-human-patient-2021-10-19/ |access-date=2 November 2021}}</ref> The procedure used a donor pig genetically engineered not to have a specific carbohydrate that the human body considers a threat–].<ref>{{cite web |title=Progress in Xenotransplantation Opens Door to New Supply of Critically Needed Organs |url=https://nyulangone.org/news/progress-xenotransplantation-opens-door-new-supply-critically-needed-organs |access-date=2 November 2021 |website=NYU Langone News}}</ref> Pigs are good for human donation as the risk of cross-species disease transmission is reduced by the considerable ] distance from humans.<ref name="Dooldeniya2003">{{cite journal |last1=Dooldeniya |first1=M. D. |last2=Warrens |first2=A. N. |year=2003 |title=Xenotransplantation: Where are we today? |journal=] |volume=96 |issue=3 |pages=111–117 |doi=10.1177/014107680309600303 |pmc=539416 |pmid=12612110}}</ref> They are readily available, and the danger of creating new human diseases is low as domesticated pigs have been in close contact with humans for thousands of years.<ref name="Taylor2007">Taylor, L. (2007) . Emedicine.com</ref> | |||
== Impact of pig husbandry == | |||
=== On public health === | |||
{{further|Zoonotic disease}} | |||
Pig farms can serve as reservoirs of viral diseases that are dangerous to humans and so contribute to their outbreaks in human populations.<ref name="McLean2022" /> The ] was caused by an ] variant which had first emerged in pigs.<ref>{{cite journal |last1=Mena |first1=Ignacio |last2=Nelson |first2=Martha I. |last3=Quezada-Monroy |first3=Francisco |last4=Dutta |first4=Jayeeta |last5=Cortes-Fernández |first5=Refugio |last6=Lara-Puente |first6=J Horacio |last7=Castro-Peralta |first7=Felipa |last8=Cunha |first8=Luis F. |last9=Trovão |first9=Nídia S. |last10=Lozano-Dubernard |first10=Bernardo |last11=Rambaut |first11=Andrew |last12=van Bakel |first12=Harm |last13=García-Sastre |first13=Adolfo |display-authors=6 |title=Origins of the 2009 H1N1 influenza pandemic in swine in Mexico |journal=eLife |volume=5 |date=28 June 2016 |issn=2050-084X |pmid=27350259 |pmc=4957980 |doi=10.7554/eLife.16777 |doi-access=free}}</ref> Pigs were also essential to the first outbreak of the ] in 1999, with 93% of the infected humans having had contact with pigs.<ref name="McLean2022" /> While ] is primarily spread by ]es, pigs are a known intermediary host.<ref name="WHO2015">{{cite web |title=Japanese encephalitis |url=https://www.who.int/mediacentre/factsheets/fs386/en/ |website=] |access-date=29 October 2017 |date=December 2015 |url-status=live |archive-url=https://web.archive.org/web/20170713162555/http://www.who.int/mediacentre/factsheets/fs386/en/ |archive-date=13 July 2017 |df=dmy-all}}</ref> There is also a potential for porcine ]es such as ] or ] to spill over into human populations.<ref name="McLean2022">{{cite journal |last1=McLean |first1=Rebecca |last2=Graham |first2=Simon P. |date=4 April 2022 |title=The pig as an amplifying host for new and emerging zoonotic viruses |journal=One Health |volume=14 |doi=10.1016/j.onehlt.2022.100384 |pmid=35392655 |pmc=8975596 }}</ref> | |||
=== On the environment === | |||
{{Main|Environmental impact of pig farming}} | |||
] | |||
As with the other forms of meat, producing pork is more energy-intensive than plant-based foods, and it is associated with more ] per ]. However, emissions from pork are many times smaller than those of ], ] and ], though larger than of ] meat.<ref>{{cite report |title=World Food and Agriculture – Statistical Yearbook 2023 |publisher=FAO |year=2023 |doi=10.4060/cc8166en |isbn=978-92-5-138262-2 }}</ref> | |||
Intensive pig production is also associated with ] concerns, as the swine waste is often stored above ground in so-called lagoons. These lagoons typically have high levels of ] and ], and can contain ]s like ] and ], microbial ]s, or hold elevated concentrations of pharmaceuticals from ].<ref name="Burkholder2007" /> This wastewater from lagoons is liable to reach ] on farms, though there is little evidence for it reaching deeper into local ] supplies.<ref>{{cite journal |last1=Fridrich |first1=Beata |last2=Krčmar |first2=Dejan |last3=Dalmacija |first3=Božo |last4=Molnar |first4=Jelena |last5=Pešić |first5=Vesna |last6=Kragulj |first6=Marijana |last7=Varga |first7=Nataša |date=19 January 2014 |title=Impact of wastewater from pig farm lagoons on the quality of local groundwater |journal=Agricultural Water Management |volume=135 |pages=40–53 |doi=10.1016/j.agwat.2013.12.014 |bibcode=2014AgWM..135...40F }}</ref> However, lagoon spills, such as from heavy rains in the wake of a ], can lead to fish kills and algal blooms in local rivers.<ref name="Burkholder2007" /> In the ], {{cvt|35,000|mi}} of river across over 20 states were estimated to have been contaminated by manure leakage as of 2015.<ref>{{cite web |url=https://www.epa.gov/agriculture |title=Agriculture |website=www.epa.gov |date=19 March 2015 |access-date=2017-04-23}}</ref> There is also evidence that evaporation from lagoons can cause nitrogen and phosphorus to spread through the air as dry particles then reach other water basins when they fall out through ]. This process then also contributes to water ].<ref name="Burkholder2007">{{cite journal |last1=Burkholder |first1=JoAnn |last2=Libra |first2=Bob |last3=Weyer |first3=Peter |last4=Heathcote |first4=Susan |last5=Kolpin |first5=Dana |last6=Thorne |first6=Peter S. |last7=Wichman |first7=Michael |date=14 November 2007 |title=Impacts of waste from concentrated animal feeding operations on water quality |journal=Environmental Health Perspectives |volume=115 |issue=2 |pages=308–12 |doi=10.1289/ehp.8839 |pmid=17384784 |pmc=1817674 |bibcode=2007EnvHP.115..308B }}</ref> | |||
=== On animal welfare === | |||
{{further|Cruelty to animals#Welfare concerns of farm animals|Intensive pig farming}} | |||
]s, United States, 2010]] | |||
Intensive pig production involves practices such as ], ], tattooing for litter identification, tail ], which are often done without the use of ].<ref>{{cite web |last1=Cutler |first1=R |last2=Holyoake |first2=P |date=2007 |title=The Structure and Dynamics of the Pig Meat Industry, prepared for Department of Agriculture, Fisheries and Forestry |url=http://www.agriculture.gov.au/SiteCollectionDocuments/animal-plant/animal-health/livestock-movement/pig-movement-ead.pdf |website=agriculture.gov.au}}</ref> <ref>{{cite web |url=https://www.aussiepigs.com/gallery/videos |title=Video Gallery |website=aussiepigs.com |access-date=2019-06-02 |archive-date=2019-06-02 |archive-url=https://web.archive.org/web/20190602015527/https://www.aussiepigs.com/gallery/videos |url-status=dead }}</ref> Painful ] of piglets is also done to curtail ], behavioural instability and aggression, and ], which are induced by the cramped environment.<ref name="Industry Focus" /><ref>{{Cite journal |doi = 10.2903/j.efsa.2007.611|title = The risks associated with tail biting in pigs and possible means to reduce the need for tail docking considering the different housing and husbandry systems - Scientific Opinion of the Panel on Animal Health and Welfare|journal = EFSA Journal|volume = 5|issue = 12|pages = 611|year = 2007|doi-access = free}}</ref> In English indoor farming, young pigs (less than 110kg in weight) are allowed to be kept with less than one square meter of space per pig.<ref>{{Cite web|url=https://pork.ahdb.org.uk/media/276311/keyfiguresbooklet_181001_web.pdf|title=Key figures for pig accommodation in England – legislative requirements|website=AHDB Pork|archive-url=https://web.archive.org/web/20190603062725/https://pork.ahdb.org.uk/media/276311/keyfiguresbooklet_181001_web.pdf|archive-date=3 June 2019}}</ref> | |||
Pigs often begin life in a ], which is a small pen with a central cage, designed to allow the piglets to feed from their mother while preventing her from attacking or crushing them.<ref name="auto">{{cite web |last1=Cutler |first1=R |last2=Holyoake |first2=P. |date=2007 |title=The Structure and Dynamics of the Pig Meat Industry, prepared for Department of Agriculture, Fisheries and Forestry |url=http://www.agriculture.gov.au/SiteCollectionDocuments/animal-plant/animal-health/livestock-movement/pig-movement-ead.pdf}}</ref> The crates are so small that the mother sows cannot turn around.<ref>{{cite web |url=https://assets.publishing.service.gov.uk/government/uploads/system/uploads/attachment_data/file/478588/Opinion_on_Free_Farrowing_Systems.pdf |title=Opinion on Free Farrowing Systems |website=Farm Animal Welfare Committee |archive-url=https://web.archive.org/web/20190403101822/https://assets.publishing.service.gov.uk/government/uploads/system/uploads/attachment_data/file/478588/Opinion_on_Free_Farrowing_Systems.pdf |archive-date=3 April 2019}}</ref><ref>{{Cite web |url=https://www.viva.org.uk/pigs/farrowing-crate-fact-sheet |title=Farrowing Fact Sheet |date=2016-01-06|website=Viva! - The Vegan Charity |access-date=2019-06-03 |archive-date=1 October 2020 |archive-url=https://web.archive.org/web/20201001234048/https://www.viva.org.uk/pigs/farrowing-crate-fact-sheet |url-status=dead}}</ref> While wild piglets remain with their mothers for around 12 to 14 weeks, farmed piglets are ]ed and removed from their mothers at between two and five weeks old.<ref name=":0">{{Cite web |url=https://www.ciwf.org.uk/farm-animals/pigs/pig-welfare/ |title=Pig welfare |publisher=Compassion in World Farming |access-date=3 June 2019}}</ref><ref>{{cite web |url=http://nationalhogfarmer.com/mag/farming_revisiting_weaning_age/ |title=Revisiting Weaning Age Trends, Dynamics |date=15 October 2005 |website=Nationalhogfarmer.com |access-date=28 July 2017}}</ref> Of the piglets born alive, 10% to 18% will not reach weaning age, instead succumbing to disease, starvation, dehydration, or accidental crushing by their mothers.<ref name="Industry Focus">{{cite web |publisher=Australian Pork Limited |title=Industry Focus |url=http://australianpork.com.au/industry-focus/animal-welfare/housing/ |website=australianpork.com.au}}</ref><ref>{{cite web |publisher=Pig Progress |title=Pre-weaning mortality |url=https://www.pigprogress.net/Health/Health-Tool/diseases/Pre-weaning-mortality-/ |website=pigprogress.net |date=13 July 2010 }}</ref> Unusually small ] piglets are typically killed immediately by staff through blunt trauma to the head.<ref>{{cite book |title=Model code of practice for the welfare of animals: Pigs, Primary Industries Report Series third edition |date=2008 |url=https://www.publish.csiro.au/book/5698 }}</ref><ref>{{cite web |url=https://www.animalaid.org.uk/wp-content/uploads/2016/08/Pigreport.pdf |title=Best of British? The Pig Industry Exposed |publisher=Animal Aid}}</ref> Further, intensive farming involves sows giving birth to large litter sizes at an unnatural frequency, which increases the rate of ] piglets, and causes as many as 25%-50% of sows to die of ].<ref>{{cite web |publisher=Australian Pork Limited |title=Fact Sheet – 'Reproductive Health' |date=2012 |url=https://australianpork.com.au/wp-content/uploads/2013/09/FACT-SHEET-Sectn2-Reproductive-loss__APL-Final_Jan-2012.pdf |website=australianpork.com.au |access-date=2019-06-02 |archive-date=2019-03-30 |archive-url=https://web.archive.org/web/20190330010949/https://australianpork.com.au/wp-content/uploads/2013/09/FACT-SHEET-Sectn2-Reproductive-loss__APL-Final_Jan-2012.pdf |url-status=dead }}</ref><ref>{{cite web |url=https://www.theguardian.com/environment/2018/oct/01/death-rates-surge-female-pigs-us |title='We've bred them to their limit': death rates surge for female pigs in the US |last=Greenaway |first=Twilight |date=2018-10-01 |website=] |access-date=17 November 2018}}</ref> | |||
== In culture == | |||
{{main|Pigs in culture}} | |||
Pigs, widespread in societies around the world since ] times, have been used for many purposes in art, literature, and other expressions of human ]. In classical times, the ] considered ] the finest of meats, enjoying ]s, and depicting them in their art.<ref name="MacKinnon 2001">{{cite journal |last=MacKinnon |first=Michael |title=High on the Hog: Linking Zooarchaeological, Literary, and Artistic Data for Pig Breeds in Roman Italy |journal=American Journal of Archaeology |volume=105 |issue=4 |year=2001 |pages=649–673 |doi=10.2307/507411 |jstor=507411 |s2cid=193116973}}</ref> Across Europe, pigs have been celebrated in ]s since the ],<ref name="Komins 2001">{{cite journal |last=Komins |first=Benton Jay |title=Western Culture and the Ambiguous Legacies of the Pig |journal=CLCWeb: Comparative Literature and Culture |volume=3 |issue=4 |year=2001 |doi=10.7771/1481-4374.1137 |doi-access=free }}</ref> becoming specially important in ] in cities such as ],<ref name="Newey 2014">{{cite news |last=Newey |first=Adam |title=Nuremberg, Germany: celebrating the city's sausage |url=https://www.telegraph.co.uk/travel/destinations/europe/germany/articles/Nuremberg-Germany-In-praise-of-the-citys-sausage/ |work=] |date=8 December 2014}}</ref> and in ] Italy in cities such as ].<ref>{{cite web |title=Eventi: Pane e salame |date=August 2009 |url=http://www.bibliotechebologna.it/eventi/51430/luogo/51660/date/2017-02-14:2018-03-14/date_from/2017-02-14/date_to/2018-03-14/id/100097 |publisher=Istituzione Biblioteche Bologna |access-date=31 December 2019 |language=it}}</ref><ref name="NYT Mortadella 1988">{{cite news |last1=Virbila |first1=S. Irene |title=Fare of the Country; Mortadella: Bologna's Bologna |url=https://www.nytimes.com/1988/08/07/travel/fare-of-the-country-mortadella-bologna-s-bologna.html |work=] |date=7 August 1988}}</ref> Pigs, especially miniature breeds, are occasionally kept as pets.<ref name=cabi>{{cite book |last1=Porter |first1=Valerie |last2=Alderson |first2=Lawrence |last3=Hall |first3=Stephen J.G. |last4=Sponenberg |first4=D. Phillip |chapter=Pig |date=2016 |url=https://books.google.com/books?id=2UEJDAAAQBAJ |title=Mason's World Encyclopedia of Livestock Breeds and Breeding |edition=6 | publisher=CABI |isbn=978-1-7806-4794-4 |page=616}}</ref><ref name=pbs>. PBS Nature. Accessed June 2017.</ref> | |||
In literature, both for children<ref>{{cite journal |last=Robinson |first=Robert D. |title=The Three Little Pigs: From Six Directions |journal=Elementary English |date=March 1968 |volume=45 |issue=3 |pages=356–359 |jstor=41386323}}</ref> and adults, pig characters appear in allegories, comic stories, and serious novels.<ref name="Komins 2001"/><ref name="Mullan 2010">{{cite news |last1=Mullan |first1=John |author-link1=John Mullan (academic) |title=Ten of the best pigs in literature |url=https://www.theguardian.com/books/2010/aug/21/ten-best-pigs-in-literature |work=] |date=21 August 2010}}</ref><ref name="BBC">{{cite web |last=Bragg |first= Melvyn |author-link=Melvyn Bragg |title=Topics - Pigs in literature |url=https://www.bbc.co.uk/programmes/b006qykl/topics/Pigs_in_literature |publisher=] Radio 4 |access-date=1 January 2020 |quote=Animal Farm ... Sir Gawain and the Green Knight ... The Mabinogion ... The Odyssey ... (''In Our Time'')}}</ref> In art, pigs have been represented in a wide range of media and styles from the earliest times in many cultures.<ref name="MMA">{{cite web |title=Pig |url=https://www.metmuseum.org/search-results#!/search?q=pig |publisher=] |access-date=2 January 2020}}</ref> Pig names are used in idioms and ]s, often derogatory, since pigs have long been linked with dirtiness and greed,<ref name="Telegraph">{{cite news |title=Fine Swine |date=2 February 2001 |work=] |url=https://www.telegraph.co.uk/comment/4259823/Fine-swine.html}}</ref><ref>{{cite book |author=Horwitz, Richard P. |title=Hog Ties: Pigs, Manure, and Mortality in American Culture |publisher=University of Minnesota Press |date=2002 |isbn=0816641838 |page=23}}</ref> while places such as ] are named for their association with swine.<ref name="Mills 1993">{{cite book |last1=Mills |first1=A. D. |title=A Dictionary of English Place-Names |date=1993 |publisher=Oxford University Press |isbn=0192831313 |pages=150, 318}}</ref> The eating of pork is ] and ],<ref>] 2:173, 5:3, 6:145, and 16:115.</ref><ref>{{Bibleverse||Leviticus|11:3–8|HE}}</ref> but pigs are sacred in some other religions.<ref name="Dalal2011">{{cite book |last=Dalal |first=Roshen |title=Hinduism: An Alphabetical Guide |url=https://books.google.com/books?id=DH0vmD8ghdMC&pg=PA444 |date=2011 |publisher=Penguin Books India |isbn=978-0-14-341421-6 |pages=444–445}}</ref><ref>{{cite web |last1=Bonwick |first1=James |title=Sacred Pigs |url=https://www.libraryireland.com/Druids/Sacred-Pigs.php |publisher=Library Ireland |date=1894}}</ref> | |||
<gallery class=center mode=nolines widths=220 heights=220> | |||
File:CapitalMuseum11.jpg|Bronze pig sculpture, ] | |||
File:Demeter1.jpg|Two men sacrificing a pig to ]. ], ] | |||
File:Piero di Cosimo 025.jpg|Painting of Saint Anthony with a pig in background by ] c. 1480 | |||
File:Canzone Sopra La Porcellina 1622.jpg|''Canzone Sopra La Porcellina'' ("Song Upon the Piglet") by ], ], 1622 | |||
File:Pigling Bland pg 4 Enh.jpg|] setting out on his adventures | |||
File:Bologna amazing hams sausages mortadella.jpg|]s, ]s, ]s, and ] in ], 2019 | |||
</gallery> | |||
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Revision as of 03:20, 7 December 2024
Domesticated omnivorous even-toed ungulate This article is about the domestic farm animal. For other animals known as "swine", see Suina. For other uses, see Swine (disambiguation) and Pig (disambiguation).
Pig | |
---|---|
Domestic pigs | |
Conservation status | |
Domesticated | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Family: | Suidae |
Genus: | Sus |
Species: | S. domesticus |
Binomial name | |
Sus domesticus Erxleben, 1777 | |
Synonyms | |
The pig (Sus domesticus), also called swine (pl.: swine) or hog, is an omnivorous, domesticated, even-toed, hoofed mammal. It is named the domestic pig when distinguishing it from other members of the genus Sus. It is considered a subspecies of Sus scrofa (the wild boar or Eurasian boar) by some authorities, but as a distinct species by others. Pigs were domesticated in the Neolithic, both in East Asia and in the Near East. When domesticated pigs arrived in Europe, they extensively interbred with wild boar but retained their domesticated features.
Pigs are farmed primarily for meat, called pork. The animal's skin or hide is used for leather. China is the world's largest pork producer, followed by the European Union and then the United States. Around 1.5 billion pigs are raised each year, producing some 120 million tonnes of meat, often cured as bacon. Some are kept as pets.
Pigs have featured in human culture since Neolithic times, appearing in art and literature for children and adults, and celebrated in cities such as Bologna for their meat products.
Description
The pig has a large head, with a long snout strengthened by a special prenasal bone and a disk of cartilage at the tip. The snout is used to dig into the soil to find food and is an acute sense organ. The dental formula of adult pigs is 3.1.4.33.1.4.3, giving a total of 44 teeth. The rear teeth are adapted for crushing. In males, the canine teeth can form tusks, which grow continuously and are sharpened by grinding against each other. There are four hoofed toes on each foot; the two larger central toes bear most of the weight, while the outer two are also used in soft ground. Most pigs have rather sparsely bristled hair on their skin, though there are some woolly-coated breeds such as the Mangalitsa. Adult pigs generally weigh between 140 and 300 kg (310 and 660 lb), though some breeds can exceed this range. Exceptionally, a pig called Big Bill weighed 1,157 kg (2,551 lb) and had a shoulder height of 1.5 m (4 ft 11 in).
Pigs possess both apocrine and eccrine sweat glands, although the latter are limited to the snout. Pigs, like other "hairless" mammals such as elephants, do not use thermal sweat glands in cooling. Pigs are less able than many other mammals to dissipate heat from wet mucous membranes in the mouth by panting. Their thermoneutral zone is 16–22 °C (61–72 °F). At higher temperatures, pigs lose heat by wallowing in mud or water via evaporative cooling, although it has been suggested that wallowing may serve other functions, such as protection from sunburn, ecto-parasite control, and scent-marking. Pigs are among four mammalian species with mutations in the nicotinic acetylcholine receptor that protect against snake venom. Mongooses, honey badgers, hedgehogs, and pigs all have different modifications to the receptor pocket which prevents α-neurotoxin from binding. Pigs have small lungs for their body size, and are thus more susceptible than other domesticated animals to fatal bronchitis and pneumonia. The genome of the pig has been sequenced; it contains about 22,342 protein-coding genes.
Evolution
Phylogeny
Domestic pigs are related to other pig species as shown in the cladogram, based on phylogenetic analysis using mitochondrial DNA.
Suidae |
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Taxonomy
The pig is most often considered to be a subspecies of the wild boar, which was given the name Sus scrofa by Carl Linnaeus in 1758; following from this, the formal name of the pig is Sus scrofa domesticus. However, in 1777, Johann Christian Polycarp Erxleben classified the pig as a separate species from the wild boar. He gave it the name Sus domesticus, still used by some taxonomists. The American Society of Mammalogists considers it a separate species.
Domestication in the Neolithic
Further information: DomesticationArchaeological evidence shows that pigs were domesticated from wild boar in the Near East in or around the Tigris Basin, being managed in a semi-wild state much as they are managed by some modern New Guineans. There were pigs in Cyprus more than 11,400 years ago, introduced from the mainland, implying domestication in the adjacent mainland by then. Pigs were separately domesticated in China, starting some 8,000 years ago. In the Near East, pig husbandry spread for the next few millennia. It reduced gradually during the Bronze Age, as rural populations instead focused on commodity-producing livestock, but it was sustained in cities.
Domestication did not involve reproductive isolation with population bottlenecks. Western Asian pigs were introduced into Europe, where they crossed with wild boar. There appears to have been interbreeding with a now extinct ghost population of wild pigs during the Pleistocene. The genomes of domestic pigs show strong selection for genes affecting behavior and morphology. Human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars and created domestication islands in the genome. Pigs arrived in Europe from the Near East at least 8,500 years ago. Over the next 3,000 years they interbred with European wild boar until their genome showed less than 5% Near Eastern ancestry, yet retained their domesticated features.
DNA evidence from subfossil remains of teeth and jawbones of Neolithic pigs shows that the first domestic pigs in Europe were brought from the Near East. This stimulated the domestication of local European wild boar, resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. More recently there have been complex exchanges, with European domesticated lines being exported, in turn, to the ancient Near East. Historical records indicate that Asian pigs were again introduced into Europe during the 18th and early 19th centuries.
History
Columbian Exchange
Among the animals that the Spanish introduced to the Chiloé Archipelago in the 16th century Columbian Exchange, pigs were the most successful in adapting to local conditions. The pigs benefited from abundant shellfish and algae exposed by the large tides of the archipelago. Pigs were brought to southeastern North America from Europe by de Soto and other early Spanish explorers. Escaped pigs became feral.
Feral pigs
Main article: Feral pigPigs have escaped from farms and gone feral in many parts of the world. Feral pigs in the southeastern United States have migrated north to the Midwest, where many state agencies have programs to remove them. Feral pigs in New Zealand and northern Queensland have caused substantial environmental damage. Feral hybrids of the European wild boar with the domestic pig are disruptive to both environment and agriculture, as they destroy crops, spread animal diseases including Foot-and-mouth disease, and consume wildlife such as juvenile seabirds and young tortoises. Feral pig damage is especially an issue in southeastern South America.
Reproduction
Physiology
Reproductive system of the boarReproductive system of the sowFemale pigs reach sexual maturity at 3–12 months of age and come into estrus every 18–24 days if they are not successfully bred. The variation in ovulation rate can be attributed to intrinsic factors such as age and genotype, as well as extrinsic factors like nutrition, environment, and the supplementation of exogenous hormones. The gestation period averages 112–120 days.
Estrus lasts two to three days, and the female's displayed receptiveness to mate is known as standing heat. Standing heat is a reflexive response that is stimulated when the female is in contact with the saliva of a sexually mature boar. Androstenol is one of the pheromones produced in the submaxillary salivary glands of boars that trigger the female's response. The female cervix contains a series of five interdigitating pads, or folds, that hold the boar's corkscrew-shaped penis during copulation. Females have bicornuate uteruses and two conceptuses must be present in both uterine horns to enable pregnancy to proceed. The mother's body recognises that it is pregnant on days 11 to 12 of pregnancy, and is marked by the corpus luteum's producing the sex hormone progesterone. To sustain the pregnancy, the embryo signals to the corpus luteum with the hormones estradiol and prostaglandin E2. This signaling acts on both the endometrium and luteal tissue to prevent the regression of the corpus luteum by activation of genes that are responsible for corpus luteum maintenance. During mid to late pregnancy, the corpus luteum relies primarily on luteinizing hormone for maintenance until birth.
Archeological evidence indicates that medieval European pigs farrowed, or bore a litter of piglets, once per year. By the nineteenth century, European piglets routinely double-farrowed, or bore two litters of piglets per year. It is unclear when this shift occurred. Pigs have a maximum life span of about 27 years.
Nest-building
A characteristic of pigs which they share with carnivores is nest-building. Sows root in the ground to create depressions the size of their body, and then build nest mounds, using twigs and leaves, softer in the middle, in which to give birth. When the mound reaches the desired height, she places large branches, up to 2 metres in length, on the surface. She enters the mound and roots around to create a depression within the gathered material. She then gives birth in a lying position, unlike other artiodactyls which usually stand while birthing.
Nest-building occurs during the last 24 hours before the onset of farrowing, and becomes most intense 12 to 6 hours before farrowing. The sow separates from the group and seeks a suitable nest site with well-drained soil and shelter from rain and wind. This provides the offspring with shelter, comfort, and thermoregulation. The nest provides protection against weather and predators, while keeping the piglets close to the sow and away from the rest of the herd. This ensures they do not get trampled on, and prevents other piglets from stealing milk from the sow. The onset of nest-building is triggered by a rise in prolactin level, caused by a decrease in progesterone and an increase in prostaglandin; the gathering of nest material seems to be regulated more by external stimuli such as temperature.
Nursing and suckling
Pigs have complex nursing and suckling behaviour. Nursing occurs every 50–60 minutes, and the sow requires stimulation from piglets before milk let-down. Sensory inputs (vocalisation, odours from mammary and birth fluids, and hair patterns of the sow) are particularly important immediately post-birth to facilitate teat location by the piglets. Initially, the piglets compete for position at the udder; then the piglets massage around their respective teats with their snouts, during which time the sow grunts at slow, regular intervals. Each series of grunts varies in frequency, tone and magnitude, indicating the stages of nursing to the piglets.
The phase of competition for teats and of nosing the udder lasts for about a minute, ending when milk begins to flow. The piglets then hold the teats in their mouths and suck with slow mouth movements (one per second), and the rate of the sow's grunting increases for approximately 20 seconds. The grunt peak in the third phase of suckling does not coincide with milk ejection, but rather the release of oxytocin from the pituitary into the bloodstream. Phase four coincides with the period of main milk flow (10–20 seconds) when the piglets suddenly withdraw slightly from the udder and start sucking with rapid mouth movements of about three per second. The sow grunts rapidly, lower in tone and often in quick runs of three or four, during this phase. Finally, the flow stops and so does the grunting of the sow. The piglets may dart from teat to teat and recommence suckling with slow movements, or nosing the udder. Piglets massage and suckle the sow's teats after milk flow ceases as a way of letting the sow know their nutritional status. This helps her to regulate the amount of milk released from that teat in future sucklings. The more intense the post-feed massaging of a teat, the more milk that teat later releases.
Teat order
In pigs, dominance hierarchies are formed at an early age. Piglets are precocious, and attempt to suckle soon after being born. The piglets are born with sharp teeth and fight for the anterior teats, as these produce more milk. Once established, this teat order remains stable; each piglet tends to feed on a particular teat or group of teats. Stimulation of the anterior teats appears to be important in causing milk letdown, so it might be advantageous to the entire litter to have these teats occupied by healthy piglets. Piglets locate teats by sight and then by olfaction.
Behaviour
Social
Pig behaviour is intermediate between that of other artiodactyls and of carnivores. Pigs seek out the company of other pigs and often huddle to maintain physical contact, but they do not naturally form large herds. They live in groups of about 8–10 adult sows, some young individuals, and some single males. Pigs confined in a simplified, crowded, or uncomfortable environment may resort to tail-biting; farmers sometimes dock the tails of pigs to prevent the problem, or may enrich the environment with toys or other objects to reduce the risk.
Temperature control
Because of their relative lack of sweat glands, pigs often control their body temperature using behavioural thermoregulation. Wallowing, coating the body with mud, is a common behaviour. They do not submerge completely under the mud, but vary the depth and duration of wallowing depending on environmental conditions. Adult pigs start wallowing once the ambient temperature is around 17–21 °C (63–70 °F). They cover themselves in mud from head to tail. They may use mud as a sunscreen, or to keep parasites away. Most bristled pigs "blow their coat", meaning that they shed most of the longer, coarser stiff hair once a year, usually in spring or early summer, to prepare for the warmer months ahead.
Eating, feeding, sleeping
Where pigs are allowed to roam freely, they walk roughly 4 km daily, scavenging within a home range of around a hectare. Farmers in Africa often choose such a low-input, free-range production system.
If conditions permit, pigs feed continuously for many hours and then sleep for many hours, in contrast to ruminants, which tend to feed for a short time and then sleep for a short time. Pigs are omnivorous and versatile in their feeding behaviour. They primarily eat leaves, stems, roots, fruits, and flowers.
Rooting is an instinctual comforting behaviour in pigs characterized by nudging the snout into something. It first happens when piglets are born to obtain their mother's milk, and can become a habitual, obsessive behaviour, most prominent in animals weaned too early. Pigs root and dig into the ground to forage for food. Rooting is also a means of communication.
Intelligence
Pigs are relatively intelligent animals, roughly on par with dogs. They distinguish each other as individuals, spend time in play, and form structured communities. They have good long-term memory and they experience emotions, changing their behaviour in response to the emotional states of other pigs. In terms of experimental tasks, pigs can perform tasks that require them to identify the locations of objects; they can solve mazes; and they can work with a simple language of symbols. They display self-recognition in a mirror. Pigs have been trained to associate different sorts of music (Bach and a military march) with food and social isolation respectively, and could communicate the resulting positive or negative emotion to untrained pigs. Pigs can be trained to use a joystick with their snout to select a target on screen.
Senses
Pigs have panoramic vision of approximately 310° and binocular vision of 35° to 50°. It is thought they have no eye accommodation. Other animals that have no accommodation, e.g. sheep, lift their heads to see distant objects. The extent to which pigs have colour vision is still a source of some debate; however, the presence of cone cells in the retina with two distinct wavelength sensitivities (blue and green) suggests that at least some colour vision is present.
Pigs have a well-developed sense of smell; this is exploited in Europe where trained pigs find underground truffles. Pigs have 1,113 genes for smell receptors, compared to 1,094 in dogs; this may indicate an acute sense of smell, but against this, insects have only around 50 to 100 such genes but make extensive use of olfaction. Olfactory rather than visual stimuli are used in the identification of other pigs. Hearing is well developed; sounds are localised by moving the head. Pigs use auditory stimuli extensively for communication in all social activities. Alarm or aversive stimuli are transmitted to other pigs not only by auditory cues but also by pheromones. Similarly, recognition between the sow and her piglets is by olfactory and vocal cues.
Pests and diseases
Pigs are subject to many pests and diseases which can seriously affect productivity and cause death. These include parasites such as Ascaris roundworms, virus diseases such as the tick-borne African Swine Fever, bacterial infections such as Clostridium, arthritis caused by Mycoplasma, and stillbirths caused by Parvovirus.
Some parasites of pigs are a public health risk as they can be transmitted to humans in undercooked pork. These are the pork tapeworm Taenia solium; a protozoan, Toxoplasma gondii; and a nematode, Trichinella spiralis. Transmission can be prevented by thorough sanitation on the farm; by meat inspection and careful commercial processing; and by thorough cooking, or alternatively by sufficient freezing and curing.
In agriculture
Further information: Pig farming § TerminologyProduction
Main article: Pig farmingPigs have been raised outdoors, and sometimes allowed to forage in woods or pastures. In industrialized nations, pig production has largely switched to large-scale intensive pig farming. This has lowered production costs but has caused concern about possible cruelty. As consumers have become concerned with the humane treatment of livestock, demand for pasture-raised pork in these nations has increased. Most pigs in the US receive ractopamine, a beta-agonist drug, which promotes muscle instead of fat and quicker weight gain, requiring less feed to reach finishing weight, and producing less manure. China has requested that pork exports be ractopamine-free. With a population of around 1 billion individuals, the domesticated pig is one of the most numerous large mammals on the planet.
Like all animals, pigs are susceptible to adverse impacts from climate change, such as heat stress from increased annual temperatures and more intense heatwaves. Heat stress has increased rapidly between 1981 and 2017 on pig farms in Europe. Installing a ground-coupled heat exchanger is an effective intervention.
- Indoor pig farm, Sweden, 1911
- Sow in stall with separate piglet balcony to prevent crushing, Germany, 1959
- Free range pigs with field shelters, England, 2006
- FAO data for 2021
- Pork is tied with chicken as the most commonly consumed meat worldwide.
- Pork production has grown substantially over the recent 60 years.
- Production of pork worldwide, by country in 2021.
Breeds
Main article: List of pig breedsAround 600 breeds of pig have been created by farmers around the world, mainly in Europe and Asia, differing in coloration, shape, and size. According to The Livestock Conservancy, as of 2016, three breeds of pig are critically rare (having a global population of fewer than 2000). They are the Choctaw hog, the Mulefoot, and the Ossabaw Island hog. The smallest known pig breed in the world is the Göttingen minipig, typically weighing about 26 kilograms (57 lb) as a healthy, full-grown adult.
As pets
Vietnamese Pot-bellied pigs, a miniature breed of pig, have been kept as pets in the United States, beginning in the latter half of the 20th century.
Pigs are intelligent, social creatures. They are considered hypoallergenic and are known to do quite well with people who have the usual animal allergies. Since these animals are known to have a life expectancy of 15 to 20 years, they require a long-term commitment.
Given pigs are bred primarily as livestock and have not been bred as companion animals for very long, selective breeding for a placid or biddable temperament is not well established. Pigs have radically different psychology and behaviours compared to dogs, and exhibit fight-or-flight instincts, an independent nature, and natural assertiveness. Male and female swine that have not been de-sexed may express unwanted aggressive behavior, and are prone to developing serious health issues. As rooting is found to be comforting, pigs kept in the house may root household objects, furniture or surfaces. Pet pigs should be let outside to allow them to fulfill their natural desire of rooting around.
Economy
Global pig stock | |
---|---|
in 2019 | |
Number in millions | |
1. China (Mainland) | 310.4 (36.5%) |
2. European Union | 143.1 (16.83%) |
3. United States | 78.7 (9.26%) |
4. Brazil | 40.6 (4.77%) |
5. Russia | 23.7 (2.79%) |
6. Myanmar | 21.6 (2.54%) |
7. Vietnam | 19.6 (2.31%) |
8. Mexico | 18.4 (2.16%) |
9. Canada | 14.1 (1.66%) |
10. Philippines | 12.7 (1.49%) |
World total | 850.3 |
Source: UN Food and Agriculture Organization |
Approximately 1.5 billion pigs are slaughtered each year for meat.
The pork belly futures contract became an icon of commodities trading. It appears in depictions of the arena in popular entertainment, such as the 1983 film Trading Places. Trade in pork bellies declined, and they were delisted from the Chicago Mercantile Exchange in 2011.
In 2023, China produced more pork than any other country, 55 million tonnes, followed by the European Union with 22.8 million tonnes and the United States with 12.5 million tonnes. Global production in 2023 was 120 million tonnes. India, despite its large population, consumed under 0.3 million tonnes of pork in 2023. International trade in pork (meat not consumed in the producing country) reached 13 million tonnes in 2020.
Uses
Products
Pigs are farmed primarily for meat, called pork. Pork is eaten in the form of pork chops, loin or rib roasts, shoulder joints, steaks, and loin (also called fillet). The many meat products made from pork include ham, bacon (mainly from the back and belly), and sausages. Pork is further made into charcuterie products such as terrines, galantines, pâtés and confits. Some sausages such as salami are fermented and air-dried, to be eaten raw. There are many types, the original Italian varieties including Genovese, Milanese, and Cacciatorino, with spicier kinds from the South of Italy including Calabrese, Napoletano, and Peperone.
The hide is made into pigskin leather, which is soft and durable; it can be brushed to form suede leather. These are used for products such as gloves, wallets, suede shoes, and leather jackets. In the 16th century, pig skin was the most popular book-binding material in Germany, though calf skin was more common elsewhere.
- Pork chops
- Streaky or side bacon
- Salami, a fermented and air-dried sausage, originally made in Italy
- A 16th century book bound in pig skin
- A woman's suede gloves, England, c. 1820
In medicine
Pigs, both as live animals and as a source of post-mortem tissues, are valuable animal models because of their biological, physiological, and anatomical similarities to human beings. For instance, human skin is very similar to the pigskin, therefore pigskin has been used in many preclinical studies.
Pigs are good non-human candidates for organ donation to humans, and in 2021 became the first animal to successfully donate an organ to a human body. The procedure used a donor pig genetically engineered not to have a specific carbohydrate that the human body considers a threat–Galactose-alpha-1,3-galactose. Pigs are good for human donation as the risk of cross-species disease transmission is reduced by the considerable phylogenetic distance from humans. They are readily available, and the danger of creating new human diseases is low as domesticated pigs have been in close contact with humans for thousands of years.
Impact of pig husbandry
On public health
Further information: Zoonotic diseasePig farms can serve as reservoirs of viral diseases that are dangerous to humans and so contribute to their outbreaks in human populations. The 2009 swine flu pandemic was caused by an influenza A variant which had first emerged in pigs. Pigs were also essential to the first outbreak of the Nipah virus in 1999, with 93% of the infected humans having had contact with pigs. While Japanese encephalitis is primarily spread by mosquitoes, pigs are a known intermediary host. There is also a potential for porcine coronaviruses such as porcine epidemic diarrhea virus or swine acute diarrhea syndrome coronavirus to spill over into human populations.
On the environment
Main article: Environmental impact of pig farmingAs with the other forms of meat, producing pork is more energy-intensive than plant-based foods, and it is associated with more greenhouse gas emissions per calorie. However, emissions from pork are many times smaller than those of beef, veal and mutton, though larger than of chicken meat.
Intensive pig production is also associated with water pollution concerns, as the swine waste is often stored above ground in so-called lagoons. These lagoons typically have high levels of nitrogen and phosphorus, and can contain toxic heavy metals like zinc and copper, microbial pathogens, or hold elevated concentrations of pharmaceuticals from subtherapeutic antibiotic use in swine. This wastewater from lagoons is liable to reach groundwater on farms, though there is little evidence for it reaching deeper into local drinking water supplies. However, lagoon spills, such as from heavy rains in the wake of a hurricane, can lead to fish kills and algal blooms in local rivers. In the United States, 35,000 mi (56,000 km) of river across over 20 states were estimated to have been contaminated by manure leakage as of 2015. There is also evidence that evaporation from lagoons can cause nitrogen and phosphorus to spread through the air as dry particles then reach other water basins when they fall out through dry deposition. This process then also contributes to water eutrophication.
On animal welfare
Further information: Cruelty to animals § Welfare concerns of farm animals, and Intensive pig farmingIntensive pig production involves practices such as castration, earmarking, tattooing for litter identification, tail docking, which are often done without the use of anesthetic. Painful teeth clipping of piglets is also done to curtail cannibalism, behavioural instability and aggression, and tail biting, which are induced by the cramped environment. In English indoor farming, young pigs (less than 110kg in weight) are allowed to be kept with less than one square meter of space per pig.
Pigs often begin life in a farrowing or gestation crate, which is a small pen with a central cage, designed to allow the piglets to feed from their mother while preventing her from attacking or crushing them. The crates are so small that the mother sows cannot turn around. While wild piglets remain with their mothers for around 12 to 14 weeks, farmed piglets are weaned and removed from their mothers at between two and five weeks old. Of the piglets born alive, 10% to 18% will not reach weaning age, instead succumbing to disease, starvation, dehydration, or accidental crushing by their mothers. Unusually small runt piglets are typically killed immediately by staff through blunt trauma to the head. Further, intensive farming involves sows giving birth to large litter sizes at an unnatural frequency, which increases the rate of stillborn piglets, and causes as many as 25%-50% of sows to die of prolapse.
In culture
Main article: Pigs in culturePigs, widespread in societies around the world since Neolithic times, have been used for many purposes in art, literature, and other expressions of human culture. In classical times, the Romans considered pork the finest of meats, enjoying sausages, and depicting them in their art. Across Europe, pigs have been celebrated in carnivals since the Middle Ages, becoming specially important in Medieval Germany in cities such as Nuremberg, and in Early Modern Italy in cities such as Bologna. Pigs, especially miniature breeds, are occasionally kept as pets.
In literature, both for children and adults, pig characters appear in allegories, comic stories, and serious novels. In art, pigs have been represented in a wide range of media and styles from the earliest times in many cultures. Pig names are used in idioms and animal epithets, often derogatory, since pigs have long been linked with dirtiness and greed, while places such as Swindon are named for their association with swine. The eating of pork is forbidden in Islam and Judaism, but pigs are sacred in some other religions.
- Bronze pig sculpture, Zhou dynasty
- Two men sacrificing a pig to Demeter. Red-figure pot, Ancient Greece
- Painting of Saint Anthony with a pig in background by Piero di Cosimo c. 1480
- Canzone Sopra La Porcellina ("Song Upon the Piglet") by Giulio Cesare Croce, Bologna, 1622
- Pigling Bland setting out on his adventures
- Hams, pig's trotters, sausages, and mortadella in Bologna, 2019
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External links
Taxon identifiers | |
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Sus scrofa domesticus | |
Sus scrofa scrofa |