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{{Short description|Neural pathway which controls a reflex}} | |||
A ] is the neural pathway ] a ]. It generally does not involve the brain. Instead of the brain it can include a ] ] composed of ] to connect ] and ] signals. While the ] generation may be initiated by ] input, an extensive processing takes place within the ]. The neural connection from the primary sensory neurons to the motor neurons is a ] (Andersen). | |||
] | |||
A '''reflex arc''' is a ] that controls a ]. In ]s, most ]s do not pass directly into the ], but ] in the ]. This allows for faster reflex actions to occur by activating spinal ]s without the delay of routing signals through the brain. The brain will receive the input while the reflex is being carried out and the analysis of the signal takes place after the reflex action. | |||
A stimulus follows the following path through the reflex arc: | |||
There are two types: ] reflex arc (affecting inner organs) and ] reflex arc (affecting muscles). Autonomic reflexes sometimes involve the spinal cord and some somatic reflexes are mediated more by the brain than the spinal cord.<ref name=":0">{{Cite book|title=Anatomy & Physiology: The Unity of Form and Function|last=Saladin|first=Kenneth|publisher=McGraw-Hill|year=2015|isbn=978-0073403717|location=New York|pages=496–497}}</ref> | |||
Stimulus (ex.a light) -----> Sensory neuron (photoreceptors in the eyes for light) -----> Interneuron (connects neurons) ----> Motor neuron (causes pupil to dilate) | |||
During a somatic reflex, nerve signals travel along the following pathway:<ref name=":0" /> | |||
A reflex to a stimulus is almost simultaneous, as the reflex arc doesn't involve the brain at all. | |||
# ''Somatic receptors'' in the skin, muscles and tendons | |||
# ''Afferent nerve fibers'' carry signals from the somatic receptors to the ] of the spinal cord or to the brainstem | |||
# An ''integrating center'', the point at which the neurons that compose the gray matter of the spinal cord or brainstem synapse | |||
# ''Efferent nerve fibers'' carry motor nerve signals from the ] to the muscles | |||
# ''Effector'' muscle innervated by the efferent nerve fiber carries out the response. | |||
A reflex arc, then, is the pathway followed by nerves which (a.) carry sensory information from the receptor to the spinal cord, and then (b.) carry the response generated by the spinal cord to effector organs during a reflex action. | |||
The pathway taken by the nerve impulse to accomplish a reflex action is called the reflex arc. | |||
==Monosynaptic vs. polysynaptic== | |||
] | |||
When a reflex arc in an animal consists of only one ] and one ], it is defined as '''monosynaptic''', referring to the presence of a single ]. In the case of peripheral muscle reflexes (], ]), brief stimulation to the ] results in contraction of the agonist or effector muscle. | |||
By contrast, in '''polysynaptic''' reflex pathways, one or more interneurons connect ] (]) and ] (]) signals. All but the most simple reflexes are polysynaptic, allowing processing or inhibition of polysynaptic reflexes within the brain.{{Clarify | date=August 2019 | reason=This sentence is ambiguous: doesn't being 'monosynaptic' allow the brain to inhibit reflexes? Does the brain always process polysynaptic reflexes? }} | |||
== The patellar reflex (aka "knee jerk") == | |||
{{main article|Patellar reflex}} | |||
] | |||
When the ] is tapped just below the knee, the tap initiates an ] in a specialized structure known as a ] located within the ]. This action potential travels to the L3 and L4 nerve roots of the spinal cord,<ref name=":1"> | |||
{{Cite web | |||
| title= Deep Tendon Reflexes | |||
| date= November 28, 2016 | |||
| website= The Precise Neurological Exam | |||
| publisher= New York University School of Medicine | |||
| url= https://informatics.med.nyu.edu/modules/pub/neurosurgery/reflexes.html | |||
| access-date= November 28, 2016 | |||
}} | |||
</ref> | |||
via a sensory ] which chemically communicates by releasing ] onto a ]. The result of this motor nerve activity is contraction of the quadriceps muscle, leading to extension of the lower leg at the knee (i.e. the lower leg kicks forward). Ultimately, an improper patellar reflex may indicate an injury of the ].<ref name=":1"/> | |||
The sensory input from the quadriceps also activates local ]s that release the inhibitory ] ] onto motor neurons of ] muscles, blocking their stimulation (in this case the ]). The relaxation of the opposing muscle facilitates (by not opposing) the extension of the lower leg. | |||
In invertebrates reflex interneurons do not necessarily reside in the spinal cord, for example as in the ] of ]. | |||
==See also== | ==See also== | ||
*] | * ] | ||
*] | |||
== |
==References== | ||
{{Reflist}} | |||
⚫ | |||
==External links== | |||
] | |||
* {{Cite thesis | |||
| author = Ole Kæseler Andersen | |||
| url = http://person.hst.aau.dk/oka/oka_phd_thesis.pdf | |||
⚫ | | title = Physiological and Pharmacological modulation of the human nociceptive withdrawal reflex | ||
| type = PhD Thesis | |||
| publisher = Center for Sansory-Motor Interaction, Aalborg University | |||
| date = 1996 | |||
}} | |||
* | |||
] | |||
<div class="boilerplate metadata" id="stub"><table cellpadding="0" cellspacing="0" style="background-color: transparent;"><tr><td>]</td><td>'' This ] article is a ]. You can ] Misplaced Pages by ''.</td></tr></table></div>] |
Latest revision as of 12:01, 19 January 2024
Neural pathway which controls a reflexA reflex arc is a neural pathway that controls a reflex. In vertebrates, most sensory neurons do not pass directly into the brain, but synapse in the spinal cord. This allows for faster reflex actions to occur by activating spinal motor neurons without the delay of routing signals through the brain. The brain will receive the input while the reflex is being carried out and the analysis of the signal takes place after the reflex action.
There are two types: autonomic reflex arc (affecting inner organs) and somatic reflex arc (affecting muscles). Autonomic reflexes sometimes involve the spinal cord and some somatic reflexes are mediated more by the brain than the spinal cord.
During a somatic reflex, nerve signals travel along the following pathway:
- Somatic receptors in the skin, muscles and tendons
- Afferent nerve fibers carry signals from the somatic receptors to the posterior horn of the spinal cord or to the brainstem
- An integrating center, the point at which the neurons that compose the gray matter of the spinal cord or brainstem synapse
- Efferent nerve fibers carry motor nerve signals from the anterior horn to the muscles
- Effector muscle innervated by the efferent nerve fiber carries out the response.
A reflex arc, then, is the pathway followed by nerves which (a.) carry sensory information from the receptor to the spinal cord, and then (b.) carry the response generated by the spinal cord to effector organs during a reflex action. The pathway taken by the nerve impulse to accomplish a reflex action is called the reflex arc.
Monosynaptic vs. polysynaptic
When a reflex arc in an animal consists of only one sensory neuron and one motor neuron, it is defined as monosynaptic, referring to the presence of a single chemical synapse. In the case of peripheral muscle reflexes (patellar reflex, achilles reflex), brief stimulation to the muscle spindle results in contraction of the agonist or effector muscle. By contrast, in polysynaptic reflex pathways, one or more interneurons connect afferent (sensory) and efferent (motor) signals. All but the most simple reflexes are polysynaptic, allowing processing or inhibition of polysynaptic reflexes within the brain.
The patellar reflex (aka "knee jerk")
Main article: Patellar reflexWhen the patellar tendon is tapped just below the knee, the tap initiates an action potential in a specialized structure known as a muscle spindle located within the quadriceps. This action potential travels to the L3 and L4 nerve roots of the spinal cord, via a sensory axon which chemically communicates by releasing glutamate onto a motor nerve. The result of this motor nerve activity is contraction of the quadriceps muscle, leading to extension of the lower leg at the knee (i.e. the lower leg kicks forward). Ultimately, an improper patellar reflex may indicate an injury of the central nervous system.
The sensory input from the quadriceps also activates local interneurons that release the inhibitory neurotransmitter glycine onto motor neurons of antagonist muscles, blocking their stimulation (in this case the hamstring muscles). The relaxation of the opposing muscle facilitates (by not opposing) the extension of the lower leg.
In invertebrates reflex interneurons do not necessarily reside in the spinal cord, for example as in the lateral giant neuron of crayfish.
See also
References
- ^ Saladin, Kenneth (2015). Anatomy & Physiology: The Unity of Form and Function. New York: McGraw-Hill. pp. 496–497. ISBN 978-0073403717.
- ^ "Deep Tendon Reflexes". The Precise Neurological Exam. New York University School of Medicine. November 28, 2016. Retrieved November 28, 2016.
External links
- Ole Kæseler Andersen (1996). Physiological and Pharmacological modulation of the human nociceptive withdrawal reflex (PDF) (PhD Thesis). Center for Sansory-Motor Interaction, Aalborg University.
- Tutorial at wisc-online.com