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Capra Temporal range: 2.58–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ Early Pleistocene-Present
Capra, St. Leonhard in Passeier, Italy
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Family:	Bovidae
Subfamily:	Caprinae
Tribe:	Caprini
Genus:	Capra Linnaeus, 1758
Type species
Capra hircus Linnaeus, 1758
Species
See text.
Approximate range of the Capra species

Capra is a genus of mammals, the goats, composed of up to nine species, including the markhor and many species known as ibexes. The domestic goat (Capra hircus) is a domesticated species derived from the wild goat (Capra aegagrus). Evidence of goat domestication dates back more than 8,500 years.

Wild goats are animals of mountain habitats. They are very agile and hardy, able to climb on bare rock and survive on sparse vegetation. They can be distinguished from the genus Ovis, which includes sheep, by the presence of scent glands close to the feet, in the groin, and in front of the eyes, and the absence of other facial glands, and by the presence of a beard in some specimens, and of hairless calluses on the knees of the forelegs.

The Rocky Mountain goat is in a separate genus, Oreamnos. Present-day genetic and phenotypic differences between the Capra species are largely related to (1) discontinuity of and impeded migration between Capra populations during glacial periods, and (2) insufficient time in the postglacial period for now-adjoining Capra populations to overcome behavioral mechanisms impeding hybridization in the wild so as to erase these differences.

Taxonomy

All members of the genus Capra are bovids (members of the family Bovidae), and more specifically caprids (subfamily Caprinae). As such they are ruminants, meaning they chew the cud, and have four-chambered stomachs which play a vital role in digesting, regurgitating, and redigesting their food.

The genus has sometimes been taken to include Ovis (sheep) and Ammotragus (Barbary sheep), but these are usually regarded as distinct genera, leaving Capra for goats and ibexes. In this smaller genus, some authors have recognized only two species, the markhor on one side and all other forms included in one species on the other side. Today, nine species are usually accepted:

• Wild goat (Capra aegagrus)
  • Bezoar ibex (Capra aegagrus aegagrus)
  • Sindh ibex (Capra aegagrus blythi)
• West Caucasian tur (Capra caucasica)
• East Caucasian tur (Capra cylindricornis)
• Markhor (Capra falconeri)
• Domestic goat (Capra hircus; includes feral goat, sometimes considered a subspecies of C. aegagrus)
• Alpine ibex (Capra ibex)
• Nubian ibex (Capra nubiana)
• Spanish ibex (Capra pyrenaica)
• Siberian ibex (Capra sibirica)
• Walia ibex (Capra walie)

The goats of the genus Capra have complex systematic relationships, which are still not completely resolved. Recent studies based on mitochondrial DNA suggest that the Siberian ibex and the Nubian ibex represent distinct species, which are not very closely related to the physically similar Alpine ibex. The Alpine ibex forms a group with the Spanish ibex. The West Caucasian tur appears to be more closely related to the wild goat than to the East Caucasian tur. The markhor is relatively little separated from other forms—previously it had been considered to be a separate branch of the genus.

Almost all wild goat species are allopatric (geographically separated)—the only geographical overlaps are the wild goat (Capra hircus) with the East Caucasian tur (Capra caucasica cylindricornis), and the markhor (Capra falconeri) with the Siberian ibex (Capra siberica). In both cases, the overlapping species do not usually interbreed in the wild, but in captivity, all Capra species can interbreed, producing fertile offspring.

Species and subspecies

Subgenus	Image	Scientific name	Common Name	Distribution
Brachyceros		Capra caucasica	West Caucasian tur	Caucasus Mountains range.
		Capra cylindricornis	East Caucasian tur	Greater Caucasus Mountains
Aegoceros		Capra falconeri	Markhor	Central Asia, Karakoram and the Himalayas
Capra		Capra aegagrus	Wild goat	Turkey and the Caucasus in the west to Turkmenistan, Afghanistan and Pakistan
		Capra hircus	Domestic goat	Domesticated
		Capra sibirica	Siberian ibex	central and northern Asia, Afghanistan, western and northern China (Primarily Xinjiang), north-western India, south-eastern Kazakhstan, Kyrgyzstan, Tajikistan, eastern Uzbekistan, Mongolia, northern Pakistan, and south-central Russia.
		Capra pyrenaica	Spanish ibex	Iberian Peninsula
		Capra walie	Walia ibex	Ethiopia
		Capra ibex	Alpine ibex	France, Italy, Switzerland, Liechtenstein, Bavaria, Austria and Slovenia
		Capra nubiana	Nubian ibex	Algeria, Egypt, Ethiopia, Eritrea, Israel, Jordan, Lebanon, Oman, Saudi Arabia, Sudan, and Yemen

Domestication and uses

Along with sheep, goats were among the first domesticated animals. The domestication process started at least 10,000 years ago in what is now northern Iran. Easy human access to goat hair, meat, and milk were the primary motivations. Goat skins were popularly used until the Middle Ages for water and wine bottles when traveling and camping, and in certain regions as parchment for writing.

References

1. Parrini, F.; et al. (2009). "Capra ibex (Artiodactyla: Bovidae)". Mammalian Species. 830: 1–12. doi:10.1644/830.1.
2. Gavashelishvili, A.; Yarovenko, Y. A.; Babayev, E. A.; Mikeladze, G.; Gurielidze, Z.; Dekanoidze, D.; Kerdikoshvili, N.; Ninua, L.; Paposhvili, N. (2018). "Modeling the distribution and abundance of eastern tur (Capra cylindricornis) in the Caucasus". Journal of Mammalogy. 99 (4): 885–897. doi:10.1093/jmammal/gyy056.
3. Ansell, W. F. H. 1972. Order Artiodactyla. Part 15. Pp. 1–84, in The mammals of Africa: An identification manual (J. Meester and H. W. Setzer, eds.) . Smithsonian Institution Press, Washington, D.C., not continuously paginated. (quoted in Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN 0-8018-8221-4. OCLC 62265494.)
4. Haltenorth, T. 1963. Klassifikation der Säugetiere: Artiodactyla I. Handbuch der Zoologie, 8(32):1–167 (quoted in Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN 0-8018-8221-4. OCLC 62265494.)
5. Nathalie Pidancier, Steve Jordan, Gordon Luikart, Pierre Taberlet: Evolutionary history of the genus Capra (Mammalia, Artiodactyla): Discordance between mitochondrial DNA and Y-chromosome. Molecular Phylogenetics and Evolution 40 (2006) 739–749 online
6. Phylogenetic Reconstructions in the Genus Capra (Bovidae, Artiodactyla) Based on the Mitochondrial DNA Analysis. Russian Journal of Genetics, 2007, Vol. 43, No. 2, pp. 181–189. online
7. V. G. Heptner: Mammals of the Sowjetunion Vol. I UNGULATES. Leiden, New York, 1989 ISBN 90-04-08874-1
8. Melinda A. Zeder, Brian Hesse: The Initial Domestication of Goats (Capra hircus) in the Zagros Mountains 10,000 Years Ago. Science 24 March 2000: Vol. 287. no. 5461, pp. 2254–2257 online abstract

External links

• "Ibex" . New International Encyclopedia. 1905.

• Capra (genus)
• Goats
• Mammals of Asia
• Mammals of Africa
• Mammal genera
• Extant Pleistocene first appearances
• Taxa named by Carl Linnaeus