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Capra Temporal range: 2.6–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ Pleistocene-Present
Female and male Siberian ibex at the Berlin Zoological Garden
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Family:	Bovidae
Subfamily:	Caprinae
Tribe:	Caprini
Genus:	Capra Linnaeus, 1758
Species
See text.
Approximate range of the Capra species

Capra is a genus of mammals, the goats or wild goats, composed of up to nine species, including the wild goat, the markhor, and several species known as ibex. The domestic goat (Capra aegagrus hircus) is a domesticated subspecies of the wild goat (Capra aegagrus). Evidence of goat domestication dates back more than 8,500 years.

Wild goats are animals of mountain habitats. They are very agile and hardy, able to climb on bare rock and survive on sparse vegetation. They can be distinguished from the genus Ovis, which includes sheep, by the presence of scent glands close to the feet, in the groin, and in front of the eyes, and the absence of other facial glands, and by the presence of a beard in the males, and of hairless calluses on the knees of the forelegs.

The Rocky Mountain goat is in a separate genus, Oreamnos.

Taxonomy

All members of the Capra genus are bovids (members of the family Bovidae), and more specifically caprids (subfamily Caprinae). As such they are ruminants, meaning they chew the cud, and have four-chambered stomachs which play a vital role in digesting, regurgitating, and redigesting their food.

The genus has sometimes been taken to include Ovis (sheep) and Ammotragus (Barbary sheep), but these are usually regarded as distinct genera, leaving Capra for goats and ibexes. In this smaller genus, some authors have recognized only two species, the markhor on one side and all other forms included in one species on the other side. Today, nine species are usually accepted:

• West Caucasian tur (Capra caucasica)
  • East Caucasian tur (Capra caucasica cylindricornis)
• Markhor (Capra falconeri)
• Alpine ibex (Capra ibex)
• Nubian ibex (Capra nubiana)
• Spanish ibex (Capra pyrenaica)
• Siberian ibex (Capra sibirica)
• Walia ibex (Capra walie)
• Wild goat (Capra aegagrus)
  • Domestic goat (Capra aegagrus hircus; includes feral goat)
  • Bezoar ibex (Capra aegagrus aegagrus)

The goats of the genus Capra have complex systematic relationships, which are still not completely resolved. Recent studies based on mitochondrial DNA suggest that the Siberian ibex and the Nubian ibex represent distinct species, which are not very closely related to the physically similar Alpine ibex. The Alpine ibex forms a group with the Spanish ibex. The West Caucasian tur appears to be more closely related to the wild goat than to the East Caucasian tur. The markhor is relatively little separated from other forms—previously it had been considered to be a separate branch of the genus.

Almost all wild goat species are allopatric (geographically separated)—the only geographical overlaps are the wild goat (Capra hircus) with the East Caucasian tur (Capra caucasica cylindricornis), and the markhor (Capra falconeri) with the Siberian ibex (Capra siberica). In both cases, the overlapping species do not usually interbreed in the wild, but in captivity, all Capra species can interbreed, producing fertile offspring.

Species and subspecies

	Capra caucasica	West Caucasian tur
	Capra cylindricornis	East Caucasian tur
	Capra falconeri	Markhor
	Capra aegagrus	Wild goat
	Capra (aegagrus) hircus	Domestic goat
	Capra sibirica	Siberian ibex
	Capra pyrenaica	Spanish ibex
	Capra walie	Walia ibex
	Capra ibex	Alpine ibex
	Capra nubiana	Nubian ibex

Domestication and uses

Along with sheep, goats were among the first domesticated animals. The domestication process started at least 10,000 years ago in what is now northern Iran. Easy human access to goat hair, meat, and milk were the primary motivations. Goat skins were popularly used until the Middle Ages for water and wine bottles when traveling and camping, and in certain regions as parchment for writing.

In ancient history

Evidence of the ibex is widely present in the archaeological record, particularly in the Near East and Mediterranean regions. Ibex motifs are very common on cylinder seals and pottery, both painted and embossed. Excavations from Minoan Crete at Knossos, for example, have yielded specimens from about 1800 BC, including one cylinder seal depicting an ibex defending itself from a hunting dog. From the similar age a gold jewelry ibex image was found at the Akrotiri archaeological site on Santorini in present-day Greece.

An Iron Age Capra ibex specimen was recovered at the Aq Kupruk archaeological site in present-day Afghanistan, illustrating either domestication or hunting of the ibex by these early peoples. However, archaeological records of ibex can be difficult to separate from those of domestic goats. A 4,000-year-old rock drawing of an ibex deer, complete with long curled horns, was discovered near Khomeyn, Iran.

References

1. Parrini, F.; et al. (2009). "Capra ibex (Artiodactyla: Bovidae)". Mammalian Species. 830: 1–12. doi:10.1644/830.1.
2. Ansell, W. F. H. 1972. Order Artiodactyla. Part 15. Pp. 1–84, in The mammals of Africa: An identification manual (J. Meester and H. W. Setzer, eds.) . Smithsonian Institution Press, Washington, D.C., not continuously paginated. (quoted in Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN 0-8018-8221-4. OCLC 62265494.)
3. Haltenorth, T. 1963. Klassifikation der Säugetiere: Artiodactyla I. Handbuch der Zoologie, 8(32):1–167 (quoted in Grubb, P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore, MD: Johns Hopkins University Press. ISBN 0-8018-8221-4. OCLC 62265494.)
4. Nathalie Pidancier, Steve Jordan, Gordon Luikart, Pierre Taberlet: Evolutionary history of the genus Capra (Mammalia, Artiodactyla): Discordance between mitochondrial DNA and Y-chromosome. Molecular Phylogenetics and Evolution 40 (2006) 739–749 online
5. Phylogenetic Reconstructions in the Genus Capra (Bovidae, Artiodactyla) Based on the Mitochondrial DNA Analysis. Russian Journal of Genetics, 2007, Vol. 43, No. 2, pp. 181–189. online
6. V. G. Heptner: Mammals of the Sowjetunion Vol. I UNGULATES. Leiden, New York, 1989 ISBN 90-04-08874-1
7. Melinda A. Zeder, Brian Hesse: The Initial Domestication of Goats (Capra hircus) in the Zagros Mountains 10,000 Years Ago. Science 24 March 2000: Vol. 287. no. 5461, pp. 2254–2257 online abstract
8. C. Michael Hogan, Knossos fieldnotes, Modern Antiquarian (2007)
9. M. Uda, G. Demortier, I. Nakai, X-rays for archaeology, 2005, Springer, ISBN 1-4020-3580-2
10. Pam J. Crabtree, Douglas V. Campana, Kathleen Ryan, Early Animal Domestication and Its Cultural Context, 1989, University of Pennsylvania Museum of Archaeology ISBN 0-924171-96-0
11. http://www.independent.co.uk/news/world/middle-east/world-oldest-rock-drawings-archaeologist-iran-khomeyn-mohammed-naserifard-a7470321.html

External links

• "Ibex" . New International Encyclopedia. 1905.

• Capra
• Goats
• Mammals of Asia
• Mammals of Africa
• Mammal genera
• Extant Pleistocene first appearances
• Taxa named by Carl Linnaeus