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Spinosaurus Temporal range: mid Cretaceous
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Sauropsida
Superorder:	Dinosauria
Order:	Saurischia
Suborder:	Theropoda
Superfamily:	Megalosauroidea
Family:	Spinosauridae
Genus:	Spinosaurus Stromer, 1915
Species
S. aegyptiacus (type) ?S. marocannus

Spinosaurus (meaning "spine lizard") was a theropod dinosaur genus that lived in what is now Egypt, from the Albian to early Cenomanian stages of the Cretaceous period, about 95 to 93 million years ago. According to a study by dal Sasso et al. (2006), it was the largest of all carnivorous dinosaurs by a significant margin, even larger than Tyrannosaurus rex and Giganotosaurus. Spinosaurus was the longest theropod, measuring 16 to 18 metres (52 to 60 feet) long and standing at 5 to 6 metres (16 to 20 feet) in height. Spinosaurus reached weights up to 9 tons, although further comparisons with related species suggest that adults might have reached sizes up to 20 tons and 21 m (70 ft) in length.

The distinctive spines of these animals (large bones extending from the vertebrae) grew up to 2 m long and were likely to have had skin stretching between them, forming a sail-like structure, although some have suggested they were covered in muscle and formed a hump or ridge.

Spinosaurus provides the name of a family of dinosaurs, the Spinosauridae, of which other members include Angaturama (probably synonymous with Irritator), Baryonyx, Irritator, Suchomimus, and Siamosaurus.

Description

Although Spinosaurus is well-known to dinosaur enthusiasts, because of its unusual features, it is mostly known from remains that have been destroyed, aside from a few more recently discovered teeth and skull elements. Jaw and skull material published in 2006 show that it had one of the longest skulls of any carnivorous dinosaur, estimated by dal Sasso et. al. at about 175 cm (5.7 ft).

Originally found in the Bahariya Valley of Egypt in 1912, it was named by German paleontologist Ernst Stromer in 1915. Some of the fossils were damaged during transport back to the Deutsches Museum, Munich, Germany and the remaining bones were completely lost due to Allied bombing in 1944.

Aside from its 'sail', notable characteristics of Spinosaurus include:

• A long, narrow snout, similar to other Spinosaurids and, like them, filled with conical teeth.
• One enlarged, hook-like claw on each of its front limbs, perhaps for catching fish.
• Relatively short legs and long 'arms', leading some paleontologists to suggest it may have been quadrupedal, rather than strictly bipedal (though it was undoubtedly capable of at least facultative bipedality).

Much of this is speculation based on Baryonyx and other spinosaurids, as no limb material has ever been attributed to Spinosaurus itself.

Diet

It is unclear whether Spinosaurus was a cursorial predator or a fisher, as indicated by its elongated jaws, conical teeth and raised nostrils. The only direct evidence for spinosaur diet comes from related European and South American species. Baryonyx was found with both fish scales and bones from juvenile Iguanodon in its stomach, while a tooth embedded in a South American pterosaur bone suggests that spinosaurs occasionally preyed on these flying archosaurs. Spinosaurus was likely to have been a more generalized and opportunistic predator, possibly a Cretaceous equivalent of large grizzly bears, being biased toward fishing, although it undoubtedly took many kinds of small-to-medium-sized prey.

Sail

Spinosaurus sails were unusual, although other dinosaurs of the same time and area, namely the ornithopod Ouranosaurus and the sauropod Rebbachisaurus, developed a similar structural adaptation of their dorsal vertebrae. The sail is possibly analogous (not homologous) to that of the Permian mammal-like reptile, Dimetrodon, which lived before the dinosaurs even appeared (these similarities are presumably due to parallel evolution).

The purpose of these sails is uncertain; scientists have proposed several hypotheses:

• Heat regulator. If the sail contained abundant blood vessels, the animal could have used the sail's large surface area to absorb heat. This would imply that the animal was only partly warm-blooded at best and lived in climates where nighttime temperatures were cool or low and the sky usually not cloudy. It is thought that Spinosaurus and Ouranosaurus both lived in or at the margins of an earlier version of the Sahara Desert, which could explain this. It is also possible that the sail was used to radiate excess heat from the body, rather than to collect it. Large animals, due to the relatively small ratio of surface area of their body compared to the overall volume (Haldane's principle), face far greater problems of dissipating excess heat at higher temperatures than gaining it at lower. Sails of these dinosaurs added considerably to the skin area of the body, with minimum increase of volume. Furthermore, if the sail was turned away from the sun, or positioned at a 90 degree angle towards a cooling wind, the animal would quite effectivly cool itself in the warm climate of Cretaceous Africa.
• Sexual display. Elaborate body structures of many modern-day animals usually serve to attract members of opposite sex during mating. It is quite possible that the sails of these dinosaurus were used for courtship, in a way similar to a peacock's tail. If this was the case, the sails may have been brightly colored, but this is purely speculative.
• Intimidating device. The sail was possibly used to intimidate rivals or frighten enemies, making the animal appear to be bigger than it was. The dinosaur could display its sail as a final warning signal, before it would resort to open attack, like modern-day rattlesnakes use their tail.

Finally, since things in nature rarely develop for a singular reason, it is quite possible that the sail combined all these functions, acting normally as a heat regulator, becoming a courting aid during the mating season, being used to cool itself and, on occasions, turning into an intimidating device when an animal was feeling threathened. Conjecture may even allow that the sail may have changed colour, during any of these functions.

Species and Specimens

Two species of Spinosaurus have been named: Spinosaurus aegyptiacus ("Egyptian spine lizard") and Spinosaurus marocannus ("Moroccan spine lizard"). S. marocannus was originally described by Russell as a new species based on the length of its neck vertebrae. However, several later authors considered the length of the neck vertebrae to be variable from individual to individual and therefore consider S. marocannus to be a synonym of S. aegyptiacus .

Five partial specimens of Spinosaurus have been found, the first having been destroyed during World War II (luckily, detailed drawings and descriptions of the specimen remain). The probable size of these individual spinosaurs can be estimated using comparison to known material from other spinosaurid dinosaurs.

• IPHG 1912 VIII 19 (Stromer, 1915) (destroyed)
  • Size: 17.4 m, 12-19 tons (subadult)
  • Material: (skull ~1.45 m) maxillary fragment, incomplete dentary (mandible ~1.34 m), nineteen teeth (62, 126 mm), two incomplete cervical vertebrae, seven dorsal vertebrae (190-210 mm), dorsal ribs, gastralia, eight caudal centra.
• CMN 50791 (Russell, 1996)
  • Material: mid cervical vertebra (195 mm), anterior dorsal neural arch, anterior dentary, mid dentary.
  • Note: holotype of Spinosaurus marocannus.
• MNHN SAM 124 (Taquet and Russell, 1998)
  • Size: ~17 m, ~11-18 tons (adult)
  • Material: (skull ~1.42 m) partial premaxillae, partial maxillae, vomers, dentary fragment.
• Office National des Mines nBM231 (Buffetaut and Ouaja, 2002)
  • Material: anterior dentary.
• MSNM V4047 (Dal Sasso et al., 2006)
  • Size: ~21 m, ~20-32 tons
  • Material: (skull ~1.75 m) premaxillae, partial maxillae, partial nasals

Rauhut (2003) suggested that Stromer's Spinosaurus holotype was a chimera, consisting of dorsal vertebrae from a carcharodontosaurid similar to Acrocanthosaurus and a dentary from a large theropod similar to Baryonyx. This analysis, however, was rejected by dal Sasso and most other researchers.

]Spinosaurus in Jurassic Park III==

Spinosaurus achieved widespread fame as the main antagonist in Jurassic Park III. It is portrayed as larger, more powerful and more vicious than Tyrannosaurus, epitomized by a scene in which the two resurrected predators battle and Spinosaurus emerges victorious. No such battle could ever have taken place in real life, since Spinosaurus and Tyrannosaurus lived thousands of miles and millions of years apart. The Spinosaurus depicted in the film was shown to be more aggressive and powerful than what most scientists believe the actual animal to be. Recent evidence suggest that Spinosaurus may have grown up to 60 feet long, although the size is misleading as the animal is relatively lightweight, and the tail may make up 50% or more of the total length.

Spinosaurus did, however, share its habitat with two other theropods that equalled or exceeded T. rex in size: the massive carnosaur Carcharodontosaurus and the large but relatively lightweight abelisaur Deltadromeus. Although the three gigantic predators probably occupied different ecological niches, they may have occasionally come into conflict over prey or territory.

References

• dal Sasso, C., S. Maganuco, E. Buffetaut and M. A. Mendez (2006). "New information on the skull of the enigmatic theropod Spinosaurus, with remarks on its sizes and affinities". Journal of Vertebrate Paleontology. 25(4): 888–896. {{cite journal}}: Check date values in: |year= (help)CS1 maint: multiple names: authors list (link) CS1 maint: year (link)
• Rauhut, O.W.M. (2003). "The interrelationships and evolution of basal theropod dinosaurs". Special Papers in Palaeontology. 69: 1–213. {{cite journal}}: Check date values in: |year= (help)CS1 maint: year (link)

External links

• "Spinosaurus". DinoData.
• "The Biggest Carnivore: Dinosaur History Rewritten

1. http://www.gavinrymill.com/dinosaurs/carnivores/

• Cretaceous dinosaurs
• Spinosaurids
• African dinosaurs
• Jurassic Park species