Misplaced Pages

This is an old revision of this page, as edited by Danny Bierek (talk | contribs) at 09:26, 29 May 2009 (→Origin of life: This link referred to Origin of Life on Earth, while this article refers to life in general. This is worth mentioning, but maybe somewhere else.). The present address (URL) is a permanent link to this revision, which may differ significantly from the current revision.

Life (Biota)
Life on a rocky peak in the Waitakere Ranges
Scientific classification
Domains and Kingdoms
Life on Earth: Non-cellular life (viruses) ** Cellular life Bacteria Archaea Eukarya Protista Fungi Plantae Animalia

Life is a characteristic that distinguishes objects that have self-sustaining biological processes ("alive," "living"), from those which do not —either because such functions have ceased (death), or else because they lack such functions and are classified as "inanimate."

In biology, "life" (cf. biota) is a quantitative and qualitative characteristic of all functioning organisms — (excluding viruses, cf. "non-cellular life") —such that exhibit continued self-sustaining (biological) processes. A diverse array of living organisms (life forms) can be found in the biosphere on Earth. Properties common to these organisms—plants, animals, fungi, protists, archaea, and bacteria — are a carbon- and water-based cellular form with complex organization and heritable genetic information. They undergo metabolism, maintain homeostasis, possess a capacity to grow, respond to stimuli, reproduce and, through natural selection, adapt to their environment in successive generations. More complex living organisms can communicate through various means.

Definitions

To define life in unequivocal terms is still a challenge for scientists, as the definition must be sufficiently broad that would encompass all life with which we are familiar. It should be sufficiently general that, with it, scientists would not miss life that may be fundamentally different from earthly life. In addition, defining life requires measurable terms, and when derived from analysis of known organisms, life is usually defined at cellular level.

Biology

The consensus is that life is a characteristic of organisms that exhibit all or most of the following phenomena:

1. Homeostasis: Regulation of the internal environment to maintain a constant state; for example, electrolyte concentration or sweating to reduce temperature.
2. Organization: Being structurally composed of one or more cells, which are the basic units of life.
3. Metabolism: Transformation of energy by converting chemicals and energy into cellular components (anabolism) and decomposing organic matter (catabolism). Living things require energy to maintain internal organization (homeostasis) and to produce the other phenomena associated with life.
4. Growth: Maintenance of a higher rate of anabolism than catabolism. A growing organism increases in size in all of its parts, rather than simply accumulating matter.
5. Adaptation: The ability to change over a period of time in response to the environment. This ability is fundamental to the process of evolution and is determined by the organism's heredity as well as the composition of metabolized substances, and external factors present.
6. Response to stimuli: A response can take many forms, from the contraction of a unicellular organism to external chemicals, to complex reactions involving all the senses of higher animals. A response is often expressed by motion, for example, the leaves of a plant turning toward the sun (phototropism) and by chemotaxis.
7. Reproduction: The ability to produce new individual organisms either asexually, from a single parent organism, or sexually, from at least two parent organisms.

Template:Fixbunching

Template:Fixbunching

Template:Fixbunching

Template:Fixbunching

Proposed

To reflect the minimum phenomena required, some have proposed other biological definitions of life:

1. Living things are systems that tend to respond to changes in their environment, and inside themselves, in such a way as to promote their own continuation.
2. A network of inferior negative feedbacks (regulatory mechanisms) subordinated to a superior positive feedback (potential of expansion, reproduction).
3. A systemic definition of life is that living things are self-organizing and autopoietic (self-producing). Variations of this definition include Stuart Kauffman's definition as an autonomous agent or a multi-agent system capable of reproducing itself or themselves, and of completing at least one thermodynamic work cycle.

Viruses

Viruses are most often considered replicators rather than forms of life. They have been described as "organisms at the edge of life", since they possess genes, evolve by natural selection, and replicate by creating multiple copies of themselves through self-assembly. However, viruses do not metabolise and require a host cell to make new products. Virus self-assembly within host cells has implications for the study of the origin of life, as it may support the hypothesis that life could have started as self-assembling organic molecules.

Biophysics

Biophysicists have also commented on the nature and qualities of life forms—notably that they function on negative entropy. In more detail, according to physicists such as John Bernal, Erwin Schrödinger, Eugene Wigner, and John Avery, life is a member of the class of phenomena which are open or continuous systems able to decrease their internal entropy at the expense of substances or free energy taken in from the environment and subsequently rejected in a degraded form (see: entropy and life).

Early theories about life

Materialism

Some of the earliest theories of life were materialist, holding that all that exists is matter, and that all life is merely a complex form or arrangment of matter. Empedocles (430 B.C.) argued that every thing in the universe is made up of a combination of four eternal 'elements' or 'roots of all': earth, water, air, and fire. All change is explained by the arrangement and rearrangement of these four elements. The various forms of life are caused by an appropriate mixture of elements. For example, growth in plants is explained by the natural downward movement of earth and the natural upward movement of fire.

Democritus (460 B.C.), the disciple of Leucippus, thought that the essential characteristic of life is having a soul (psychê). In common with other ancient writers, he used the term to mean the principle of living things that causes them to function as a living thing. He thought the soul was composed of fire atoms, because of the apparent connection between life and heat, and because fire moves. He also suggested that humans originally lived like animals, gradually developing communities to help one another, originating language, and developing crafts and agriculture.

In the scientific revolution of the seventeenth century, mechanistic ideas were revived by philosophers like Descartes.

Hylomorphism

Hylomorphism is the theory (originating with Aristotle) that all things are a combination of matter and form. Aristotle was one of the first ancient writers to approach the subject of life in a scientific way. Biology was one of his main interests, and there is extensive biological material in his extant writings. According to him, all things in the material universe have both matter and form. The form of a living thing is its soul (Greek 'psyche', Latin 'anima'). There are three kinds of souls: the 'vegetative soul' of plants, which causes them to grow and decay and nourish themselves, but does not cause motion and sensation; the 'animal soul' which causes animals to move and feel; and the rational soul which is the source of consciousness and reasoning which (Aristotle believed) is found only in man. Each higher soul has all the attributes of the lower one. Aristotle believed that while matter can exist without form, form cannot exist without matter, and therefore the soul cannot exist without the body.

Consistent with this account is a teleological explanation of life. A teleological explanation accounts for phenomena in terms of their purpose or goal-directedness. Thus, the whiteness of the polar bear's coat is explained by its purpose of camouflage. The direction of causality is the other way round from materialistic science, which explains the consequence in terms of a prior cause. Most modern biologists now reject this functional view in terms of a material and causal one: biological features are to be explained not by looking forward to future optimal results, but by looking backwards to the past evolutionary history of a species, which led to the natural selection of the features in question.

Vitalism

Vitalism is the belief that the life-principle is essentially immaterial. This originated with Stahl, and held sway until the middle of the nineteenth century. It appealed to philosophers such as Henri Bergson, Nietzsche, Wilhelm Dilthey , anatomists like Bichat, and chemists like Liebig.

Vitalism underpinned the idea of a fundamental separation of 'organic' and inorganic material, and the belief that organic material can only be derived from living things. This was disproved in 1828 when Wöhler prepared urea from inorganic materials. This so-called Wöhler synthesis is considered the starting point of modern organic chemistry. It is of great historical significance because for the first time an organic compound was produced from inorganic reactants.

Later, Helmholtz, anticipated by Mayer, demonstrated that no energy is lost in muscle movement, suggesting that there were no vital forces necessary to move a muscle. These empirical results led to the abandonment of scientific interest in vitalistic theories, although the belief lingered on in non-scientific theories such as Homeopathy, which interprets diseases and sickness as caused by disturbances in a hypothetical vital force or life force.

Origin of life

For religious beliefs about the creation of life, see creation myth.

Although it has not been pinpointed exactly, evidence suggests that life on Earth has existed for about 3.7 billion years. All known life forms share fundamental molecular mechanisms, and based on these observations, theories on the origin of life attempt to find a mechanism explaining the formation of a primordial single cell organism from which all life originates. There are many different hypotheses regarding the path that might have been taken from simple organic molecules via pre-cellular life to protocells and metabolism. Many models fall into the "genes-first" category or the "metabolism-first" category, but a recent trend is the emergence of hybrid models that do not fit into either of these categories.

There is no scientific consensus as to how life originated and all proposed theories are highly speculative. However, most currently accepted scientific models build in one way or another on the following theories:

1. Plausible pre-biotic conditions result in the creation of the basic small molecules of life. This was demonstrated in the Miller-Urey experiment, and in the work of Sidney Fox.
2. Phospholipids spontaneously form lipid bilayers, the basic structure of a cell membrane.
3. Procedures for producing random RNA molecules can produce ribozymes, which are able to produce more of themselves under very specific conditions.

Classification of life

Traditionally, people have divided organisms into the classes of plants and animals, based mainly on their ability of movement. The first known attempt to classify organisms was conducted by the Greek philosopher Aristotle (384-322 BC). He classified all living organisms known at that time as either a plant or an animal. Aristotle distinguished animals with blood from animals without blood (or at least without red blood), which can be compared with the concepts of vertebrates and invertebrates respectively. He divided the blooded animals into five groups: viviparous quadrupeds (mammals), birds, oviparous quadrupeds (reptiles and amphibians), fishes and whales. The bloodless animals were also divided into five groups: cephalopods, crustaceans, insects (which also included the spiders, scorpions, and centipedes, in addition to what we now define as insects), shelled animals (such as most molluscs and echinoderms) and "zoophytes". Though Aristotle's work in zoology was not without errors, it was the grandest biological synthesis of the time and remained the ultimate authority for many centuries after his death.

The exploration of the American continent revealed large numbers of new plants and animals that needed descriptions and classification. In the latter part of the 16th century and the beginning of the 17th, careful study of animals commenced and was gradually extended until it formed a sufficient body of knowledge to serve as an anatomical basis for classification.

In the late 1740s, Carolus Linnaeus introduced his method, still used, to formulate the scientific name of every species. Linnaeus took every effort to improve the composition and reduce the length of the many-worded names by abolishing unnecessary rhetoric, introducing new descriptive terms and defining their meaning with an unprecedented precision. By consistently using his system, Linnaeus separated nomenclature from taxonomy. This convention for naming species is referred to as binomial nomenclature.

The fungi were originally treated as plants. For a short period Linnaeus had placed them in the taxon Vermes in Animalia. He later placed them back in Plantae. Copeland classified the Fungi in his Protoctista, thus partially avoiding the problem but acknowledged their special status. The problem was eventually solved by Whittaker, when he gave them their own kingdom in his five-kingdom system. As it turned out, the fungi are more closely related to animals than to plants.

As new discoveries enabled us to study cells and microorganisms, new groups of life were revealed, and the fields of cell biology and microbiology were created. These new organisms were originally described separately in protozoa as animals and protophyta/thallophyta as plants, but were united by Haeckel in his kingdom protista, later the group of prokaryotes were split off in the kingdom Monera, eventually this kingdom would be divided in two separate groups, the Bacteria and the Archaea, leading to the six-kingdom system and eventually to the current three-domain system. The classification of eukaryotes is still controversial, with protist taxonomy especially problematic.

As microbiology, molecular biology and virology developed, non-cellular reproducing agents were discovered, such as viruses and viroids. Sometimes these entities are considered to be alive but others argue that viruses are not living organisms since they lack characteristics such as cell membrane, metabolism and do not grow or respond to their environments. Viruses can however be classed into "species" based on their biology and genetics but many aspects of such a classification remain controversial.

Since the 1960s a trend called cladistics has emerged, arranging taxa in an evolutionary or phylogenetic tree. It is unclear, should this be implemented, how the different codes will coexist.

Extraterrestrial life

Earth is the only planet in the universe known to harbour life. The Drake equation, which relates the number of extraterrestrial civilizations in our galaxy with which we might come in contact, has been used to discuss the probability of life elsewhere, but scientists disagree on many of the values of variables in this equation. Depending on those values, the equation may either suggest that life arises frequently or infrequently.

Panspermia and exogenesis are theories proposing that life originated elsewhere in the universe and was subsequently transferred to Earth in the form of spores perhaps via meteorites, comets or cosmic dust. However those theories do not help explain the ultimate origin of life.

See also

References

1. ^ Koshland Jr, Daniel E. (March 22, 2002). "The Seven Pillars of Life". Science. Vol. 295. (no. 5563): pp. 2215 - 2216. doi:10.1126/science.1068489. Retrieved 2009-05-25. {{cite journal}}: |issue= has extra text (help); |pages= has extra text (help); |volume= has extra text (help); Cite has empty unknown parameter: |coauthors= (help)
2. The American Heritage Dictionary of the English Language, 4th edition, published by Houghton Mifflin Company, via Answers.com:
   • "The property or quality that distinguishes living organisms from dead organisms and inanimate matter, manifested in functions such as metabolism, growth, reproduction, and response to stimuli or adaptation to the environment originating from within the organism."
   • "The characteristic state or condition of a living organism."
3. "Merriam-Webster Dictionary". Merriam-Webster Dictionary. Retrieved 2009-05-25.
4. "organism". Chambers 21st Century Dictionary (online ed.). 1999.
5. Defining Life : Astrobiology Magazine - earth science - evolution distribution Origin of life universe - life beyond
6. Defining Life, Explaining Emergence
7. Nealson KH, Conrad PG (1999). "Life: past, present and future" (PDF). Philos. Trans. R. Soc. Lond., B, Biol. Sci. 354 (1392): 1923–39. doi:10.1098/rstb.1999.0532. PMC 1692713. PMID 10670014. {{cite journal}}: Unknown parameter |month= ignored (help)
8. Davison, Paul G. "How to Define Life". The University of North Alabama. Retrieved 2008-10-17.
9. ^ Witzany, G. (2007). The Logos of the Bios 2. Bio-Communication. Helsinki, Umweb.
10. Korzeniewski, Bernard (2001). "Cybernetic formulation of the definition of life". Journal of Theoretical Biology. 2001 April 7. 209 (3) pp. 275–86.
11. 2004, "Autonomous Agents", in John D. Barrow, P.C.W. Davies, and C.L. Harper Jr., eds., Science and Ultimate Reality: Quantum Theory, Cosmology, and Complexity, Cambridge University Press.
12. Rybicki EP (1990) "The classification of organisms at the edge of life, or problems with virus systematics." S Aft J Sci 86:182–186
13. Holmes EC (2007). "Viral evolution in the genomic age". PLoS Biol. 5 (10): e278. doi:10.1371/journal.pbio.0050278. PMC 1994994. PMID 17914905. Retrieved 2008-09-13. {{cite journal}}: Unknown parameter |month= ignored (help)CS1 maint: unflagged free DOI (link)
14. Koonin EV, Senkevich TG, Dolja VV (2006). "The ancient Virus World and evolution of cells". Biol. Direct. 1: 29. doi:10.1186/1745-6150-1-29. PMC 1594570. PMID 16984643. Retrieved 2008-09-14.{{cite journal}}: CS1 maint: multiple names: authors list (link) CS1 maint: unflagged free DOI (link)
15. Rybicki, Ed (November 1997). "Origins of Viruses". Retrieved 2009-04-12.
16. Schrödinger, Erwin (1944). What is Life?. Cambridge University Press. ISBN 0-521-42708-8.
17. Margulis, Lynn (1995). What is Life?. University of California Press. ISBN 0-520-22021-8. {{cite book}}: Unknown parameter |coauthors= ignored (|author= suggested) (help)
18. Lovelock, James (2000). Gaia – a New Look at Life on Earth. Oxford University Press. ISBN 0-19-286218-9.
19. Avery, John (2003). Information Theory and Evolution. World Scientific. ISBN 9812383999.
20. SEP:
21. SEP
22. Ibidem
23. Aristotle, De Anima, Book II
24. Introduction to Ancient Philosophy, Don Marietta, p.104.
25. "History of life through time". University of California Museum of Paleontology.
26. Coveney, Peter V.; Philip W. Fowler. "Modelling biological complexity: a physical scientist's perspective". Journal of the Royal Society Interface. 2005. 2 (4) pp. 267–280. doi:10.1098/rsif.2005.0045
27. "Aristotle -biography". University of California Museum of Paleontology. Retrieved 2008-10-20. {{cite news}}: Cite has empty unknown parameter: |coauthors= (help)
28. Knapp S, Lamas G, Lughadha EN, Novarino G (2004). "Stability or stasis in the names of organisms: the evolving codes of nomenclature". Philosophical transactions of the Royal Society of London. Series B, Biological sciences. 359 (1444): 611–22. doi:10.1098/rstb.2003.1445. PMC 1693349. PMID 15253348. {{cite journal}}: Unknown parameter |month= ignored (help)CS1 maint: multiple names: authors list (link)
29. ^ Copeland, H. F. (1938). "The Kingdoms of Organisms". Quarterly Review of Biology. 13 (4): 383. doi:10.1086/394568. Cite error: The named reference "Copeland1938" was defined multiple times with different content (see the help page).
30. Whittaker RH (1969). "New concepts of kingdoms or organisms. Evolutionary relations are better represented by new classifications than by the traditional two kingdoms". Science. 163 (863): 150–60. doi:10.1126/science.163.3863.150. PMID 5762760. {{cite journal}}: Unknown parameter |month= ignored (help)
31. ^ Woese, C.; Kandler, O.; Wheelis, M. (1990). "Towards a natural system of organisms:proposal for the domains Archaea, Bacteria, and Eucarya". Proceedings of the National Academy of Sciences of the United States of America. 87 (12): 4576–9. Bibcode:1990PNAS...87.4576W. doi:10.1073/pnas.87.12.4576. PMC 54159. PMID 2112744.
32. Adl SM, Simpson AG, Farmer MA; et al. (2005). "The new higher level classification of eukaryotes with emphasis on the taxonomy of protists". J. Eukaryot. Microbiol. 52 (5): 399–451. doi:10.1111/j.1550-7408.2005.00053.x. PMID 16248873. {{cite journal}}: Explicit use of et al. in: |author= (help)CS1 maint: multiple names: authors list (link)
33. Van Regenmortel MH (2007). "Virus species and virus identification: past and current controversies". Infection, genetics and evolution : journal of molecular epidemiology and evolutionary genetics in infectious diseases. 7 (1): 133–44. doi:10.1016/j.meegid.2006.04.002. PMID 16713373. {{cite journal}}: Unknown parameter |month= ignored (help)
34. Pennisi E (2001). "Taxonomy. Linnaeus's last stand?". Science. 291 (5512). New York, N.Y.: 2304–7. doi:10.1126/science.291.5512.2304. PMID 11269295. {{cite journal}}: Unknown parameter |month= ignored (help)
35. Linnaeus, C. (1735). Systemae Naturae, sive regna tria naturae, systematics proposita per classes, ordines, genera & species.
36. Haeckel, E. (1866). Generelle Morphologie der Organismen. Reimer, Berlin.
37. Chatton, É. (1925). "Pansporella perplexa. Réflexions sur la biologie et la phylogénie des protozoaires". Annales des Sciences Naturelles - Zoologie et Biologie Animale. 10-VII: 1–84.
38. Whittaker, R. H. (January 1969). "New concepts of kingdoms of organisms". Science. 163 (3863): 150–60. Bibcode:1969Sci...163..150W. doi:10.1126/science.163.3863.150. PMID 5762760.
39. Cavalier-Smith, T. (1998). "A revised six-kingdom system of life". Biological Reviews. 73 (3): 203–66. doi:10.1111/j.1469-185X.1998.tb00030.x. PMID 9809012. S2CID 6557779.
40. Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M.; Thuesen, Erik V. (2015). "A higher level classification of all living organisms". PLOS ONE. 10 (4): e0119248. Bibcode:2015PLoSO..1019248R. doi:10.1371/journal.pone.0119248. PMC 4418965. PMID 25923521.

Further reading

• Kauffman, Stuart. The Adjacent Possible: A Talk with Stuart Kauffman
• Nealson KH, Conrad PG (1999). "Life: past, present and future". Philos. Trans. R. Soc. Lond., B, Biol. Sci. 354 (1392): 1923–39. doi:10.1098/rstb.1999.0532. PMC 1692713. PMID 10670014. {{cite journal}}: Unknown parameter |month= ignored (help)
• Walker, Martin G. LIFE! Why We Exist...And What We Must Do to Survive Dog Ear Publishing, 2006, ISBN 1598582437

External links

• Wikispecies - a free directory of life
• "The Adjacent Possible: A Talk with Stuart Kauffman"
• Stanford Encyclopedia of Philosophy entry
• Life under extreme conditions An in depth look at how life can form under the most extreme conditions.

• Life
• Biology
• Biological systems