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Spinosaurus

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Spinosaurus
Temporal range: mid Cretaceous
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Sauropsida
Superorder: Dinosauria
Order: Saurischia
Suborder: Theropoda
Superfamily: Megalosauroidea
Family: Spinosauridae
Genus: Spinosaurus
Stromer, 1915
Species
  • S. aegyptiacus (type)
  • ?S. marocannus

Spinosaurus (meaning 'spine lizard') was a theropod dinosaur genus that lived in what is now Egypt, from the Albian to early Cenomanian stages of the Cretaceous Period, about 95 to 93 million years ago. According to a study by dal Sasso et al. (2006), it was the largest of all carnivorous dinosaurs by a significant margin, even larger than Tyrannosaurus rex and Giganotosaurus. Spinosaurus was the longest theropod, measuring 16 to 18 metres (53.3 to 60 feet) long and standing at 5 to 6 metres (16.6 to 20 feet) in height. Spinosaurus reached weights up to 9 tons, although further comparisons with related species suggest that adults might have reached sizes up to 20 tons and 21 m (70 ft) in length.

The distinctive spines of these animals (large bones extending from the vertebrae) grew up to 2 m (6.6ft) long and were likely to have had skin stretching between them, forming a sail-like structure, although some have suggested they were covered in muscle and formed a hump or ridge.

Spinosaurus provides the name of a family of dinosaurs, the Spinosauridae, of which other members include Angaturama (probably synonymous with Irritator), Baryonyx, Irritator, Suchomimus, and Siamosaurus.

Description: Spinosaurus was a huge gracile theropod dinosaur from the Middle Cretaceous period in North Africa and possibly Argentina. It was the largest meat eating dinosaur that ever lived which could reach a length of 66-69 feet and weigh at 12-15 tons. It was a semi-aquatic dinosaur capable of diving to depths of 100 feet while searching for its favorite food: coelacanths. An addition to being a swimmer, it was a capable runner and hunter of medium sized sauropods, ornithropods (like Ouranosaurus), and sometimes smaller theropods. Its jaws were narrow and weak in comparison to other theropods but the slender lower jaw was anchored by powerful neck muscles which reinforced the jaw muscles for a more powerful bite. Its teeth varied in size from 4 to 6 inches in length but were smoothed edged which was ideal for severing the spinal cord of a young sauropod. The strong, three fingered arms bore huge claws which acted like grappling hooks to hold onto the prey. The extremely long tail acted like a balancing rod which allowed the animal to make a sharp turn while chasing prey (see also Cheetah). The tail also acts as a weapon to knock their prey of their feet. Another useful purpose for such a long tail is that it makes a good propeller when swimming in the lagoons. The most distinctive feature of Spinosaurus was its sail that stood 6 feet, 6 inches. No one knows the true purpose of the so-called sail but it was probably used to regulate body temperature. Heat regulation seems most logical suggested that this animal was cold blooded.

Diet

It is unclear whether Spinosaurus was a cursorial predator or a fisher, as indicated by its elongated jaws, conical teeth and raised nostrils. The only direct evidence for spinosaur diet comes from related European and South American species. Baryonyx was found with both fish scales and bones from juvenile Iguanodon in its stomach, while a tooth embedded in a South American pterosaur bone suggests that spinosaurs occasionally preyed on these flying archosaurs. Spinosaurus was likely to have been a more generalized and opportunistic predator, possibly a Cretaceous equivalent of large grizzly bears, being biased toward fishing, although it undoubtedly took many kinds of small-to-medium-sized prey.

Diet: Spinosaurus was a hunter and fisherman who adapted hunt in both land and water. On land it hunted like a tyrannosaur by blending itself with the environment using the climatophores (see chameleon, octopus, or cuttlefish) in its scales. Than it charges into the open usually preying on the smaller dinosaurs. In the water it swam like a crocodile and was capable of chasing the sharks and coelacanths. It was a fast swimmer and can move as fast as 25 miles an hour when chasing their underwater quarry. On land it was fast for an animal that weighs 15 tons; with its sleek built, gracile legs, and long tail it could run as fast as 30 miles an hour while in pursuit.

Species and Specimens

Two species of Spinosaurus have been named: Spinosaurus aegyptiacus ("Egyptian spine lizard") and Spinosaurus marocannus ("Moroccan spine lizard"). S. marocannus was originally described by Russell as a new species based on the length of its neck vertebrae. However, several later authors considered the length of the neck vertebrae to be variable from individual to individual and therefore consider S. marocannus to be a synonym of S. aegyptiacus .

Five partial specimens of Spinosaurus have been found, the first having been destroyed during World War II (luckily, detailed drawings and descriptions of the specimen remain). The probable size of these individual spinosaurs can be estimated using comparison to known material from other spinosaurid dinosaurs.

  • IPHG 1912 VIII 19 (Stromer, 1915) (destroyed)
    • Size: 17.4 m, 12-19 tons (subadult)
    • Material: (skull ~1.45 m) maxillary fragment, incomplete dentary (mandible ~1.34 m), nineteen teeth (62, 126 mm), two incomplete cervical vertebrae, seven dorsal vertebrae (190-210 mm), dorsal ribs, gastralia, eight caudal centra.
  • CMN 50791 (Russell, 1996)
    • Material: mid cervical vertebra (195 mm), anterior dorsal neural arch, anterior dentary, mid dentary.
    • Note: holotype of Spinosaurus marocannus.
  • MNHN SAM 124 (Taquet and Russell, 1998)
    • Size: ~17 m, ~11-18 tons (adult)
    • Material: (skull ~1.42 m) partial premaxillae, partial maxillae, vomers, dentary fragment.
  • Office National des Mines nBM231 (Buffetaut and Ouaja, 2002)
    • Material: anterior dentary.
  • MSNM V4047 (Dal Sasso et al., 2006)
    • Size: ~21 m, ~20-32 tons
    • Material: (skull ~1.75 m) premaxillae, partial maxillae, partial nasals

Rauhut (2003) suggested that Stromer's Spinosaurus holotype was a chimera, consisting of dorsal vertebrae from a carcharodontosaurid similar to Acrocanthosaurus and a dentary from a large theropod similar to Baryonyx. This analysis, however, was rejected by dal Sasso and most other researchers.

Spinosaurus in Jurassic Park III

File:JurassicParkSPINO.jpg
A computer generated Spinosaurus from Jurassic Park III.
File:JurassicParkSPINO2.jpg
The animatronic Spinosaurus from JP3.

Spinosaurus achieved widespread fame as the main antagonist in Jurassic Park III. It is portrayed as larger, more powerful and more vicious than Tyrannosaurus, epitomized by a scene in which the two resurrected predators battle and Spinosaurus emerges victorious. No such battle could ever have taken place in real life, since Spinosaurus and Tyrannosaurus lived thousands of miles and millions of years apart.

Spinosaurus did, however, share its habitat with two other theropods that equalled or exceeded T. rex in size: the massive carnosaur Carcharodontosaurus and the large but relatively lightweight abelisaur Deltadromeus. Although the three gigantic predators probably occupied different ecological niches, they may have occasionally come into conflict over prey or territory.

References

  • dal Sasso, C., S. Maganuco, E. Buffetaut and M. A. Mendez (2006). "New information on the skull of the enigmatic theropod Spinosaurus, with remarks on its sizes and affinities". Journal of Vertebrate Paleontology. 25(4): 888–896. {{cite journal}}: Check date values in: |year= (help)CS1 maint: multiple names: authors list (link) CS1 maint: year (link)
  • Rauhut, O.W.M. (2003). "The interrelationships and evolution of basal theropod dinosaurs". Special Papers in Palaeontology. 69: 1–213. {{cite journal}}: Check date values in: |year= (help)CS1 maint: year (link)

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