Alcobaça Formation | |
---|---|
Stratigraphic range: Kimmeridgian ~156.9–152.5 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ | |
Type | Geological formation |
Sub-units | Camadas de Guimarota |
Underlies | Lourinhã Formation |
Overlies | Montejunto Formation |
Lithology | |
Primary | Marl |
Other | Mudstone, coal, siltstone, limestone, sandstone |
Location | |
Coordinates | 39°36′N 9°00′W / 39.6°N 9.0°W / 39.6; -9.0 |
Approximate paleocoordinates | 33°42′N 1°36′E / 33.7°N 1.6°E / 33.7; 1.6 |
Region | Estremadura, Leiria, Lisboa |
Country | Portugal |
Extent | Lusitanian Basin |
Type section | |
Named for | Alcobaça, Portugal |
Named by | Choffat |
Year defined | 1885 |
Alcobaça Formation (Portugal) |
The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit, is a geological formation in Portugal. It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of information on the diversity of Late Jurassic mammals. Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine.
Vertebrate paleofauna
Dinosaur eggs are geographically located in Lisbon District, Portugal. Dinosaur tracks are geographically located in Leiria District, Portugal.
Chondrichthyes
Chondrichthyes of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
Hybodus | Indeterminate | Guimarota | Isolated Teeth | |||
Asteracanthus | A. biformatus | Guimarota | isolated teeth, fin-spine and a head-spine | |||
Polyacrodus | Indeterminate | Guimarota | Isolated teeth and a fin-spine | |||
Orectolobiformes | Indeterminate | Guimarota | One isolated tooth | |||
Batomorphii | Indeterminate | Guimarota | Isolated teeth |
Osteichthyes
Osteichthyes of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
Macromesodon | Indeterminate | Guimarota | Isolated teeth | |||
Proscinetes | Indeterminate | Guimarota | Isolated teeth and prearticulars | |||
"Lepidotes" | Indeterminate | Guimarota | isolated lateral teeth, lower jaws, ganoid scales and isolated skull bones | Two species of different size can be distinguished from the fossil material | ||
Ionoscopidae | Indeterminate | Guimarota | Isolated teeth | |||
Macrosemiidae | Indeterminate | Guimarota | Isolated grasping tooth | |||
Caturus? | Indeterminate | Guimarota | Isolated Teeth and a partial skull | |||
Pachycormidae | Indeterminate | Guimarota |
Amphibians
Albanerpetontids are one of the most numerous faunal components of the Guimarota mine with around 9000 isolated remains, including parts of the skull and limbs. Remains of indeterminate salamanders and frogs are also present but far rarer.
Amphibians of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
cf. Celtedens | Indeterminate | Porto Pinheiro | Albanerpetontid | |||
cf. Marmorerpeton | Indeterminate | Guimarota | Indeterminate salamander | |||
Discoglossidae | Indeterminate | Guimarota | Indeterminate frog |
Choristoderes
Choristoderes of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Cteniogenys | Indeterminate | Guimarota |
Ornithischians
Indeterminate euornithopod remains located in Lisbon District. Indeterminate stegosaurid remains present in Lisbon District.
Ornithischians of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | District | Stratigraphic position | Abundance | Notes | Images |
Dacentrurus | D. armatus | Lisboa | ||||
Ornithopoda | Indeterminate | Lisboa | ||||
Stegosaurus | S. cf. ungulatus | "...one tooth, five cervical vertebrae (including the axis), five dorsal vertebrae, cervical and dorsal ribs, three caudal vertebrae, chevrons, preacetabular process of the left ilium, right tibia, fibula, astragalus and , a cervical plate, and several fragments of plates." | ||||
Trimucrodon | T. cuneatus | Lisboa | Later determined to be indeterminate ornithischian remains. |
Saurischians
Indeterminate sauropod remains located in Leiria and Lisboa.
Saurischians of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | District | Stratigraphic position | Abundance | Notes | Images |
Allosaurus | Indeterminate | Leiria | ||||
Aviatyrannis | A. jurassica | Guimarota | ||||
Lusotitan | L. atalaiensis | Lisboa | ||||
Ceratosaurus | Indeterminate | Lisboa | ||||
Dinheirosaurus | D. lourinhanensis | Lisboa | " cervicals, dorsals, ribs, limb elements." | |||
Lourinhasaurus | L. alenquerensis | Lisboa | ||||
Megalosaurus | M. insignis | Lisboa | Later determined to be indeterminate theropod remains. |
Pterosaurs
Pterosaurs of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
aff. Rhamphorhynchus | Indeterminate | Guimarota | Isolated Fragments | |||
Pterodactyloidea | Indeterminate | Guimarota | Teeth |
Turtles
Turtles of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Peltochelys | P. duchastelli | Guimarota | ||||
Platychelys | Indeterminate | Guimarota | ||||
Pleurosternidae | Indeterminate | Guimarota |
Lepidosauromorphs
Lepidosauromorphs of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Becklesius | B. hoffstetteri | Guimarota | Paramacellodid lizard | |||
Dorsetisaurus | D. purbeckensis | Guimarota | Anguimorph lizard | |||
Marmoretta | Indeterminate | Guimarota | Basal Lepidosauromorph | |||
Portugalophis | P. lignites | Guimarota | Stem-snake | |||
Saurillodon | S. proraformis | Guimarota | A lizard | |||
Saurillus | S. henkeli, cf. obtusus | Guimarota | A lizard | |||
cf. Opisthias | Indeterminate | Andrés, Pombal | A rhynchocephalian |
Crocodyliformes
Crocodyliformes of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Goniopholis | G. baryglyphaeus | Guimarota | A goniopholidid | |||
Knoetschkesuchus | K. guimarotae | Guimarota | An atoposaurid | |||
Lisboasaurus | L. mitracostatus | Guimarota | ||||
Machimosaurus | M. hugii | Guimarota | A marine machimosaurid |
Mammals
Mammals of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Bathmochoffatia | B. hapax | Guimarota | A paulchoffatiid multituberculate | |||
Bernardodon | B. sp, B. atlanticus | Porto Pinheiro | A pinheirodontid multituberculate | |||
Drescheratherium | D. acutum | Guimarota | A paurodontid dryolestidan | |||
Dryolestes | D. leirensis | Guimarota | A dryolestid dryolestidan | |||
Ecprepaulax | E. anomala | Porto Pinheiro | A pinheirodontid multituberculate | |||
Guimarotodon | G. leiriensis | Guimarota | A paulchoffatiid multituberculate | |||
Guimarotodus | G. inflatus | Guimarota | A dryolestid dryolestidan | |||
Haldanodon | H. exspectatus | Guimarota | A docodont | |||
Henkelodon | H. naias, H. sp | Guimarota | A paulchoffatiid multituberculate | |||
Henkelotherium | H. guimarotae | Guimarota | A dryolestidan | |||
Iberodon | I. quadrituberculatus | Porto Pinheiro | A pinheirodontid multituberculate | |||
Kielanodon | K. hopsoni | Guimarota | A paulchoffatiid multituberculate | |||
Krebsotherium | K. lusitanicum | Guimarota | A dryolestid dryolestidan | |||
Kuehneodon | K. guimarotensis, K. dietrichi, K. simpsoni, K. uniradiculatus, K. sp | Guimarota | A paulchoffatiid multituberculate | |||
Laolestes | L. andresi | Porto Pinheiro | A dryolestid dryolestidan | |||
Meketibolodon | M. robustus | Guimarota | A paulchoffatiid multituberculate | |||
Meketichoffatia | M. krausei, M. sp | Guimarota | A paulchoffatiid multituberculate | |||
Nanolestes | N. krusati | Porto Pinheiro | A cladotherian | |||
N. drescherae | Guimarota | |||||
Paulchoffatia | P. delgadoi | Guimarota | A paulchoffatiid multituberculate | |||
Pinheirodon | P. sp, P. vastus, P. pygmaeus | Porto Pinheiro | A pinheirodontid multituberculate | |||
Plesiochoffatia | P. thoas, P. peperethos | Guimarota | A paulchoffatiid multituberculate | |||
Portopinheirodon | P. asymmetricus | Porto Pinheiro | A dryolestid dryolestidan | |||
Priacodon | Indeterminate | Porto Pinheiro | A triconodontid | |||
Proalbionbaatar | P. plagiocyrtus | Guimarota | A multituberculate of uncertain placement | |||
Pseudobolodon | P. dryas, P krebsi, P. oreas | Guimarota | A paulchoffatiid multituberculate | |||
Renatodon | R. amalthea | Guimarota | A paulchoffatiid multituberculate | |||
? Tinodon | Indeterminate | Porto Pinheiro | A tinodontid symmetrodont | |||
Xenachoffatia | X. oinopion | Guimarota | A paulchoffatiid multituberculate | |||
Paulchoffatiidae | Indeterminate | Guimarota | 4 distinct genera with 7 species | |||
Pinheirodontidae | Indeterminate | Porto Pinheiro |
Color key
|
Notes Uncertain or tentative taxa are in small text; |
Invertebrate Paleofauna
Ostracods
Ostracods of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
Cetacella | C. armata | Cetacella is a typical component of Late Jurassic brackish water environments in Europe. Member of the Cypridacea | ||||
C. inermis | ||||||
C. striata | ||||||
Theriosynoecum | T. wyomingense | Member of the Cytheracea | ||||
Bisulcocypris | B. pahasapensis | Member of the Cytheracea | ||||
Timiriasevia | T. guimarotensis | Member of the Cytheracea | ||||
Poisia | P. bicostata | Member of the Cytheracea | ||||
P. clivosa | ||||||
Dicrorygima | D. reticulata | Member of the Cytheracea | ||||
Darwinula | D. leguminella | Member of the Darwinulacea |
Mollusca
Bivalves
Bivalves of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
Isognomon | I. rugosus | Guimarota | Dozens of well preserved shells sometimes making up entire stratigraphic layers | This genus is regarded as an indicator for brachyhaline water conditions and it was likely buried in its natural habitat | ||
"Unio" | "U." cf. alcobacensis | Guimarota | Dozens of well preserved shells with articulated valves | freshwater clam |
Gastropods
Gastropods of the Alcobaça Formation | ||||||
---|---|---|---|---|---|---|
Genus | Species | Location | Stratigraphic Position | Material | Notes | Images |
Teinostoma | Indeterminate | Guimarota | Shells | Boring traces of clionidid sponges on the shells of this taxon indicate a marine lifestyle | ||
Cryptaulax | Indeterminate | Guimarota | Shells | |||
Melampoides | M. jurassicus | Guimarota | Shells | Boring traces of clionidid sponges on the shells of this taxon indicate a marine lifestyle, modern relatives like Melampus also venture onto land, which can also be expected from M. jurassicus |
See also
References
- Mateus, O.; Dinis, J.; Cunha, P. P. (2017-09-28). "The Lourinhã Formation: the Upper Jurassic to lower most Cretaceous of the Lusitanian Basin, Portugal – landscapes where dinosaurs walked". Ciências da Terra / Earth Sciences Journal. 19 (1): 75–97. doi:10.21695/cterra/esj.v19i1.355. hdl:10316/79879. ISSN 2183-4431.
- Fürsich, Franz T.; Schneider, Simon; Werner, Winfried; Lopez-Mir, Berta; Pierce, Colm S. (2021-09-13). "Life at the continental–marine interface: palaeoenvironments and biota of the Alcobaça Formation (Late Jurassic, Central Portugal), with a formal definition of the unit appended". Palaeobiodiversity and Palaeoenvironments. 102 (2): 265–329. doi:10.1007/s12549-021-00496-x. ISSN 1867-1608.
- "Guimarota". Paleobiology Database. Retrieved 31 August 2018.
- "Porto Pinheiro, Lourinha". Paleobiology Database. Retrieved 31 August 2018.
- ^ "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
- ^ "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," in Weishampel, et al. (2004). Page 548.
- ^ Palmer, Douglas (September 2001). "M ARTIN , T. & K REBS , B. (eds) 2000. Guimarota. A Jurassic Ecosystem . 155 pp. München: Verlag Dr. Friedrich Pfeil. Price DM 120.00 (hard covers). ISBN 3 931516 80 6". Geological Magazine. 138 (5): 619–630. doi:10.1017/S0016756801316084. ISSN 0016-7568.
- Escaso, Fernando; Ortega, Francisco; Dantas, Pedro; Malafaia, Elisabete; Pimentel, Nuno L.; Pereda-Suberbiola, Xabier; Sanz, José Luis; Kullberg, José Carlos; Kullberg, María Carla (2006-12-23). "New evidence of shared dinosaur across Upper Jurassic Proto-North Atlantic: Stegosaurus from Portugal". Naturwissenschaften. 94 (5): 367–374. Bibcode:2007NW.....94..367E. doi:10.1007/s00114-006-0209-8. PMID 17187254. S2CID 10930309.
- "16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549.
- "Table 13.1," in Weishampel, et al. (2004). Page 265.
- Malafaia, E.; Ortega, F.; Escaso, F.; Dantas, P.; Pimentel, N.; Gasulla, J. M.; Ribeiro, B.; Barriga, F.; Sanz, J. L. (2010). "Vertebrate fauna at the Allosaurus fossil-site of Andrés (Upper Jurassic), Pombal, Portugal". Journal of Iberian Geology. 36 (2): 193–204. doi:10.5209/rev_JIGE.2010.v36.n2.7. ISSN 1698-6180.
Bibliography
- Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (2004), The Dinosauria, 2nd edition, Berkeley: University of California Press, pp. 1–880, ISBN 0-520-24209-2, retrieved 2019-02-21