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Sierra Negra giant tortoise
Chelonoidis guentheri
Conservation status
Critically Endangered  (IUCN 3.1)
CITES Appendix I (CITES)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Superfamily:	Testudinoidea
Family:	Testudinidae
Genus:	Chelonoidis
Species:	C. niger
Subspecies:	C. n. guentheri
Trinomial name
Chelonoidis niger guentheri (Baur, 1889)
Synonyms
Testudo güntheri Baur, 1889 Testudo wallacei Rothschild, 1902 Geochelone elephantopus guntheri Pritchard, 1967 Geochelone nigra guntheri Iverson, 1992 Geochelone guntheri Cisneros-Heredia, 2006

Chelonoidis niger guentheri, commonly known as the Sierra Negra giant tortoise or Günther's giant tortoise, is a subspecies of Galápagos tortoise endemic to the Galápagos archipelago in the eastern Pacific Ocean. The specific epithet guentheri honours zoologist Albert Günther.

Taxonomy

This tortoise is a subspecies of Chelonoidis niger, and is sometimes considered a distinct species.

Description

Male tortoises grow to about 120 cm and females to about 92 cm in length and may have either domed or flattened carapaces.

Behaviour

Feeding

The tortoises graze on shrubs and low-growing herbaceous vegetation, consuming herbs, grass, cacti, lichens, and fruit.

Breeding

Male tortoises compete with each other by extending their necks, gaping, biting and shell-bumping. They produce loud guttural noises while mating. Females lay clutches of usually 6–11 (occasionally up to 17) eggs. Juvenile tortoises tend to remain in the warmer, lowland, part of the subspecies' range for the first 10–15 years of their lives.

Distribution and habitat

The tortoise's range is limited to an area of about 800 km on the slopes of the Sierra Negra volcano, from near sea-level to an altitude of around 1,000 m, at the southern end of Isabela Island. There it inhabits deciduous and evergreen forests, dry grassland, introduced vegetation and agricultural land. The population is estimated to comprise some 400–700 mature individuals, a decline of 99% from an estimated 71,000 individuals prior to human contact with the islands.

Conservation

The subspecies is considered to be Critically Endangered. The population was historically overexploited for food and oil by sailors and settlers, and some poaching continues. Other threats include predation of eggs and hatchlings, as well as habitat degradation, by introduced animals, including dogs, rodents, pigs, goats and fire ants. Volcanic eruptions also threaten the tortoises and their habitat.

References

1. Cayot, L.J., Gibbs, J.P., Tapia, W. & Caccone, A. (2018). "Sierra Negra Giant Tortoise". IUCN Red List of Threatened Species. 2018. Retrieved 26 June 2021.{{cite journal}}: CS1 maint: multiple names: authors list (link)
2. "Appendices | CITES". cites.org. Retrieved 2022-01-14.
3. "Chelonoidis guntheri (BAUR, 1889)". Reptile Database. Peter Uetz and Jakob Hallermann. Retrieved 26 June 2021.
4. ^ Arteaga, A; Bustamante, L; Vieira, J; Guayasamin, JM (2020). Reptiles of Ecuador: Life in the middle of the world. Quito: Universidad Tecnológica Indoamereica.

v t e Tortoise family
Kingdom: Animalia Phylum: Chordata Class: Reptilia Subclass: Anapsida Order: Testudines Suborder: Cryptodira Superfamily: Testudinoidea Family: Testudinidae
Genera	Species of the tortoise family
Agrionemys	Russian tortoise
Aldabrachelys	Aldabra giant tortoise (subspecies: A. g. arnoldi †A. g. daudinii A. g. hololissa) †Aldabrachelys abrupta †Aldabrachelys grandidieri
Astrochelys	Angonoka tortoise Radiated tortoise †Astrochelys rogerbouri
Centrochelys	African spurred tortoise †Centrochelys atlantica †Centrochelys burchardi †Centrochelys vulcanica
Chelonoidis	Chaco tortoise Red-footed tortoise Yellow-footed tortoise †Chelonoidis alburyorum †Chelonoidis cubensis †Chelonoidis lutzae †Chelonoidis monensis †Chelonoidis sellovii †Chelonoidis sombrerensis
Galápagos tortoise Chelonoidis complex	Cerro Azul giant tortoise Chatham Island giant tortoise Eastern Santa Cruz tortoise Fernandina Island tortoise †Floreana Island tortoise Hood Island giant tortoise (Diego) Santa Fe Island tortoise †Pinta Island tortoise †(Lonesome George) Santiago Island giant tortoise Pinzón Island giant tortoise Sierra Negra giant tortoise Volcán Alcedo giant tortoise Volcán Darwin giant tortoise Volcán Wolf giant tortoise Western Santa Cruz tortoise
†Cheirogaster	†Cheirogaster bacharidisi †Cheirogaster gymnesica †Cheirogaster schafferi
Chersina	Angulate tortoise
Chersobius	Nama padloper Karoo padloper Speckled padloper
†Cylindraspis	†Domed Mauritius giant tortoise †Domed Rodrigues giant tortoise †Réunion giant tortoise †Saddle-backed Mauritius giant tortoise †Saddle-backed Rodrigues giant tortoise
Geochelone	Burmese star tortoise Indian star tortoise
Gopherus	Bolson tortoise Desert tortoise Goode's thornscrub tortoise Gopher tortoise Sonoran Desert tortoise Texas tortoise
†Hadrianus	†Hadrianus corsoni †Hadrianus majusculus †Hadrianus octonaria †Hadrianus robustus †Hadrianus schucherti †Hadrianus utahensis †Hadrianus vialovi
†Hesperotestudo	†Hesperotestudo percrassa
Homopus	Common padloper Greater padloper
Indotestudo	Elongated tortoise Forsten's tortoise Travancore tortoise
Kinixys	Bell's hinge-back tortoise Forest hinge-back tortoise Home's hinge-back tortoise Lobatse hinge-back tortoise Natal hinge-back tortoise Speke's hinge-back tortoise
Malacochersus	Pancake tortoise
Manouria	Asian forest tortoise Impressed tortoise
†Megalochelys	†Megalochelys atlas †Megalochelys cautleyi †Megalochelys margae †Megalochelys sondaari
Psammobates	Geometric tortoise Serrated tortoise Tent tortoise
Pyxis	Flat-backed spider tortoise Spider tortoise
Stigmochelys	Leopard tortoise
†Stylemys	†Stylemys botti †Stylemys calaverensis †Stylemys canetotiana †Stylemys capax †Stylemys conspecta †Stylemys copei †Stylemys emiliae †Stylemys frizaciana †Stylemys karakolensis †Stylemys nebrascensis †Stylemys neglectus †Stylemys oregonensis †Stylemys pygmea †Stylemys uintensis †Stylemys undabuna
Testudo	Hermann's tortoise Kleinmann's tortoise Marginated tortoise Spur-thighed tortoise †Testudo brevitesta
Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. Key: †=extinct.

• IUCN Red List critically endangered species
• Chelonoidis
• Subspecies
• Turtles of South America
• Endemic reptiles of the Galápagos Islands
• Reptiles of Ecuador
• Taxa named by Georg Baur
• Reptiles described in 1889