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Frullania nisquallensis
Scientific classification
Kingdom:	Plantae
Division:	Marchantiophyta
Class:	Jungermanniopsida
Order:	Frullaniales
Family:	Frullaniaceae
Genus:	Frullania
Species:	F. nisquallensis
Binomial name
Frullania nisquallensis Sull.

Frullania nisquallensis, commonly known as hanging millipede liverwort, is a reddish-brown species of liverwort in the family Frullaniaceae. It is found in western Washington and British Columbia, including Vancouver Island. The plant grows in mats, sometimes in mats that hang from tree branches (particularly those of alders, or maples), or growing close to the substrate. The leaves are small (1 mm long) and flat, with the lower leaves being slightly smaller than those growing higher up the stem.

Secondary metabolites

This species contains the tridepside compound tenuiron, and the sesquiterpenes (-)-frullanolide and costunolide, the latter of which has been shown to be damaging to DNA.

References

1. Plants of the Pacific Northwest Coast. Lone Pine Publishing, 1994. p. 444
2. Kim YC, da S Bolzani V, Baj N, Gunatilaka AA, Kingston DG (February 1996). "A DNA-damaging sesquiterpene and other constituents from Frullania nisquallensis". Planta Med. 62 (1): 61–3. doi:10.1055/s-2006-957800. PMID 17252410.

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