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Geomitridae
Cernuella virgata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Superfamily:	Helicoidea
Family:	Geomitridae Boettger, 1909
Type genus
Geomitra Swainson, 1840
Genera
Actinella Backeljaia Candidula Caseolus Cernuella Cernuellopsis Cochlicella Discula Geomitra Helicella Helicopsis Microxeromagna Orexana Plentuisa Ponentina Trochoidea Xerocrassa Xerogyra Xerolenta Xeroplexa Xerosecta Xerotricha Zarateana

Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.

Anatomy

The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus. A free right ommatophoral retractor has been linked to adaptation to xeric habitats.

Taxonomy

The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015)

• Subfamily Geomitrinae
  • tribe Cochlicellini Schileyko, 1972
  • tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)
  • tribe Ponentinini Razkin et al., 2015
• Subfamily Helicellinae
  • tribe Cernuellini Schileyko, 1991
  • tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)
  • tribe Helicopsini H. Nordsieck, 1987
  • tribe Plentuisini Razkin et al., 2015
  • tribe Trochoideini H. Nordsieck, 1987

Genera

Genera with the family Geomitridae include:

Not belonging to a subfamily:

• Keraea Gude, 1911
• Peridotitea Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018
• Subfamily Geomitrinae
• tribe Cochlicellini
  • Cochlicella Férussac, 1821
  • Monilearia Mousson, 1872
  • Obelus W. Hartmann, 1842
  • Ripkeniella Hutterer & E. Gittenberger, 1998
• tribe Geomitrini
  • Actinella R. T. Lowe, 1852
  • Callina R. T. Lowe, 1855
  • Caseolus R. T. Lowe, 1852
  • Discula R. T. Lowe, 1852
  • Disculella Pilsbry, 1895
  • Domunculifex Brozzo, De Mattia, Harl & Neiber, 2020
  • Geomitra Swainson, 1840
  • Helicomela R. T. Lowe, 1855
  • Heterostoma W. Hartmann, 1843: synonym of Steenbergia Mandahl-Barth, 1950 (invalid: junior homonym of Heterostoma de Filippi, 1837 )
  • Hystricella R. T. Lowe, 1855
  • Lemniscia R. T. Lowe, 1855
  • Moreletina de Frias Martins, 2002
  • Plebecula R. T. Lowe, 1852
  • Pseudocampylaea L. Pfeiffer, 1877
  • Serratorotula Groh & Hemmen, 1986
  • Spirorbula R. T. Lowe, 1852
  • Steenbergia Mandahl-Barth, 1950
  • Testudodiscula Brozzo, De Mattia, Harl & Neiber, 2020
  • Wollastonaria De Mattia, Neiber & Groh, 2018
• tribe Ponentinini
  • Ponentina P. Hesse, 1921

subfamily Helicellinae

• tribe Cernuellini
  • Alteniella Clerx & Gittenberger, 1977
  • Cernuella Schlüter, 1838
  • Microxeromagna Ortiz de Zárate López, 1950
  • Xeroplana Monterosato, 1892
  • Xerosecta Monterosato, 1892
• tribe Helicellini
  • Backeljaia Chueca et al., 2018
  • Candidula Kobelt, 1871
  • Helicella Férussac, 1821
  • Orexana Chueca et al., 2018
  • Xerogyra Monterosato, 1892
  • Xeroleuca Kobelt, 1877
  • Xeroplexa Monterosato, 1892
  • Xerotricha Monterosato, 1892
  • Zarateana Chueca et al., 2018
• tribe Helicopsini H. Nordsieck, 1987
  • Helicopsis Fitzinger, 1833
  • Pseudoxerophila Westerlund, 1879
  • Xerolenta Monterosato, 1892
  • Xeromunda Monterosato, 1892
  • Xeropicta Monterosato, 1892
• tribe Plentuisini
  • Plentuisa Puente & Prieto, 1992
• tribe Trochoideini
  • Trochoidea T. Brown, 1827
  • Xerocrassa Monterosato, 1892

References

1. ^ Razkin, Oihana; Gómez-Moliner, Benjamín Juán; Prieto, Carlos Enrique; Martínez-Ortí, Alberto; Arrébola, José Ramón; Muñoz, Benito; Chueca, Luis Javier; Madeira, María José (February 2015). "Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution. 83: 99–117. doi:10.1016/j.ympev.2014.11.014. PMID 25485783.
2. "WoRMS - World Register of Marine Species - Geomitridae C. R. Boettger, 1909". www.marinespecies.org. Retrieved 5 September 2024.
3. Giusti, F; Manganelli, G (1987). "Notulae malacologicae, XXXVI. On some Hygromiidae (Gastropoda: Helicoidea) living in Sardinia and in Corsica.(Studies on the Sardinian and Corsican malacofauna VI)". Bollettino Malacologico. 23: 123–206.
4. Chueca, Luis J.; Gómez-Moliner, Benjamín J.; Madeira, María José; Pfenninger, Markus (January 2018). "Molecular phylogeny of Candidula (Geomitridae) land snails inferred from mitochondrial and nuclear markers reveals the polyphyly of the genus". Molecular Phylogenetics and Evolution. 118: 357–368. doi:10.1016/j.ympev.2017.10.022. ISSN 1055-7903. PMID 29107619.
5. ^ "Geomitrini". Fauna Europaea, last update 27 January 2011, accessed 4 August 2011.

• Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

External links

• https://liferecovernatura.madeira.gov.pt
• Razkin, O., Gómez-Moliner, B. J., Prieto, C. E., Martínez-Ortí, A., Arrébola, J. R., Muñoz, B., Chueca, L. J. & Madeira, M. J. (2015). Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora). Molecular Phylogenetics and Evolution. 83: 99-117

• Geomitridae
• Helicoidea
• Gastropod families