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Argiles et Grès à Reptiles Formation
Stratigraphic range: Campanian-Early Maastrichtian, 74–70 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Type	Geological formation
Underlies	Calcaire de Rognac
Thickness	Over 40 m (130 ft)
Lithology
Primary	Clay, shale
Other	Sandstone
Location
Coordinates	43°30′N 5°30′E / 43.5°N 5.5°E / 43.5; 5.5
Approximate paleocoordinates	36°06′N 4°00′E / 36.1°N 4.0°E / 36.1; 4.0
Region	Provence-Alpes-Cote d'Azur, Bouches-du-Rhône
Country	France
Extent	Aix-en-Provence Basin
Argiles et Grès à Reptiles Formation (France)

Argiles et Grès à Reptiles Formation (meaning Reptile Clay and Sandstone) also known as the Argiles Rutilantes Formation is an early Maastrichtian French geologic formation in the département of Var preserving the remains of several types of dinosaurs and other extinct organisms.

Fossil content

An abelisaurid similar to Arcovenator is known from Département des Bouches-du-Rhone, while enantiornithean and indeterminate avialan remains are known from Département du Var.

Dinosaurs

Dinosaurs of the Grès à Reptiles
Genus	Species	Location	Stratigraphic position	Abundance	Notes	Images
Abelisauridae	Indeterminate	Bouches du Rhône	Lower		A large abelisaurid that was similar to Arcovenator.
Arcovenator	A. escotae	Var	Lower		A large abelisaurid.
Atsinganosaurus	G. meridionalis	Bouches du Rhône	Lower		A titanosaur Lirainosaurinae.
Castignovolucris	C. sebei	Castigno valley, Villespassans, Hérault	Lower (i.e. continental red clays)		An enantiornithine bird.
Garrigatitan	A. velauciensis	Bouches du Rhône	Lower		A titanosaur Lirainosaurinae.
Gargantuavis	G. philoinos	Var
Hypselosaurus	H. priscus	Bouches-du-Rhône Var			Actually indeterminate sauropod remains.
Martinavis	M. cruzyensis				An enantiornithe bird
Matheronodon	M. provincialis	Bouches-du-Rhône			A rhabdodontid ornithopod
Theropoda	Indeterminate	Var			Previously referred to "Megalosaurus" pannoniensis
Pyroraptor	P. olympius	Bouches-du-Rhône		"Fragmentary postcrania and teeth"
Rhabdodon	R. priscus	Bouches-du-Rhône Var
Rhodanosaurus	R. lugdunensis	Bouches-du-Rhône
Variraptor	V. mechinorum	Var Bouches-du-Rhône

Crocodylians

• Acynodon iberoccitanus
• Allodaposuchus sp.
• Alligatoridae indet.
• Crocodylidae indet.

Turtles

• Foxemys mechinorum
• cf. Polysternon sp.
• cf. Solemys sp.

Pterosaurs

• Mistralazhdarcho maggii

Fish

• Hybodontiformes indet.
• Lepisosteidae indet.

Fossil eggs

• Megaloolithus aureliensis
• Megaloolithus petralta
• Megaloolithus sp.

Flora

• Peckichara cancellata
• P. pectinata
• Platychara caudata
• Saportanella maslovi
• ?Amblyochara begudiana

See also

• List of dinosaur-bearing stratigraphic units
• List of fossiliferous stratigraphic units in France

References

1. Tabuce et al., 2013
2. ^ Weishampel et al., 2004, pp.588-593
3. Argiles et Grès à Reptiles Formation at Fossilworks.org
4. Tortosa, Thierry; Buffetaut, Eric; Vialle, Nicolas; Dutour, Yves; Turini, Eric; Cheylan, Gilles (2014-01-01). "A new abelisaurid dinosaur from the Late Cretaceous of southern France: Palaeobiogeographical implications". Annales de Paléontologie. 100 (1): 63–86. Bibcode:2014AnPal.100...63T. doi:10.1016/j.annpal.2013.10.003. ISSN 0753-3969.
5. Díez Díaz, Verónica; Garcia, Géraldine; Pereda Suberbiola, Xabier; Jentgen-Ceschino, Benjamin; Stein, Koen; Godefroit, Pascal; Valentin, Xavier (2020-12-21). "A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France)". Historical Biology. 33 (11): 2998–3017. doi:10.1080/08912963.2020.1841184. ISSN 0891-2963. S2CID 234404741.
6. ^ Buffetaut, E.; Angst, D.; Tong, H. (2023). "A new enantiornithine bird from Upper Cretaceous non-marine deposits at Villespassans (Hérault, southern France)". Annales de Paléontologie. 109 (1). 102585. Bibcode:2023AnPal.10902585B. doi:10.1016/j.annpal.2022.102585. S2CID 257863645.
7. B. Vila; A. Galobart; J.U. Canudo; J. Le Loeff (2012). "The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe: Clues to demise and extinction". Palaeogeography, Palaeoclimatology, Palaeoecology. 350–352 (15): 19–38. Bibcode:2012PPP...350...19V. doi:10.1016/j.palaeo.2012.06.008.
8. ^ "14.8 Département du Var, France; 2. Grès à Reptiles Formation," in Weishampel et al., 2004, p.589
9. ^ "14.8 Département du Var, France; 2. Grès à Reptiles Formation" and "14.38 Département des Bouches-du-Rhone, France; 1. Grès à Reptiles Formation," in Weishampel et al., 2004, pp.589-590
10. ^ "14.38 Département des Bouches-du-Rhone, France; 1. Grès à Reptiles Formation," in Weishampel et al., 2004, p.590
11. "Table 10.1," in Weishampel et al., 2004, p.199
12. Martin, 2007
13. Blanco & Brochu, 2016
14. ^ Le Loeuff & Buffetaut, 1998
15. Tong et al., 1998
16. ^ Martin et al., 2016
17. Vullo et al., 2018
18. ^ Viany-Liaud et al., 1994

Bibliography

• Vullo, R.; Garcia, G.; Godefroit, P.; Cincotta, A.; Valentin, X. (2018), "Mistralazhdarcho maggii, gen. et sp. nov., A New Azhdarchid Pterosaur from the Upper Cretaceous of Southeastern France", Journal of Vertebrate Paleontology, 38 (4): 1–16, doi:10.1080/02724634.2018.1502670
• Blanco, A.; Brochu, C. A. (2016), "Intra- and interspecific variability in allodaposuchid crocodylomorphs and the status of western European taxa", Historical Biology, 29: 395–408
• Martin, J. E.; Delfino, M.; Garcia, G.; Godefroit, P.; Berton, S.; Valentin, X. (2016), "New specimens of Allodaposuchus precedens from France: intraspecific variability and the diversity of European Late Cretaceous eusuchians", Zoological Journal of the Linnean Society, 176 (3): 607–631, doi:10.1111/zoj.12331
• Tabuce, Rodolphe; Tortosa, Thierry; Vianey-Liaud, Monique; Garcia, Géraldine; Lebrun, Renaud; Godefroit, Pascal; Dutour, Yves; Berton, Sévérine; Valentin, Xavier; Cheylan, Gilles (2013), "New eutherian mammals from the Late Cretaceous of Aix-en-Provence Basin, south-eastern France", Zoological Journal of the Linnean Society, 169 (3): 653–672, doi:10.1111/zoj.12074, ISSN 0024-4082
• Martin, J. E (2007), "New material of the Late Cretaceous globidontan Acynodon iberoccitanus (Crocodylia) from southern France", Journal of Vertebrate Paleontology, 27 (2): 362–372, doi:10.1671/0272-4634(2007)27[362:NMOTLC]2.0.CO;2
• Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (2004), The Dinosauria, 2nd edition, Berkeley: University of California Press, pp. 1–880, ISBN 0-520-24209-2, retrieved 2019-02-21
• Le Loeuff, J.; Buffetaut, É. (1998), "A new dromaeosaurid theropod from the Upper Cretaceous of southern France.", Oryctos, 1: 105–112
• Tong, H.; Gaffney, E. S.; Buffetaut, É. (1998), "Foxemys, a new side-necked turtle (Bothremydidae: Pelomedusoides) from the Late Cretaceous of France", American Museum Novitates (3251): 1–19
• Vianey-Liaud, M.; Mallan, P.; Buscail, O.; Montgelard, C. (1994), Review of French dinosaur eggshells: morphology, structure, mineral, and organic composition. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, Cambridge University Press, Cambridge, pp. 151–183

Further reading

• R. Cousin, G. Breton, R. Fournier and J.-P. Watte. 1994. Dinosaur egglaying and nesting in France. K. Carpenter, K.F. Hirsch & J.R. Horner (eds.) Dinosaur Eggs and Babies. Cambridge University Press, Cambridge 56–74
• J. Le Loeuff, E. Buffetaut, P. Méchin and A. Méchin-Salessy. 1992. The first record of dromaeosaurid dinosaurs (Saurischia, Theropoda) in the Maastrichtian of southern Europe: palaeobiogeographical implications. Bulletin de la Société géologique de la France 163(3):337-343
• R. Dughi and F. Sirugue. 1958. Sur les oeufs de dinosaures du bassin fluvio-lacustre de Basse-Provence. Comptes Rendus du Congrès National des Sociétés Savantes, Section des Sciences, Paris 83:183-20

• Geologic formations of France
• Upper Cretaceous Series of Europe
• Cretaceous France
• Maastrichtian Stage
• Sandstone formations
• Shale formations
• Fluvial deposits
• Ooliferous formations
• Fossiliferous stratigraphic units of Europe
• Paleontology in France