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Hadar

Site location in Ethiopia
Location	Mille district, Afar Region,  Ethiopia
Region	Eastern Africa
Coordinates	11°10′01″N 40°37′59″E / 11.167°N 40.633°E / 11.167; 40.633
Type	Archaeological Paleontology
History
Periods	Middle - Late Pliocene
Site notes
Excavation dates	1970, 1972, 1973, 1974, 2000
Archaeologists	Donald Johanson Maurice Taieb Denis Geraads Zeresenay Alemseged
Condition	Excavated
Ownership	Ethiopian Government

Hadar or Hadar Formation (also spelled Qad daqar, Qadaqar; Afar "white stream ") is a paleontological fossil site located in Mille district, Administrative Zone 1 of the Afar Region, Ethiopia, 15 km upstream (west) of the A1 road's bridge across the Awash River (Adayitu kebele).

It is situated on the southern edge of the Afar Triangle (part of East Africa's Great Rift Valley), along the left banks of the Awash River, between two minor tributaries, the eponymous Kada Hadar and the Kada Gona. In 1972, Taieb organized a small exploratory reconnaissance of the Afar region to investigate more paleontological finds there. After six weeks of exploration, the party focused on the Hadar site.

The site has yielded some of the most well-known hominin fossils, including "Lucy". These hominin fossils range in age from approximately 3.42 to 2.90 million years ago. These finds give us a greater understanding of hominin evolution during this period.

It is postulated that the specimens in the region were deposited by way of a large river system with associated crevasse channels/splays, deltas, and distributary channels, as well as periodic transgressions of paleolake Hadar located east of the research area (Aronson and Taieb, 1981, Tiercelin, 1986, Campisano and Feibel, in press) possibly related to geological activity or climatic cycles in at least the Kada Hadar Member (Yemane et al., 1996, Yemane, 1997, Campisano and Feibel, in press)."

According to Jon Kalb, early maps show caravan routes passing within 10 to 15 km of Hadar but not through it. The British explorer L.M. Nesbitt passed 15 km west of Hadar in 1928.

Geology

The region's rocks consist mainly of mudstones, siltstones, fine-grained sandstones and volcanic tuffs. The region of Hadar has been divided into four geologic members — Basal (~3.8–3.42 Ma), Sidi Hakoma (~3.42–3.26 Ma), Denen Dora (~3.26–3.2 Ma), and Kada Hadar (<~3.2 Ma)—with three tuffs (Sidi Hakoma Tuff , Triple Tuff and Kada Hadar Tuff ) separating the four members.

The Sidi Hakoma member tends towards high rainfall and low seasonality. The overlying Denan Dora Member was a grassland habitat. Finally, the Kada Hadar Member was an even more open and arid habitat, as seen in the high abundance of antilopines, which frequent these types of terrains.

Paleontology

The first paleo-geological explorations of the Hadar area were conducted by Maurice Taieb. He found Hadar in December 1970 by following the Ledi River, which originates in the highlands north of Bati to empty into the Awash River. Taieb recovered several fossils in the area and led a party back to Hadar in May 1972. In October 1973, 16 individuals with the International Afar Research Expedition (IARE) arrived at Hadar and camped there for two months during which the first hominin fossil was found. (Taieb claims in his 1985 book Sur la Terre des premiers Hommes to have discovered the Hadar fields in 1968, but Kalb argues that claim to be incorrect.) The IARE party examined a series of sedimentary layers called the Hadar Formation, which was dated to the late Pliocene to early Pleistocene epochs (3.5 to 2.3 million years ago).

Discovery of Lucy

The anthropologist Donald Johanson, a member of the 1973 expedition to Hadar, returned the next year and discovered the fossil hominin "Lucy" in the late fall of 1974. He spotted a right proximal ulna in a gully, followed by an occipital bone, a femur, some ribs, a pelvis, and a lower jaw. Within two weeks, nearly 40% of the hominoid skeleton had been identified and cataloged. Lucy is the most famous fossil to have been found at Hadar. Lucy is among the oldest hominin fossils ever discovered and was later given the taxonomic classification Australopithecus afarensis. (The name 'Lucy' was inspired by the song "Lucy in the Sky with Diamonds" by The Beatles, which happened to be playing on the radio at base camp.)

Specimens and inferences

In 1975, Donald Johanson made another discovery at a nearby site in Hadar: 216 specimens from approximately 17 individuals, most likely related and varying in age, called AL 333 (colloquially referred to as the "First Family").

About thirty years later in nearby Dikika, another Australopithecus afarensis fossil skeleton was found in a separate outcrop of the Hadar Formation across the Awash River from Hadar. The skeleton is of a three-year-old girl later named "Selam," which means peace in Amharic Ethiopian languages.

In 1973 and 1974 when the first anatomical discoveries were made, their size and shape pointed towards a variety of taxa, but further research has confirmed that only one hominin taxon is present here. The first find there was a fossil knee joint estimated to date from 3.4 million years ago. Since then, the Hadar research area has yielded 370 specimens of A. afarensis, one specimen of Homo, and 7571 additional vertebrate specimens.

The specimens recovered display a variety of different primitive cranial post features, which indicate A. afarensis is distinct from other species of Australopithecus: small cranial capacity, palate similar to African apes (parallel tooth rows, shallow, long from front to back, narrow from side to side), primitive occipital, basal cranium anatomy, high frequency of unicuspid third premolars, prognathic face, and primitive mandibular anatomy. Postcranially, the pelvis, knee, ankle, and foot indicate habitual, terrestrial bipedalism, but ape-like curved finger and foot bones are retained ancestral ape-like features.

Paleofauna

Although Australopithecus is well-known for its discovery since the 1970s, other specimens from the Hadar Formation contain several fossil remains of artiodactyls, perissodactyls, carnivorans, proboscideans, and other African species that are well preserved. The bovids found in the formation included the Aepycerotini (Aepyceros), Alcelaphini (Damalborea and Parmularius), Antilopini (Gazella), Bovini (Ugandax and Pelorovis), Caprini (Budorcas), Cephalophini, Hippotragini (Oryx), Neotragini (Raphicerus and Madoqua), Reduncini (Kobus), and Tragelaphini (Tragelaphus). Artiodactyls outside the bovid family were present within the formation as well, namely the giraffids (Giraffa and Sivatherium), Hippopotamidae (Hippopotamus), and suids (Kolpochoerus, Notochoerus, and Nyanzachoerus). While a definitive list of carnivorans found within the Hadar Formation has yet to be compiled, confirmed genera that were found within the Hadar Formation include canids (Canis and Nyctereutes), felids (Dinofelis, Leptailurus, Felis, Homotherium, and Panthera), hyaenids (Chasmaporthetes, Ikelohyaena, Crocuta, Hyaena, and cf. Pliocrocuta), herpestids (Herpestes and cf. Helogale), mustelids (Mellivora, Enhydriodon, and cf. Poecilogale), and the viverrid (cf. Civettictis). Mammals within the formation outside the artiodactyl and carnivoran families include a bat (indeterminate), the leporid (Lepus), the equid (Eurygnathohippus), rhinoceroses (Ceratotherium and Diceros), old world primates (Parapapio, Theropithecus, and Cercopithecoides), proboscideans (the deinothere Deinotherium and elephants Elephas, Loxodonta, and Mammuthus) old world porcupines (Hystrix and Xenohystrix), murid rodents (Gerbilliscus, Acomys, Golunda, Oenomys, Praomys, Saidomys, Millardia, and Mus), the spalacid Tachyoryctes, a squirrel indet., and an aardvark species. Taxons within other classes are present within the Hadar Formation as well, such as birds (Plectropterus, Balearica, Anhinga, and Struthio) and reptiles (Crocodylus, Python, Varanus, and Bitis).

Mammals

Artiodactyls

Bovidae

Bovids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Aepyceros	A. datoadeni	Kada Hadar Member at Hadar	Near-complete skull lacking parts of horn cores (AL 787-1 KH1)	An impala
Beatragus	B. sp.		Detached braincase with two horn cores	A hirola
Budorcas	B. churcheri	Sida Hakoma Member.	Complete skull with horns (AL 136-5 DD2)	An extinct takin
Damalops	D. sidihakomai	Sidi Hakoma Member	AL 208–7, a skull with horn cores, right P4–M3, and left M2–3.	An extinct genus of Alcelaphinae
Damalborea	D. elisabethae	Basal and Sidi Hakoma members	A skull with horn cores (AL 208-07 SH3)	An extinct genus of Alcelaphinae
	D. grayi	Denen Dora and Kada Hadar members	Multiple postcranial fragments, such as horn cores, maxillae, and mandibles
Gazella	G. harmonae	Kada Hadar Member at Hadar	Skull remains, such as isolated teeth, maxillae, frontlets, and occipital bones (AL 444-16)	Extinct relatives of modern gazelles
	G. cf. janenschi	Denen Dora Member	Horn cores (AL 302-6 DD)
	G. cf. praethomsoni	Denen Dora and Kada Hadar members	Few Horn core specimens
Hippotragini	Praedamalis deturi	Kada Hadar, Sida Hakoma, and Denen Dora members at Hadar	Multiple specimens based on horn cores and lower teeth	Ancestor of modern oryx
Kobus	K. hadarensis	Sidi Hakoma, Kada Hadar and Denen Dora member		Extinct relatives of antelopes
	K. oricornus	Denen Dora and Kada Hadar members	Several specimens of braincase and horn core
Madoqua	M. sp.			Relative of dik-dik
Raphicerus	R. sp.	Kada Hadar and Sidi Hakoma members	Mandibles and horn cores	Relative of Neotragini
Parmularius	P. pachyceras	Sidi Hakoma and Kada Hadar members	Braincases and horn cores	Large extinct alcelphines related to topis and hartebeests
Pelorovis	P. sp.
Tragelaphus	T. lockwoodi	Basal and Sidi Hakoma members	Braincase with complete horn cores (AL 142-3 SH2)	Extinct relatives of spiral-horned antelopes
	T. rastafari	Basal, Sidi Hakoma, Denen Dora, and Kada Hadar members	Several horn core specimens
Ugandax	U. coryndonae	Denen Dora and Sidi Hakoma members at Hadar	Complete partial skull and horns (AL 194-1 DD1)	Ancestor of Cape buffalo

Giraffidae

Giraffids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Giraffa	G. jumae	Sidi Hakoma and Denen Dora members	Partial teeth, vertebrae, ossicones, limb bones, and mandibles	Extinct relatives of modern giraffe
	G. stillei	Sidi Hakoma and Denen Dora members	Partial teeth, ossicones, limb bones, and mandibles
Sivatherium	S. maurusium	Denen Dora, Kada Hadar, and Sidi Hakoma members	Partial mandibles and horns	Okapi-like giraffid

Hippopotamidae

Suidae

Suids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Kolpochoerus	K. afarensis	Kada Hadar, Sidi Hakoma, and Denen Dora members	Postcranial skulls, teeth fragments, and mandibles	Extinct relative of the pig family
Notochoerus	N. eulius	Kada Hadar, Lower and Upper Sidi Hakoma, and Denen Dora members	Postcranial skull, teeth, and mandibles	A Tetraconodontinae pig.
Nyanzachoerus	N. kanamensis	Basal and Sidi Hakoma members	Complete cranium, mandibles, and dentition	A Tetraconodontinae pig.
	N. pattersoni	Basal, Lower and Upper Sidi Hakoma, and Denen Dora members	Complete skulls, mandibles, and dentition

Carnivorans

Canidae

Felidae

Felids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Dinofelis	D. aronoki	Kada Hadar Member	Postcranial member: distal radius (AL-363-20)	A machairodontine felid
	D. petteri	Sidi Hakoma and Denen Dora members	Postcranial member: partial humerus (AL 168-30), proximal ulna (AL 262-9), and right astragalus (Al 173-11)
Felis	F. sp.	Kada Hadar, Sidi Hakoma, and Denen Dora members		A small cat relative
Homotherium	H. hadarensis	Denen Dora Member	Complete cranium skull, mandible fragments (DIK-96-1), teeth, and forelimbs	A machairodontine felid
Megantereon	M. sp.	Kada Hadar, Sidi Hakoma, and Denen Dora members		A machairodontine felid
Leptailurus	L. sp.			A serval cat
Panthera	P. sp.	Sidi Hakoma Member		Relatives of lions

Herpestidae

Hyaenidae

Mustelidae

Viverridae

Perissodactyls

Equidae

Rhinocerotidae

Primates

Cercopithecidae

Hominidae

Hominids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Australopithecus	A. afarensis	Basal, Denen Dora, Kada Hadar, and Sidi Hakoma members	Partial skeletons featuring small cranial capacity, primitive occipitals, basal cranium anatomy, high frequency of unicuspid third premolars, prognathic faces, and primitive mandibular anatomy	An australopithecine hominid

Proboscideans

Deinotheriidae

Deinotheriids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Deinotherium	D. bozasi	Denen Dora, Kada Hadar, and Sidi Hakoma members	Complete cranium skull and teeth fragments	A large extinct elephant-like proboscidean

Elephantidae

Elephantids of the Hadar Formation
Taxa	Species	Presence	Material	Notes	Images
Elephas	E. ekorensis	Sidi Hakoma and Denen Dora members at Hadar and Dikika	Mandibles and molar fragments	Extinct lineage related to Asian elephant
Loxodonta	L. adaurora	Denen Dora and Sidi Hakoma members at Hadar	Mandibles and molar fragments	Extinct subspecies of modern African elephant
	L. exoptata	Denen Dora Member	Undescribed molars
Mammuthus	M. sp.	Sidi Hakoma and Denen Dora Member.	Undescribed molars	A mammoth relative
Palaeoloxodon recki	P. r. brumpti	Sidi Hakoma Member	Mandibles and molar fragments	Extinct relative of African elephant

Rodents

Birds

Reptiles

See also

• Ledi-Geraru

References

1. ^ Jon Kalb Adventures in the Bone Trade (New York: Copernicus Books, 2001), p. 83
2. ^ DiMaggio, Erin N.; Arrowsmith, J Ramón; Campisano, Christopher J.; Johnson, Roy; Deino, Alan L.; Warren, Mark; Fisseha, Shimeles; Cohen, Andrew S. (December 2015). "Tephrostratigraphy and depositional environment of young (<2.94 Ma) Hadar Formation deposits at Ledi-Geraru, Afar, Ethiopia". Journal of African Earth Sciences. 112: 234–250. Bibcode:2015JAfES.112..234D. doi:10.1016/j.jafrearsci.2015.09.018.
3. ^ Johanson, Donald (March 2017). "The paleoanthropology of Hadar, Ethiopia". Comptes Rendus Palevol. 16 (2): 140–154. Bibcode:2017CRPal..16..140J. doi:10.1016/j.crpv.2016.10.005.
4. Halstead, Lambert Beverly (1984). A la recherche du passé: la vie sur terre des origines aux premiers hommes. Hachette. ISBN 978-2-01-009602-0. OCLC 25125386.
5. Feibel, Craig S. (November 2004). Sedimentary Patterns in the Pliocene Hadar Formation, Afar Rift, Ethiopia. 2004 Denver Annual Meeting of The Geological Society of America.
6. ^ Hogenboom, Melissa. "The 'Lucy' fossil rewrote the story of humanity". Retrieved 25 October 2018.
7. "Lucy's Story | Institute of Human Origins". iho.asu.edu. Retrieved 25 October 2018.
8. Reed, Kaye E.; Bibi, Faysal (March 2011). "Fossil Tragelaphini (Artiodactyla: Bovidae) from the Late Pliocene Hadar Formation, Afar Regional State, Ethiopia". Journal of Mammalian Evolution. 18 (1): 57–69. doi:10.1007/s10914-010-9146-6.
9. ^ Geraads, Denis; Bobe, René; Reed, Kaye (2012). "Pliocene Bovidae (mammalia) from the Hadar Formation of Hadar and Ledi-Geraru, Lower Awash, Ethiopia". Journal of Vertebrate Paleontology. 32 (1): 180–197. Bibcode:2012JVPal..32..180G. doi:10.1080/02724634.2012.632046. JSTOR 41407716.
10. Reed, Kaye E. (June 2008). "Paleoecological patterns at the Hadar hominin site, Afar Regional State, Ethiopia". Journal of Human Evolution. 54 (6): 743–768. doi:10.1016/J.JHEVOL.2007.08.013. PMID 18191177.
11. ^ Cooke, H B S (1978). "Pliocene-Pleistocene Suidae from Hadar, Ethiopia". Kirtlandia. 29: 1–63.
12. ^ Werdelin, Lars; Lewis, Margaret E. (June 2001). "A revision of the genus Dinofelis (Mammalia, Felidae)". Zoological Journal of the Linnean Society. 132 (2): 147–258. doi:10.1111/j.1096-3642.2001.tb02465.x.
13. ^ Werdelin, Lars; Lewis, Margaret E. (2008). "New Species of Crocuta from the Early Pliocene of Kenya, with an Overview of Early Pliocene Hyenas of Eastern Africa". Journal of Vertebrate Paleontology. 28 (4): 1162–1170. Bibcode:2008JVPal..28.1162W. doi:10.1671/0272-4634-28.4.1162. JSTOR 20491047.
14. ^ Geraads, Denis; Alemseged, Zeresenay; Bobe, René; Reed, Denné (17 March 2011). "Enhydriodon dikikae , sp. nov. (Carnivora: Mammalia), a gigantic otter from the Pliocene of Dikika, Lower Awash, Ethiopia". Journal of Vertebrate Paleontology. 31 (2): 447–453. Bibcode:2011JVPal..31..447G. doi:10.1080/02724634.2011.550356.
15. ^ Gilbert, W. Henry; Bernor, Raymond L. (2009). "Equidae". Homo erectus: Pleistocene Evidence from the Middle Awash, Ethiopia. pp. 133–166. doi:10.1525/california/9780520251205.003.0006. ISBN 978-0-520-25120-5.
16. ^ Geraads, Denis (2005). "Pliocene Rhinocerotidae (Mammalia) from Hadar and Dikika (Lower Awash, Ethiopia), and a Revision of the Origin of Modern African Rhinos" (PDF). Journal of Vertebrate Paleontology. 25 (2): 451–461. doi:10.1671/0272-4634(2005)025[0451:PRMFHA]2.0.CO;2. JSTOR 4524458.
17. ^ Kalb, Jon E.; Mebrate, Assefa; Tischler, Doris (1993). "Fossil Elephantoids: From the Hominid-Bearing Awash Group, Middle Awash Valley, Afar Depression, Ethiopia". Transactions of the American Philosophical Society. 83 (1): i–114. doi:10.2307/1006558. JSTOR 1006558.
18. ^ Sanders, William J.; Haile-Selassie, Yohannes (June 2012). "A New Assemblage of Mid-Pliocene Proboscideans from the Woranso-Mille Area, Afar Region, Ethiopia: Taxonomic, Evolutionary, and Paleoecological Considerations". Journal of Mammalian Evolution. 19 (2): 105–128. doi:10.1007/s10914-011-9181-y.
19. Rowan, John; Lazagabaster, Ignacio A.; Campisano, Christopher J.; Bibi, Faysal; Bobe, René; Boisserie, Jean-Renaud; Frost, Stephen R.; Getachew, Tomas; Gilbert, Christopher C.; Lewis, Margaret E.; Melaku, Sahleselasie; Scott, Eric; Souron, Antoine; Werdelin, Lars; Kimbel, William H.; Reed, Kaye E. (6 April 2022). "Early Pleistocene large mammals from Maka'amitalu, Hadar, lower Awash Valley, Ethiopia". PeerJ. 10: e13210. doi:10.7717/peerj.13210. PMC 8994497. PMID 35411256.
20. ^ Azzarà, Beatrice; Cherin, Marco; Adams, Justin; Boschian, Giovanni; Crotti, Marco; Denys, Christiane; Fressoia, Lorenzo; Kimambo, Jackson S.; Kwekason, Amandus; Iurino, Dawid A.; Manzi, Giorgio; Masao, Fidelis T.; Melaku, Sahleselasie; Menconero, Sofia; Mori, Emiliano; Zipfel, Bernhard (June 2022). "The Thorny Issue of African Porcupines: a New Mandible of Hystrix makapanensis from Olduvai Gorge (Tanzania) and Rediagnosis of the Species". Journal of Mammalian Evolution. 29 (2): 447–474. doi:10.1007/s10914-021-09588-z. PMC 8776392. PMID 35079214.
21. ^ Reed, Denné N. (2011). "New Murid (mammalia, Rodentia) Fossils from a Late Pliocene (2.4 Ma) Locality, Hadar A.l. 894, Afar Region, Ethiopia". Journal of Vertebrate Paleontology. 31 (6): 1326–1337. Bibcode:2011JVPal..31.1326R. doi:10.1080/02724634.2011.620676. JSTOR 41407688.
22. Campisano, Christopher J.; Rowan, John; Reed, Kaye E. (2022). "The Hadar Formation, Afar Regional State, Ethiopia: Geology, Fauna, and Paleoenvironmental Reconstructions". African Paleoecology and Human Evolution. pp. 214–228. doi:10.1017/9781139696470.018. ISBN 978-1-139-69647-0.
23. ^ Geraads, Denis; Alemseged, Zeresenay; Bobe, René; Reed, Denné (July 2015). "Pliocene Carnivora (Mammalia) from the Hadar Formation at Dikika, Lower Awash Valley, Ethiopia". Journal of African Earth Sciences. 107: 28–35. Bibcode:2015JAfES.107...28G. doi:10.1016/j.jafrearsci.2015.03.020.
24. ^ Geraads, Denis; Reed, Kaye; Bobe, Rene (March 2013). "Pliocene Giraffidae (Mammalia) from the Hadar Formation of Hadar and Ledi-Geraru, Lower Awash, Ethiopia". Journal of Vertebrate Paleontology. 33 (2): 470–481. Bibcode:2013JVPal..33..470G. doi:10.1080/02724634.2013.723657.
25. ^ Frost, Stephen R.; Delson, Eric (November 2002). "Fossil Cercopithecidae from the Hadar Formation and surrounding areas of the Afar Depression, Ethiopia". Journal of Human Evolution. 43 (5): 687–748. doi:10.1006/jhev.2002.0603. PMID 12457855.
26. White, Tim D.; Moore, Roderic V.; Suwa, Gen (1984). "Hadar Biostratigraphy and Hominid Evolution". Journal of Vertebrate Paleontology. 4 (4): 575–583. Bibcode:1984JVPal...4..575W. doi:10.1080/02724634.1984.10012033. JSTOR 4523018.
27. Kalb, Jon E.; Jolly, Clifford J.; Tebedge, Sleshi; Mebrate, Assefa; Smart, Charles; Oswald, Elizabeth B.; Whitehead, Paul F.; Woods, Craig B.; Adefris, Tsrha; Rawn-Schatzinger, Viola (1982). "Vertebrate Faunas from the Awash Group, Middle Awash Valley, Afar, Ethiopia". Journal of Vertebrate Paleontology. 2 (2): 237–258. Bibcode:1982JVPal...2..237K. doi:10.1080/02724634.1982.10011932. JSTOR 4522897.
28. ^ Reed, Kaye E. (June 2008). "Paleoecological patterns at the Hadar hominin site, Afar Regional State, Ethiopia". Journal of Human Evolution. 54 (6): 743–768. doi:10.1016/J.JHEVOL.2007.08.013. PMID 18191177.
29. Howell, F. C.; Petter, G. (1979). "Diversification et affinites des Carnivores pliocenes du groupe de l'Omo et de la formation d'Hadar (Ethiopie)". Bulletin de la Société Géologique de France. S7-XXI (3): 289–293. doi:10.2113/gssgfbull.s7-xxi.3.289.
30. Gentry, A.W. (1996). "A fossil Budorcas (Mammalia, Bovidae) from Africa". Palaeoecology and Palaeoenvironments of Late Cenozoic Mammals. pp. 571–587. doi:10.3138/9781487574154-028. ISBN 978-1-4875-7415-4.
31. Vrba, E. S. (1997). "New fossils of Alcelaphini and Caprinae (Bovidae: Mammalia) from Awash, Ethiopia, and phylogenetic analysis of Alcelaphini". Palaeontologia Africana. 34. hdl:10539/16416.
32. Geraads, Denis; Alemseged, Zeresenay; Bobe, René; Reed, Denné (2010). "Nyctereutes Lockwoodi , N. Sp., a New Canid (carnivora: Mammalia) from the Middle Pliocene of Dikika, Lower Awash, Ethiopia". Journal of Vertebrate Paleontology. 30 (3): 981–987. Bibcode:2010JVPal..30..981G. doi:10.1080/02724631003758326. JSTOR 40666211.
33. Getahun, Dagmawit Abebe; Delson, Eric; Seyoum, Chalachew Mesfin (July 2023). "A review of Theropithecus oswaldi with the proposal of a new subspecies" (PDF). Journal of Human Evolution. 180: 103373. doi:10.1016/j.jhevol.2023.103373. PMID 37269782.
34. Brodkorb, Pierce; Mourer-Chauviré, Cécile (January 1982). "Fossil Anhingas (Aves: Anhingidae) from Early Man Sites of Hadar and Omo (Ethiopia) and Olduvai Gorge (Tanzania)". Geobios. 15 (4): 505–515. Bibcode:1982Geobi..15..505B. doi:10.1016/S0016-6995(82)80071-5.

External links

• Photo gallery from a University of Washington archaeological field season

• Archaeological sites in Ethiopia
• Pliocene paleontological sites of Africa
• Afar Region
• Archaeological sites of Eastern Africa