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Hoolock gibbons
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Primates
Suborder:	Haplorhini
Infraorder:	Simiiformes
Family:	Hylobatidae
Genus:	Hoolock Mootnick & Groves, 2005
Type species
Simia hoolock Harlan, 1834
Species
Hoolock hoolock Hoolock leuconedys Hoolock tianxing

The hoolock gibbons are three primate species of genus Hoolock in the gibbon family, Hylobatidae, native to eastern Bangladesh, Northeast India, Myanmar, and Southwest China.

Description

Hoolocks are the second-largest of the gibbons, after the siamang. They reach a size of 60 to 90 cm and weigh 6 to 9 kg. The sexes are about the same size, but they differ considerably in coloration; males are black-colored with remarkable white brows, while females have a grey-brown fur, which is darker at the chest and neck. White rings around their eyes and mouths give their faces a mask-like appearance. The Oxford English Dictionary says that the name "hoolock" is from "a language of Assam."

Distribution

In northeast India, the hoolock is found south of Brahmaputra and the North Bank areas and east of the Dibang Rivers. Its range extends into seven states covering Arunachal Pradesh, Assam, Manipur, Meghalaya, Mizoram, Nagaland, and Tripura (The seven northeastern states of India). The species are protected in India in the Hollongapar Gibbon sanctuary, which support 125 Individuals of the species, as of 2023. In Bangladesh, the western hoolock gibbons were distributed across 35 forest fragments. The largest of these (Lawachara and Kaptai) supported 42 and 84 individuals respectively, but 17 of the fragments had less than ten individuals. For a better overview the forest fragments are grouped into 13 areas. All areas are located within Chittagong and Sylhet divisions, mostly near the eastern border of the country. In China, in historical times, the skywalker gibbon was recorded in nine counties in the west bank of the Salween River in westernmost Yunnan province. Based on more recent interview surveys conducted in October–November 2008 and field surveys conducted in Mar, Apr, and Aug 2009, the current Chinese population of hoolock gibbons was found to be restricted to three counties (Baoshan, Tengchong, and Yingjiang). The population size was estimated to be less than 200 individuals. In Myanmar, all three species of hoolock gibbon can be found in the forested areas west of Thanlwin river. Of all four distribution countries, Myanmar holds the key population of all three species.

Behaviour and diet

Like the other gibbons, they are diurnal and arboreal, brachiating through the trees with their long arms. They live together in monogamous pairs, which stake out a territory. Their calls serve to locate family members and ward off other gibbons from their territory. Their diet consists mainly of fruits, insects and leaves.

Breeding and lifecycle

Young hoolocks are born after a 7-month gestation, with milky white or buff-colored hair. After about 6 months, the hair of males darkens and turns black, while the females' hair remains buff-colored throughout their lives. After 8–9 years, they are fully mature and their fur reaches its final coloration. Their life expectancy in the wild is about 25 years.

Classification

The classification of this gibbon has changed several times in the past few years. Classically, all gibbons were classified in the genus Hylobates, with the exception of the siamang. After some studies, the genus was divided into three subgenera (including the siamangs Symphalangus), and then into four (recognizing Bunopithecus as the hoolock subgenus distinct from other gibbon subgenera). These four subgenera were elevated to full genus status. However, the type species for Bunopithecus is Bunopithecus sericus, an extinct gibbon or gibbon-like ape from Sichuan, China. Very recent investigations have shown that the hoolock gibbons are not closely related to B. sericus, so they have been placed in their own genus, Hoolock. In the process, the two subspecies of hoolock gibbons have been raised to species level. A new subspecies of the western hoolock gibbon has been described recently from northeastern India, which has been named the Mishmi Hills hoolock gibbon, H. hoolock mishmiensis. A further new species, H. tianxing, with an estimated population of about 200, was discovered in southwest China in 2017.

The species of hoolock are:

Genus Hoolock – Mootnick & Groves, 2005 – three species

Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Western hoolock gibbon	Hoolock hoolock (Harlan, 1834)	India, Bangladesh, and Myanmar	Size: Habitat: Diet:	EN
Eastern hoolock gibbon	Hoolock leuconedys (Groves, 1967)	China(Yunnan) and India	Size: Habitat: Diet:	VU
Skywalker hoolock gibbon	Hoolock tianxing Fan et al., 2017	Myanmar and southwestern China	Size: Habitat: Diet:	EN  150

References

1. ^ Groves, C. P. (2005). "Order Primates". In Wilson, D. E.; Reeder, D. M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 178–179. ISBN 978-0-8018-8221-0. OCLC 62265494.
2. ^ Mootnick, A.; Groves, C. P. (2005). "A new generic name for the hoolock gibbon (Hylobatidae)". International Journal of Primatology. 26 (4): 971–976. doi:10.1007/s10764-005-5332-4. S2CID 8394136.
3. ^ Fan, P-F; He, K; Chen, X; Ortiz, A; Zhang, B; Zhao, C; Li, Y-Q; Zhang, H-B; Kimock, C; Wang, W-Z; Groves, C; Turvey, S.T; Roos, C; Helgen, K.M; Jiang, X-L (2017). "Description of a new species of Hoolock gibbon (Primates: Hylobatidae) based on integrative taxonomy" (PDF). American Journal of Primatology. 79 (5): e22631. doi:10.1002/ajp.22631. PMID 28073165. S2CID 3882019. Archived (PDF) from the original on 2018-07-22. Retrieved 2019-02-01.
4. Choudhury, Anwaruddin (1987) "Notes on the distribution and conservation of Phayre's leaf monkey and Hoolock gibbon in India." Tigerpaper 14(2): 2–6
5. Choudhury, Anwaruddin (1991). "Ecology of the hoolock gibbon (Hylobates hoolock), a lesser ape in the tropical forests of north-eastern India". Journal of Tropical Ecology. 7 (1): 147–153. doi:10.1017/S0266467400005228. JSTOR 2559687. S2CID 86606996.
6. Choudhury, Anwaruddin (2006). "The distribution and status of hoolock gibbon, Hoolock hoolock, in Manipur, Meghalaya, Mizoram, and Nagaland in Northeast India". Primate Conservation. 20: 79–87. doi:10.1896/0898-6207.20.1.79.
7. Anand, Ashvita (2023-11-25). "In Assam, railway line expansion and electrification threaten the hoolock gibbon, India's only ape". Scroll.in. Archived from the original on 2023-11-26. Retrieved 2023-11-26.
8. Geissmann, T.; Grindley, M. E.; Lwin, Ngwe; Aung, Saw Soe; Aung, Thet Naing; Htoo, Saw Blaw; Momberg, F. (2013). The Conservation Status of Hoolock Gibbons in Myanmar. Zürich, Switzerland: Gibbon Conservation Alliance. pp. xii + 157. ISBN 978-3-033-04358-9.
9. Choudhury, A. U. (2013). "Description of a new subspecies of hoolock gibbon Hoolock hoolock from North East India". The Newsletter & Journal of the Rhino Foundation for Nat. In NE India. 9: 49–59.
10. ^ Brown, Georgia (11 January 2017). "New species of gibbon discovered in China". The Guardian. Archived from the original on 27 April 2019. Retrieved 11 January 2017.

External links

• ARKive – images and movies of the hoolock gibbon (Bunopithecus hoolock)
• Hoolock gibbon photos
• Hoolock gibbon songs
• Gibbon Conservation Center
• Hoolock gibbon photos taken in the wild
• Primate Info Net Hoolock factsheet

v t e Haplorhini
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Primates
Haplorhini †"Omomyidae" Altanius? Altiatlasius? Bownomomys Kohatius Microchoerinae Indusomys Melaneremia Microchoerus Nannopithex Necrolemur Paraloris Pseudoloris Quercyloris Vectipithex "Anaptomorphinae" Absarokius Anaptomorphus Anemorhysis Arapahovius Aycrossia Chlororhysis Gazinius Pseudotetonius Sphacorhysis Strigorhysis Tatmanius Teilhardina Tetonius Tetonoides Trogolemur Walshina "Omomyinae" Ageitodendron Asiomomys Brontomomys Chipetaia Chumashius Diablomomys Dyseolemur Ekwiiyemakius Gunnelltarsius Hemiacodon Jemezius Loveina Macrotarsius Nesomomys Omomys Ourayia Rooneyia Shoshonius Steinius Stockia Uintanius Utahia Washakius Wyomomys Yaquius Tarkadectinae Tarka Tarkadectes Tarkops Tarsiiformes †Archicebus †Afrotarsiidae? †"Omomyidae"? Tarsiidae Carlito Cephalopachus †Hesperotarsius †Oligotarsius Tarsius †Xanthorhysis? Simiiformes see below↓
Simiiformes Simiiformes †Altiatlasius? †Amamria †Aseanpithecus †Nosmips? †Phileosimias? †Afrotarsiidae? Afrasia Afrotarsius †Eosimiidae Bahinia Eosimias Phenacopithecus †Amphipithecidae Amphipithecus Bugtipithecus? Ganlea Krabia Myanmarpithecus Pondaungia Siamopithecus †Parapithecoidea Arsinoea Biretia Proteopithecidae Proteopithecus Serapia Parapithecidae Abuqatrania Apidium Parapithecus Qatrania Ucayalipithecus Platyrrhini †Parvimico †Perupithecus †Szalatavus †Canaanimico †Branisella †Lagonimico Aotidae Aotus †Tremacebus Pitheciidae †Antillothrix Cacajao Callicebus †Carlocebus †Cebupithecia Cheracebus Chiropotes †Homunculus †Insulacebus †Mazzonicebus †Miocallicebus †Nuciruptor Pithecia Plecturocebus †Proteropithecia †Soriacebus †Xenothrix Atelidae Alouatta Ateles Brachyteles †Caipora †Cartelles †Chilecebus Lagothrix †Mohanamico Oreonax †Paralouatta †Protopithecus †Solimoea †Stirtonia Cebidae †Acrecebus Cebus †Dolichocebus †Killikaike †Panamacebus †Patasola Saimiri Sapajus Callitrichidae Callimico Callithrix Cebuella Leontocebus Leontopithecus Mico †Micodon Saguinus Catarrhini see below↓
Catarrhini Catarrhini †Micropithecus †Saadanius †Oligopithecidae Catopithecus Oligopithecus Talahpithecus †Propliopithecidae Aegyptopithecus Propliopithecus †Pliopithecoidea Krishnapithecus Paidopithex †Kapi Pliopithecidae Epipliopithecus Pliopithecus Dionysopithecidae Dionysopithecus Platodontopithecus Crouzeliidae Anapithecus Barberapithecus Crouzelia Egarapithecus Fanchangia Laccopithecus Plesiopliopithecus Pliobates Cercopithecoidea †Alophe †Victoriapithecidae Noropithecus Nsungwepithecus? Prohylobates? Victoriapithecus Zaltanpithecus Colobinae †Cercopithecoides Colobus †Dolichopithecus †Kuseracolobus †Libypithecus †Mesopithecus †Microcolobus Nasalis †Paracolobus †Parapresbytis Piliocolobus Presbytis Procolobus Pygathrix †Rhinocolobus Rhinopithecus †Sawecolobus Semnopithecus Simias Trachypithecus Cercopithecinae Cercopithecini Allenopithecus Allochrocebus Cercopithecus Chlorocebus Erythrocebus Miopithecus †Nanopithecus Papionini Cercocebus †Dinopithecus †Gorgopithecus Lophocebus Macaca Mandrillus Papio †Paradolichopithecus †Parapapio †Pliopapio †Procercocebus †Procynocephalus Rungwecebus †Soromandrillus Theropithecus Hominoidea see below↓
Hominoidea Hominoidea †Afropithecus †Ekembo †Equatorius †Griphopithecus †Kamoyapithecus? †Kenyapithecus †Morotopithecus †Nacholapithecus †Otavipithecus †Proconsul †Samburupithecus †Dendropithecidae Dendropithecus Micropithecus? Nyanzapithecus Oreopithecus? Rangwapithecus Rukwapithecus Simiolus Turkanapithecus Hylobatidae †Bunopithecus Hoolock Hylobates †Junzi Nomascus Symphalangus †Yuanmoupithecus Hominidae †Buronius †Graecopithecus †Nakalipithecus †Ouranopithecus Ponginae †Ankarapithecus †Gigantopithecus †Indopithecus †Khoratpithecus †Lufengpithecus †Meganthropus Pongo †Sivapithecus Homininae †Dryopithecini Anoiapithecus Danuvius Dryopithecus Hispanopithecus? Pierolapithecus? Rudapithecus Gorillini †Chororapithecus? Gorilla †Nakalipithecus? Hominini Pan Hominina †Ardipithecus? †Australopithecus †Graecopithecus? Homo †Kenyanthropus †Orrorin? †Sahelanthropus? †Paranthropus

• Hoolock
• Gibbons
• Taxa named by Colin Groves