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Hovasaurus Temporal range: 252.3–247.2 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Life restoration of Hovasaurus boulei
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Family:	†Tangasauridae
Genus:	†Hovasaurus Piveteau, 1926
Type species
†Hovasaurus boulei Piveteau, 1926

Hovasaurus is an extinct genus of basal diapsid reptile. It lived in what is now Madagascar during the Late Permian and Early Triassic, being a survivor of the Permian–Triassic extinction event and the paleontologically youngest member of the Tangasauridae. Fossils have been found in the Permian Lower and Triassic Middle Sakamena Formations of the Sakamena Group, where it is amongst the commonest fossils. Its morphology suggests an aquatic ecology.

Description

Hovasaurus resembled a slender lizard, two thirds of total length was taken up by its long tail. It had snout-vent length up to 35 centimetres (14 in) long, with 60 centimetres (24 in) long tail. It was well adapted to an aquatic life, with the tail being laterally flattened like that of a sea snake. Some stones have been found in the abdomen of fossil Hovasaurus, indicating the creatures swallowed these for ballast, preventing them from floating to the surface when hunting fish.

Paleoenvironment

The Lower Sakamena Formation was deposited in a wetland environment situated within a North-South orientated rift valley, perhaps similar to Lake Tanganyika. The climate at the time of deposition was temperate, warm, and humid, with seasonal rainfall and possible monsoons. Flora from the formation includes the equisetalean Schizoneura, Glossopteris, and seed fern Lepidopteris. Other vertebrates known from the Lower Sakamena Formation include the palaeoniscoid fish Atherstonia, the procolophonid parareptile Barasaurus, the gliding weigeltisaurid reptile Coelurosauravus, neodiapsids Claudiosaurus, Thadeosaurus, and Acerosodontosaurus, fragments of rhinesuchid temnospondyls, an indeteriminate theriodont therapsid and the dicynodont Oudenodon.

References

1. Ketchum, H. F.; Barrett, P. M. (2004). "New reptile material from the Lower Triassic of Madagascar: implications for the Permian–Triassic extinction event". Canadian Journal of Earth Sciences. 44 (1): 1–8. Bibcode:2004CaJES..41....1K. doi:10.1139/e03-084.
2. Pritchard, Adam C.; Nesbitt, Sterling J. (2017-10-01). "A bird-like skull in a Triassic diapsid reptile increases heterogeneity of the morphological and phylogenetic radiation of Diapsida". Royal Society Open Science. 4 (10): 170499. Bibcode:2017RSOS....470499P. doi:10.1098/rsos.170499. PMC 5666248. PMID 29134065.
3. Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 85. ISBN 1-84028-152-9.
4. Buffa, Valentin; Frey, Eberhard; Steyer, J.-Sébastien; Laurin, Michel (2021-07-12). "A new cranial reconstruction of Coelurosauravus elivensis Piveteau, 1926 (Diapsida, Weigeltisauridae) and its implications on the paleoecology of the first gliding vertebrates". Journal of Vertebrate Paleontology. 41 (2): e1930020. Bibcode:2021JVPal..41E0020B. doi:10.1080/02724634.2021.1930020. ISSN 0272-4634. S2CID 237517962.
5. Smith, R. M. H. 2000. Sedimentology and taphonomy of Late Permian vertebrate fossil localities in Southwestern Madagascar. Paleontologia Africana 36:25–41

Further reading

• Currie, P. J. (1981). "Hovasaurus boulei, an aquatic eosuchian from the Upper Permian of Madagascar". Palaeontologia Africana. 24: 99–168. hdl:10539/16296.
• D. Lambert, D. Naish and E. Wyse 2001, "Encyclopedia of Dinosaurs and prehistoric life", p. 77, Dorling Kindersley Limited, London. ISBN 0-7513-0955-9

v t e Sauropsida (Reptilia sensu lato)
Kingdom: Animalia Phylum: Chordata Superclass: Tetrapoda Clade: Amniota Tetrapodomorpha see Tetrapodomorpha Reptiliomorpha see Reptiliomorpha Sauropsida see below↓
Sauropsida Sauropsida †Australothyris †Erpetonyx †Microleter †Acleistorhinidae Acleistorhinus Carbonodraco Colobomycter Delorhynchus Feeserpeton Karutia Klastomycter †Bolosauridae Belebey Bolosaurus Eudibamus †Mesosauria Brazilosaurus Mesosaurus Stereosternum †Millerettidae Broomia Heleophilus Milleretta Millerosaurus Milleropsis Nanomilleretta †Procolophonia see Procolophonia Diapsida see below↓
Diapsida Diapsida †Adelosaurus? †Claudiosaurus †Cryptovaranoides †Dolerosaurus †Elachistosuchus †Eunotosaurus? †Feralisaurus †Heleosuchus †Kenyasaurus †Kudnu †Lanthanolania †Maiothisavros †Microzemiotes †Orovenator †Palacrodon †Palaeagama †Saurosternon †Sphodrosaurus †Younginidae Galesphyrus Thadeosaurus Youngina †Tangasauridae Acerosodontosaurus? Hovasaurus Tangasaurus †Weigeltisauridae Coelurosauravus Glaurung Rautiania Weigeltisaurus †Drepanosauromorpha Ancistronychus Avicranium Dolabrosaurus Drepanosaurus Hypuronector Kyrgyzsaurus Longisquama? Megalancosaurus Protoavis? Skybalonyx Unguinychus Vallesaurus †Kuehneosauridae Icarosaurus Idiosaura? Kuehneosaurus Kuehneosuchus Pamelina Rhabdopelix? †Choristodera see Choristodera †Ichthyosauromorpha see Ichthyosauromorpha †Sauropterygia see Sauropterygia †Thalattosauria see Thalattosauria Sauria (Crown-Reptilia) Lepidosauromorpha see Lepidosauromorpha Includes Lepidosauria includes rhynchocephalians and squamates Archelosauria Archosauromorpha see Archosauromorpha Includes Archosauria includes birds and crocodilians Pantestudines see Pantestudines Includes Testudines turtles
See also Possible sauropsids Araeoscelidia Captorhinidae Protorothyrididae Recumbirostra Varanopidae Other taxa Anapsida Captorhinida Eureptilia Parareptilia Lanthanosuchoidea Millerosauria Procolophonomorpha Ankyramorpha Neodiapsida Eosuchia Younginiformes Avicephala Eolacertilia Euryapsida
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• Diapsids
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• Lopingian reptiles of Africa
• Early Triassic reptiles of Africa
• Changhsingian genus first appearances
• Induan genus extinctions
• Permian Madagascar
• Triassic Madagascar
• Fossils of Madagascar
• Fossil taxa described in 1926
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