James Watson

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This is the latest accepted revision, reviewed on 18 December 2024. American molecular biologist, geneticist, and zoologist (born 1928) For other people named James Watson, see James Watson (disambiguation).

James Watson
Watson in 2012
Born	James Dewey Watson (1928-04-06) April 6, 1928 (age 96) Chicago, Illinois, U.S.
Education	University of Chicago (BS) Indiana University (PhD)
Known for	DNA structure Molecular biology
Spouse	Elizabeth Lewis (m. 1968)
Children	2
Awards	Albert Lasker Award for Basic Medical Research (1960) Nobel Prize in Physiology or Medicine (1962) John J. Carty Award (1971) Copley Medal (1993) Lomonosov Gold Medal (1994)
Scientific career
Fields	Genetics
Institutions	Indiana University Cold Spring Harbor Laboratory Laboratory of Molecular Biology Harvard University California Institute of Technology University of Cambridge National Institutes of Health
Thesis	The Biological Properties of X-Ray Inactivated Bacteriophage (1951)
Doctoral advisor	Salvador Luria
Doctoral students	Mario Capecchi Bob Horvitz Peter B. Moore David Schlessinger Joan Steitz
Other notable students	Ewan Birney Ronald W. Davis (postdoc) Phillip Allen Sharp (postdoc) Richard J. Roberts (postdoc) John Tooze (postdoc)

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James Dewey Watson (born April 6, 1928) is an American molecular biologist, geneticist, and zoologist. In 1953, he co-authored with Francis Crick the academic paper in Nature proposing the double helix structure of the DNA molecule. Watson, Crick and Maurice Wilkins were awarded the 1962 Nobel Prize in Physiology or Medicine "for their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material". He later drew global criticism with racist pronouncements on genetics, race and intelligence.

Watson earned degrees at the University of Chicago (BS, 1947) and Indiana University (PhD, 1950). Following a post-doctoral year at the University of Copenhagen with Herman Kalckar and Ole Maaløe, Watson worked at the University of Cambridge's Cavendish Laboratory in England, where he first met his future collaborator Francis Crick. From 1956 to 1976, Watson was on the faculty of the Harvard University Biology Department, promoting research in molecular biology.

From 1968, Watson served as director of Cold Spring Harbor Laboratory (CSHL), greatly expanding its level of funding and research. At CSHL, he shifted his research emphasis to the study of cancer, along with making it a world-leading research center in molecular biology. In 1994, he started as president and served for 10 years. He was then appointed chancellor, serving until he resigned in 2007 after making comments claiming that there is a genetic link between intelligence and race. In 2019, following the broadcast of a documentary in which Watson reiterated these views on race and genetics, CSHL revoked his honorary titles and severed all ties with him.

Watson has written many science books, including the textbook Molecular Biology of the Gene (1965) and his bestselling book The Double Helix (1968). Between 1988 and 1992, Watson was associated with the National Institutes of Health, helping to establish the Human Genome Project, which completed the task of mapping the human genome in 2003.

Early life and education

Watson was born in Chicago on April 6, 1928, as the only son of Jean (née Mitchell) and James D. Watson, a businessman descended mostly from colonial English immigrants to America. His mother's father, Lauchlin Mitchell, a tailor, was from Glasgow, Scotland, and her mother, Lizzie Gleason, was the child of parents from County Tipperary, Ireland. His mother was a modestly religious Catholic and his father an Episcopalian who had lost his belief in God. Watson was raised Catholic, but he later described himself as "an escapee from the Catholic religion". Watson said, "The luckiest thing that ever happened to me was that my father didn't believe in God." By age 11, Watson stopped attending mass and embraced the "pursuit of scientific and humanistic knowledge."

Watson grew up on the South Side of Chicago and attended public schools, including Horace Mann Elementary School and South Shore High School. He was fascinated with bird watching, a hobby shared with his father, so he considered majoring in ornithology. Watson appeared on Quiz Kids, a popular radio show that challenged bright youngsters to answer questions. Thanks to the liberal policy of university president Robert Hutchins, he enrolled at the University of Chicago, where he was awarded a tuition scholarship, at the age of 15. Among his professors was Louis Leon Thurstone from whom Watson learned about factor analysis, which he would later reference on his controversial views on race.

After reading Erwin Schrödinger's book, What Is Life? in 1946, Watson changed his professional ambitions from the study of ornithology to genetics. Watson earned his BS degree in zoology from the University of Chicago in 1947. In his autobiography, Avoid Boring People, Watson described the University of Chicago as an "idyllic academic institution where he was instilled with the capacity for critical thought and an ethical compulsion not to suffer fools who impeded his search for truth", in contrast to his description of later experiences. In 1947 Watson left the University of Chicago to become a graduate student at Indiana University, attracted by the presence at Bloomington of the 1946 Nobel Prize winner Hermann Joseph Muller, who in crucial papers published in 1922, 1929, and in the 1930s had laid out all the basic properties of the heredity molecule that Schrödinger presented in his 1944 book. He received his PhD degree from Indiana University in 1950; Salvador Luria was his doctoral advisor.

Career and research

Luria, Delbrück, and the Phage Group

Originally, Watson was drawn into molecular biology by the work of Salvador Luria. Luria eventually shared the 1969 Nobel Prize in Physiology or Medicine for his work on the Luria–Delbrück experiment, which concerned the nature of genetic mutations. He was part of a distributed group of researchers who were making use of the viruses that infect bacteria, called bacteriophages. He and Max Delbrück were among the leaders of this new "Phage Group", an important movement of geneticists from experimental systems such as Drosophila towards microbial genetics. Early in 1948, Watson began his PhD research in Luria's laboratory at Indiana University. That spring, he met Delbrück first in Luria's apartment and again that summer during Watson's first trip to the Cold Spring Harbor Laboratory (CSHL).

The Phage Group was the intellectual medium where Watson became a working scientist. Importantly, the members of the Phage Group sensed that they were on the path to discovering the physical nature of the gene. In 1949, Watson took a course with Felix Haurowitz that included the conventional view of that time: that genes were proteins and able to replicate themselves. The other major molecular component of chromosomes, DNA, was widely considered to be a "stupid tetranucleotide", serving only a structural role to support the proteins. Even at this early time, Watson, under the influence of the Phage Group, was aware of the Avery–MacLeod–McCarty experiment, which suggested that DNA was the genetic molecule. Watson's research project involved using X-rays to inactivate bacterial viruses.

Watson then went to Copenhagen University in September 1950 for a year of postdoctoral research, first heading to the laboratory of biochemist Herman Kalckar. Kalckar was interested in the enzymatic synthesis of nucleic acids, and he wanted to use phages as an experimental system. Watson wanted to explore the structure of DNA, and his interests did not coincide with Kalckar's. After working part of the year with Kalckar, Watson spent the remainder of his time in Copenhagen conducting experiments with microbial physiologist Ole Maaløe, then a member of the Phage Group.

The experiments, which Watson had learned of during the previous summer's Cold Spring Harbor phage conference, included the use of radioactive phosphate as a tracer to determine which molecular components of phage particles actually infect the target bacteria during viral infection. The intention was to determine whether protein or DNA was the genetic material, but upon consultation with Max Delbrück, they determined that their results were inconclusive and could not specifically identify the newly labeled molecules as DNA. Watson never developed a constructive interaction with Kalckar, but he did accompany Kalckar to a meeting in Italy, where Watson saw Maurice Wilkins talk about X-ray diffraction data for DNA. Watson was now certain that DNA had a definite molecular structure that could be elucidated.

In 1951, the chemist Linus Pauling in California published his model of the amino acid alpha helix, a result that grew out of Pauling's efforts in X-ray crystallography and molecular model building. After obtaining some results from his phage and other experimental research conducted at Indiana University, Statens Serum Institut (Denmark), CSHL, and the California Institute of Technology, Watson now had the desire to learn to perform X-ray diffraction experiments so he could work to determine the structure of DNA. That summer, Luria met John Kendrew, and he arranged for a new postdoctoral research project for Watson in England. In 1951 Watson visited the Stazione Zoologica 'Anton Dohrn' in Naples.

Identifying the double helix

In mid-March 1953, Watson and Crick deduced the double helix structure of DNA. Crucial to their discovery were the experimental data collected at King's College London—mainly by Rosalind Franklin for which they did not provide proper attribution. Sir Lawrence Bragg, the director of the Cavendish Laboratory (where Watson and Crick worked), made the original announcement of the discovery at a Solvay conference on proteins in Belgium on April 8, 1953; it went unreported by the press. Watson and Crick submitted a paper entitled "Molecular Structure of Nucleic Acids: A Structure for Deoxyribose Nucleic Acid" to the scientific journal Nature, which was published on April 25, 1953. Bragg gave a talk at the Guy's Hospital Medical School in London on Thursday, May 14, 1953, which resulted in a May 15, 1953, article by Ritchie Calder in the London newspaper News Chronicle, entitled "Why You Are You. Nearer Secret of Life".

Sydney Brenner, Jack Dunitz, Dorothy Hodgkin, Leslie Orgel, and Beryl M. Oughton were some of the first people in April 1953 to see the model of the structure of DNA, constructed by Crick and Watson; at the time, they were working at Oxford University's chemistry department. All were impressed by the new DNA model, especially Brenner, who subsequently worked with Crick at Cambridge in the Cavendish Laboratory and the new Laboratory of Molecular Biology. According to the late Beryl Oughton, later Rimmer, they all travelled together in two cars once Dorothy Hodgkin announced to them that they were off to Cambridge to see the model of the structure of DNA.

Watson's name on New York City's Nobel Monument; it lists American laureates only, not Crick and Wilkins who shared the 1962 Physics Nobel.

The Cambridge University student newspaper Varsity ran its own short article on the discovery on Saturday, May 30, 1953. Watson subsequently presented a paper on the double-helical structure of DNA at the 18th Cold Spring Harbor Symposium on Viruses in early June 1953, six weeks after the publication of the Watson and Crick paper in Nature. Many at the meeting had not yet heard of the discovery. The 1953 Cold Spring Harbor Symposium was the first opportunity for many to see the model of the DNA double helix. Watson, Crick, and Wilkins were awarded the Nobel Prize in Physiology or Medicine in 1962 for their research on the structure of nucleic acids. Rosalind Franklin had died in 1958 and was therefore ineligible for nomination. The publication of the double helix structure of DNA has been described as a turning point in science; understanding of life was fundamentally changed and the modern era of biology began.

Interactions with Rosalind Franklin and Raymond Gosling

Watson and Crick's use of DNA X-ray diffraction data collected by Rosalind Franklin and her student Raymond Gosling has attracted scrutiny. It has been argued that Watson and his colleagues did not properly acknowledge colleague Rosalind Franklin for her contributions to the discovery of the double helix structure. Robert P. Crease notes that "Such stingy behaviour may not be unknown, or even uncommon, among scientists". Franklin's high-quality X-ray diffraction patterns of DNA were unpublished results, which Watson and Crick used without her knowledge or consent in their construction of the double helix model of DNA. Franklin's results provided estimates of the water content of DNA crystals and these results were consistent with the two sugar-phosphate backbones being on the outside of the molecule. Franklin told Crick and Watson that the backbones had to be on the outside; before then, Linus Pauling and Watson and Crick had erroneous models with the chains inside and the bases pointing outwards. Her identification of the space group for DNA crystals revealed to Crick that the two DNA strands were antiparallel.

The X-ray diffraction images collected by Gosling and Franklin provided the best evidence for the helical nature of DNA. Watson and Crick had three sources for Franklin's unpublished data:

Her 1951 seminar, attended by Watson;
Discussions with Wilkins, who worked in the same laboratory with Franklin;
A research progress report that was intended to promote coordination of Medical Research Council-supported laboratories. Watson, Crick, Wilkins and Franklin all worked in MRC laboratories.

In a 1954 article, Watson and Crick acknowledged that, without Franklin's data, "the formulation of our structure would have been most unlikely, if not impossible". In The Double Helix, Watson later admitted that "Rosy, of course, did not directly give us her data. For that matter, no one at King's realized they were in our hands". In recent years, Watson has garnered controversy in the popular and scientific press for his "misogynist treatment" of Franklin and his failure to properly attribute her work on DNA. According to one critic, Watson's portrayal of Franklin in The Double Helix was negative, giving the impression that she was Wilkins' assistant and was unable to interpret her own DNA data. Watson's accusation was indefensible since Franklin told Crick and Watson that the helix backbones had to be on the outside. From a 2003 piece by Brenda Maddox in Nature:

Other comments dismissive of "Rosy" in Watson's book caught the attention of the emerging women's movement in the late 1960s. "Clearly Rosy had to go or be put in her place ... Unfortunately Maurice could not see any decent way to give Rosy the boot". And, "Certainly a bad way to go out into the foulness of a ... November night was to be told by a woman to refrain from venturing an opinion about a subject for which you were not trained."

Robert P. Crease remarks that " was close to figuring out the structure of DNA, but did not do it. The title of 'discoverer' goes to those who first fit the pieces together". Jeremy Bernstein rejects that Franklin was a "victim" and states that " made the double-helix scheme work. It is as simple as that". Matthew Cobb and Nathaniel C. Comfort write that "Franklin was no victim in how the DNA double helix was solved" but that she was "an equal contributor to the solution of the structure".

A review of the correspondence from Franklin to Watson, in the archives at CSHL, revealed that the two scientists later exchanged constructive scientific correspondence. Franklin consulted with Watson on her tobacco mosaic virus RNA research. Franklin's letters were framed with the normal and unremarkable forms of address, beginning with "Dear Jim", and concluding with "Best Wishes, Yours, Rosalind". Each of the scientists published their own unique contributions to the discovery of the structure of DNA in separate articles, and all of the contributors published their findings in the same volume of Nature. These classic molecular biology papers are identified as: Watson J. D. and Crick F. H. C. "A Structure for Deoxyribose Nucleic Acid". Nature 171, 737–738 (1953); Wilkins M. H. F., Stokes A. R. & Wilson H. R. "Molecular Structure of Deoxypentose Nucleic Acids". Nature 171, 738–740 (1953); Franklin R. and Gosling R. G. "Molecular Configuration in Sodium Thymonucleate". Nature 171, 740–741 (1953).

Harvard University

In 1956, Watson accepted a position in the biology department at Harvard University. His work at Harvard focused on RNA and its role in the transfer of genetic information.

Watson championed a switch in focus for the school from classical biology to molecular biology, stating that disciplines such as ecology, developmental biology, taxonomy, physiology, etc. had stagnated and could progress only once the underlying disciplines of molecular biology and biochemistry had elucidated their underpinnings, going so far as to discourage their study by students.

Watson continued to be a member of the Harvard faculty until 1976, even though he took over the directorship of Cold Spring Harbor Laboratory in 1968.

During his tenure at Harvard, Watson participated in a protest against the Vietnam War, leading a group of 12 biologists and biochemists calling for "the immediate withdrawal of U.S. forces from Vietnam". In 1975, on the thirtieth anniversary of the bombing of Hiroshima, Watson was one of over 2000 scientists and engineers who spoke out against nuclear proliferation to President Gerald Ford, arguing that there was no proven method for the safe disposal of radioactive waste, and that nuclear plants were a security threat due to the possibility of terrorist theft of plutonium.

Watson's first textbook, The Molecular Biology of the Gene, used the concept of heads—brief declarative subheadings. His next textbook was Molecular Biology of the Cell, in which he coordinated the work of a group of scientist-writers. His third was Recombinant DNA, which described the ways in which genetic engineering has brought new information about how organisms function.

Publishing The Double Helix

In 1968, Watson wrote The Double Helix, listed by the board of the Modern Library as number seven in their list of 100 Best Nonfiction books. The book details the story of the discovery of the structure of DNA, as well as the personalities, conflicts and controversy surrounding their work, and includes many of his private emotional impressions at the time. Watson's original title was to have been "Honest Jim". Controversy surrounded the publication of the book. Watson's book was originally to be published by the Harvard University Press, but Francis Crick and Maurice Wilkins, among others, objected. Watson's home university dropped the project and the book was commercially published. In an interview with Anne Sayre for her book, Rosalind Franklin and DNA (published in 1975 and reissued in 2000), Francis Crick said that he regarded Watson's book as a "contemptible pack of damned nonsense".

Cold Spring Harbor Laboratory

External videos

"James Watson: Why society isn't ready for genomic-based medicine", 2012, Chemical Heritage Foundation

In 1968, Watson became the director of the Cold Spring Harbor Laboratory (CSHL). Between 1970 and 1972, the Watsons' two sons were born, and by 1974, the young family made Cold Spring Harbor their permanent residence. Watson served as the laboratory's director and president for about 35 years, and later he assumed the role of chancellor and then chancellor emeritus.

In his roles as director, president, and chancellor, Watson led CSHL to articulate its present-day mission, "dedication to exploring molecular biology and genetics in order to advance the understanding and ability to diagnose and treat cancers, neurological diseases, and other causes of human suffering." CSHL substantially expanded both its research and its science educational programs under Watson's direction. He is credited with "transforming a small facility into one of the world's great education and research institutions. Initiating a program to study the cause of human cancer, scientists under his direction have made major contributions to understanding the genetic basis of cancer." In a retrospective summary of Watson's accomplishments there, Bruce Stillman, the laboratory's president, said, "Jim Watson created a research environment that is unparalleled in the world of science."

In 2007, Watson said, "I turned against the left wing because they don't like genetics, because genetics implies that sometimes in life we fail because we have bad genes. They want all failure in life to be due to the evil system."

Human Genome Project

In 1990, Watson was appointed as the head of the Human Genome Project at the National Institutes of Health, a position he held until April 10, 1992. Watson left the Genome Project after conflicts with the new NIH Director, Bernadine Healy. Watson was opposed to Healy's attempts to acquire patents on gene sequences, and any ownership of the "laws of nature". Two years before stepping down from the Genome Project, he had stated his own opinion on this long and ongoing controversy which he saw as an illogical barrier to research; he said, "The nations of the world must see that the human genome belongs to the world's people, as opposed to its nations." He left within weeks of the 1992 announcement that the NIH would be applying for patents on brain-specific cDNAs. (The issue of the patentability of genes has since been resolved in the US by the US Supreme Court; see Association for Molecular Pathology v. U.S. Patent and Trademark Office.)

In 1994, Watson became president of CSHL. Francis Collins took over the role as director of the Human Genome Project.

Watson was quoted in The Sunday Telegraph in 1997 as stating: "If you could find the gene which determines sexuality and a woman decides she doesn't want a homosexual child, well, let her." The biologist Richard Dawkins wrote a letter to The Independent claiming that Watson's position was misrepresented by The Sunday Telegraph article, and that Watson would equally consider the possibility of having a heterosexual child to be just as valid as any other reason for abortion, to emphasise that Watson is in favor of allowing choice.

On the issue of obesity, Watson was quoted in 2000, saying: "Whenever you interview fat people, you feel bad, because you know you're not going to hire them."

Watson has repeatedly supported genetic screening and genetic engineering in public lectures and interviews, arguing that stupidity is a disease and the "really stupid" bottom 10% of people should be cured. He has also suggested that beauty could be genetically engineered, saying in 2003, "People say it would be terrible if we made all girls pretty. I think it would be great."

In 2007, Watson became the second person to publish his fully sequenced genome online, after it was presented to him on May 31, 2007, by 454 Life Sciences Corporation in collaboration with scientists at the Human Genome Sequencing Center, Baylor College of Medicine. Watson was quoted as saying, "I am putting my genome sequence on line to encourage the development of an era of personalized medicine, in which information contained in our genomes can be used to identify and prevent disease and to create individualized medical therapies".

Later life

In 2014, Watson published a paper in The Lancet suggesting that biological oxidants may have a different role than is thought in diseases including diabetes, dementia, heart disease and cancer. For example, type 2 diabetes is usually thought to be caused by oxidation in the body that causes inflammation and kills off pancreatic cells. Watson thinks the root of that inflammation is different: "a lack of biological oxidants, not an excess", and discusses this in detail. One critical response was that the idea was neither new nor worthy of merit, and that The Lancet published Watson's paper only because of his name. Other scientists have expressed their support for his hypothesis and have proposed that it can also be expanded to why a lack of oxidants can result in cancer and its progression.

In 2014, Watson sold his Nobel Prize medal to raise money after complaining of being made an "unperson" following controversial statements he had made. Part of the funds raised by the sale went to support scientific research. The medal sold at auction at Christie's in December 2014 for US$4.1 million. Watson intended to contribute the proceeds to conservation work in Long Island and to funding research at Trinity College, Dublin. He was the first living Nobel recipient to auction a medal. The medal was later returned to Watson by the purchaser, Alisher Usmanov.

Notable former students

Several of Watson's former doctoral students subsequently became notable in their own right including, Mario Capecchi, Bob Horvitz, Peter B. Moore and Joan Steitz. Besides numerous PhD students, Watson also supervised postdoctoral researchers and other interns including Ewan Birney, Ronald W. Davis, Phillip Allen Sharp (postdoc), John Tooze (postdoc) and Richard J. Roberts (postdoc).

Other affiliations

Watson is a former member of the Board of Directors of United Biomedical, Inc., founded by Chang Yi Wang. He held the position for six years and retired from the board in 1999.

In January 2007, Watson accepted the invitation of Leonor Beleza, president of the Champalimaud Foundation, to become the head of the foundation's scientific council, an advisory organ.

In March 2017, Watson was named head consultant of the Cheerland Investment Group, a Chinese investment company which sponsored his trip.

Watson has also been an institute adviser for the Allen Institute for Brain Science.

Avoid Boring People

Watson has had disagreements with Craig Venter regarding his use of EST fragments while Venter worked at NIH. Venter went on to found Celera genomics and continued his feud with Watson. Watson was quoted as calling Venter "Hitler".

In his 2007 memoir, Avoid Boring People: Lessons from a Life in Science, Watson describes his academic colleagues as "dinosaurs", "deadbeats", "fossils", "has-beens", "mediocre", and "vapid". Steve Shapin in Harvard Magazine noted that Watson had written an unlikely "Book of Manners", telling about the skills needed at different times in a scientist's career; he wrote Watson was known for aggressively pursuing his own goals at the university. E. O. Wilson once described Watson as "the most unpleasant human being I had ever met", but in a later TV interview said that he considered them friends and their rivalry at Harvard "old history" (when they had competed for funding in their respective fields).

In the epilogue to the memoir Avoid Boring People, Watson alternately attacks and defends former Harvard University president Lawrence Summers, who stepped down in 2006 due in part to his remarks about women and science. Watson also states in the epilogue, "Anyone sincerely interested in understanding the imbalance in the representation of men and women in science must reasonably be prepared at least to consider the extent to which nature may figure, even with the clear evidence that nurture is strongly implicated."

Comments on race

At a conference in 2000, Watson suggested a link between skin color and sex drive, hypothesizing that dark-skinned people have stronger libidos. His lecture argued that extracts of melanin—which gives skin its color—had been found to boost subjects' sex drive. "That's why you have Latin lovers", he said, according to people who attended the lecture. "You've never heard of an English lover. Only an English Patient." He has also said that stereotypes associated with racial and ethnic groups have a genetic basis: Jews being intelligent, Chinese being intelligent but not creative because of selection for conformity, and Indians being servile because of selection under caste endogamy. Regarding intelligence differences between blacks and whites, Watson has asserted that "all our social policies are based on the fact that their (blacks) intelligence is the same as ours (whites) – whereas all the testing says not really ... people who have to deal with black employees find this not true."

Watson has repeatedly asserted that differences in average measured IQ between blacks and whites are due to genetics. In early October 2007, he was interviewed by Charlotte Hunt-Grubbe at Cold Spring Harbor Laboratory (CSHL). He discussed his view that Africans are less intelligent than Westerners. Watson said his intention was to promote science, not racism, but some UK venues canceled his appearances, and he canceled the rest of his tour. An editorial in Nature said that his remarks were "beyond the pale" but expressed a wish that the tour had not been canceled so that Watson would have had to face his critics in person, encouraging scientific discussion on the matter. Because of the controversy, the board of trustees at Cold Spring Harbor Laboratory suspended Watson's administrative responsibilities. Watson issued an apology, then retired at the age of 79 from CSHL from what the lab called "nearly 40 years of distinguished service". Watson attributed his retirement to his age and to circumstances that he could never have anticipated or desired.

In 2008, Watson was appointed chancellor emeritus of CSHL but continued to advise and guide project work at the laboratory. In a BBC documentary that year, Watson said he did not see himself as a racist.

In January 2019, following the broadcast of a television documentary made the previous year in which he repeated his views about race and genetics, CSHL revoked honorary titles that it had awarded to Watson and cut all remaining ties with him. Watson did not respond to the developments.

Personal life

Watson is an atheist. In 2003, he was one of 22 Nobel Laureates who signed the Humanist Manifesto. Watson wrote in Time that he contributed $1,000 to Bernie Sanders' 2016 presidential campaign.

Marriage and family

Watson married Elizabeth Lewis in 1968. They have two sons, Rufus Robert Watson (b. 1970) and Duncan James Watson (b. 1972). Watson sometimes talks about his son Rufus, who has schizophrenia, seeking to encourage progress in the understanding and treatment of mental illness by determining how genetics contributes to it.

Awards and honors

Watson has won numerous awards, including:

Albert Lasker Award for Basic Medical Research, 1960
Benjamin Franklin Medal for Distinguished Achievement in the Sciences (2001)
Copley Medal of the Royal Society, 1993
CSHL Double Helix Medal Honoree, 2008
Eli Lilly Award in Biological Chemistry, 1960
EMBO Membership in 1985

Gairdner Foundation International Award, 2002
Honorary Member of Royal Irish Academy, 2005
Honorary Fellow, the Hastings Center, an independent bioethics research institution
Honorary Knight Commander of the Order of the British Empire (KBE), 2002
Irish America Hall of Fame, inducted March 2011
John J. Carty Award in molecular biology from the National Academy of Sciences
Liberty Medal, 2000
Lomonosov Gold Medal, 1994
Lotos Club Medal of Merit, 2004
National Medal of Science, 1997
Nobel Prize in Physiology or Medicine, 1962
Othmer Gold Medal (2005)
Presidential Medal of Freedom, 1977
Golden Plate Award of the American Academy of Achievement, 1986

Honorary degrees received

DSc, University of Chicago, US, 1961
DSc, Indiana University, US, 1963
LLD, University of Notre Dame, US, 1965
DSc, Long Island University (CW Post), US, 1970
DSc, Adelphi University, US, 1972
DSc, Brandeis University, US, 1973
DSc, Albert Einstein College of Medicine, US, 1974
DSc, Hofstra University, US, 1976
DSc, Harvard University, US, 1978
DSc, Rockefeller University, US, 1980
DSc, Clarkson College of Technology, US, 1981
DSc, SUNY at Farmingdale, US, 1983
MD, Buenos Aires, Argentina, 1986
DSc, Rutgers University, US, 1988
DSc, Bard College, US, 1991
DSc, University of Stellenbosch, South Africa, 1993
DSc, Fairfield University, US, 1993
DSc, University of Cambridge, United Kingdom, 1993
DrHC, Charles University in Prague, Czech Republic, 1998
ScD, University of Dublin, Ireland, 2001

Professional and honorary affiliations

American Academy of Arts and Sciences
American Association for Cancer Research
American Philosophical Society
American Society of Biological Chemists
Athenaeum Club, London, member
Cambridge University, Honorary Fellow, Clare College, Cambridge
Cold Spring Harbor Laboratory, Chancellor Emeritus; Honorary Trustee; Oliver R. Grace Professor Emeritus (all revoked in 2019)
European Molecular Biology Organization, member since 1985
National Academy of Sciences
Oxford University, Newton-Abraham Visiting Professor
Royal Danish Academy of Sciences and Letters
Royal Society, Foreign Member of the Royal Society (ForMemRS) since 1981
Russian Academy of Sciences

References

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"All text published under the heading 'Biography' on Fellow profile pages is available under Creative Commons Attribution 4.0 International License." --"Royal Society Terms, conditions and policies". Archived from the original on September 25, 2015. Retrieved March 9, 2016.{{cite web}}: CS1 maint: bot: original URL status unknown (link)
"Copley Medal". Royal Society website. The Royal Society. Retrieved April 19, 2013.
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External links

James D. Watson Collection at the Cold Spring Harbor Laboratory Library
DNA – The Double Helix Game from Nobelprize.org
MSN Encarta biography (Archived 2009-10-31)
DNA Interactive – This site from the Dolan DNA Learning Center (part of CSHL) commemorates the discovery of the structure of DNA and includes dozens of animations, as well as interviews with James Watson and others.
DNA from the Beginning – another DNA Learning Center site on the basics of DNA, genes, and heredity, from Mendel to the Human Genome Project.
Appearances on C-SPAN
James Watson on Charlie Rose
James Watson at TED
James Watson at IMDb
James D. Watson collected news and commentary at The New York Times
- A Revolution at 50, February 25, 2003
James Watson on Nobelprize.org

Articles and interviews

BBC Four Interviews Archived December 12, 2011, at the Wayback Machine – Watson and Crick speaking on the BBC in 1962, 1972, and 1974.
NPR Science Friday: "A Conversation with Genetics Pioneer James Watson" – Ira Flatow interviews Watson on the history of DNA and his recent book A Passion for DNA: Genes, Genomes, and Society. 2002-06-02
NPR Science Friday "DNA: The Secret of Life" – Ira Flatow interviews Watson on his new book. 2003-05-02
Discover "Reversing Bad Truths" – David Duncan interviews Watson. 2003-07-01
Two remembrances of James Watson by one of the founders of molecular genetics, Esther Lederberg, can be found at http://www.estherlederberg.com/Anecdotes.html#WATSON1 and http://www.estherlederberg.com/Anecdotes.html#WATSON2
James Watson telling his life story at Web of Stories
American Masters: Decoding Watson PBS film about Watson, including extensive interviews with him, his family, and colleagues. 2019-01-02.
James D. Watson, Ph.D. Biography and Interview on American Academy of Achievement

v t e Fellows of the Royal Society elected in 1981
Fellows	John Barker Eric Barnard Fraser John Bergersen Peter Bradshaw David Brink Fred Brown Ian Butterworth Brandon Carter Malcolm Clarke Robert Clayton John Conway Rex Dawson Dennis Evans Frank Farmer Ned Franklin Christopher Graham Norman Green Herbert Gutfreund Allan Hay John Hill Kenneth Holmes Robert Honeycombe Michael Horne Yuet Wai Kan Robert Langlands Simon Maddrell Ian McGregor John Nelder Mike O'Hara Autar Singh Paintal John Phillips Edward Roy Pike Walter Plowright Ken Pounds Dai Rees Colin Reese Edward Reich Wal Sargent John Taylor David Wheeler
Foreign	Britton Chance Igor Shafarevich James Watson Steven Weinberg

Fellows of the Royal Society elected in 1981

Fellows

Foreign

v t e Laureates of the Nobel Prize in Physiology or Medicine
1901–1925	1901: Emil Behring 1902: Ronald Ross 1903: Niels Finsen 1904: Ivan Pavlov 1905: Robert Koch 1906: Camillo Golgi / Santiago Ramón y Cajal 1907: Alphonse Laveran 1908: Élie Metchnikoff / Paul Ehrlich 1909: Emil Kocher 1910: Albrecht Kossel 1911: Allvar Gullstrand 1912: Alexis Carrel 1913: Charles Richet 1914: Róbert Bárány 1915 1916 1917 1918 1919: Jules Bordet 1920: August Krogh 1921 1922: Archibald Hill / Otto Meyerhof 1923: Frederick Banting / John Macleod 1924: Willem Einthoven 1925
1926–1950	1926: Johannes Fibiger 1927: Julius Wagner-Jauregg 1928: Charles Nicolle 1929: Christiaan Eijkman / Frederick Gowland Hopkins 1930: Karl Landsteiner 1931: Otto Warburg 1932: Charles Scott Sherrington / Edgar Adrian 1933: Thomas Morgan 1934: George Whipple / George Minot / William Murphy 1935: Hans Spemann 1936: Henry Dale / Otto Loewi 1937: Albert Szent-Györgyi 1938: Corneille Heymans 1939: Gerhard Domagk 1940 1941 1942 1943: Henrik Dam / Edward Doisy 1944: Joseph Erlanger / Herbert Gasser 1945: Alexander Fleming / Ernst Chain / Howard Florey 1946: Hermann Muller 1947: Carl Cori / Gerty Cori / Bernardo Houssay 1948: Paul Müller 1949: Walter Hess / António Egas Moniz 1950: Edward Kendall / Tadeusz Reichstein / Philip Hench
1951–1975	1951: Max Theiler 1952: Selman Waksman 1953: Hans Krebs / Fritz Lipmann 1954: John Enders / Thomas Weller / Frederick Robbins 1955: Hugo Theorell 1956: André Cournand / Werner Forssmann / Dickinson W. Richards 1957: Daniel Bovet 1958: George Beadle / Edward Tatum / Joshua Lederberg 1959: Severo Ochoa / Arthur Kornberg 1960: Frank Burnet / Peter Medawar 1961: Georg von Békésy 1962: Francis Crick / James Watson / Maurice Wilkins 1963: John Eccles / Alan Hodgkin / Andrew Huxley 1964: Konrad Bloch / Feodor Lynen 1965: François Jacob / André Lwoff / Jacques Monod 1966: Francis Rous / Charles B. Huggins 1967: Ragnar Granit / Haldan Hartline / George Wald 1968: Robert W. Holley / Har Khorana / Marshall Nirenberg 1969: Max Delbrück / Alfred Hershey / Salvador Luria 1970: Bernard Katz / Ulf von Euler / Julius Axelrod 1971: Earl Sutherland Jr. 1972: Gerald Edelman / Rodney Porter 1973: Karl von Frisch / Konrad Lorenz / Nikolaas Tinbergen 1974: Albert Claude / Christian de Duve / George Palade 1975: David Baltimore / Renato Dulbecco / Howard Temin
1976–2000	1976: Baruch Blumberg / Daniel Gajdusek 1977: Roger Guillemin / Andrew Schally / Rosalyn Yalow 1978: Werner Arber / Daniel Nathans / Hamilton O. Smith 1979: Allan Cormack / Godfrey Hounsfield 1980: Baruj Benacerraf / Jean Dausset / George Snell 1981: Roger Sperry / David H. Hubel / Torsten Wiesel 1982: Sune Bergström / Bengt I. Samuelsson / John Vane 1983: Barbara McClintock 1984: Niels Jerne / Georges Köhler / César Milstein 1985: Michael Brown / Joseph L. Goldstein 1986: Stanley Cohen / Rita Levi-Montalcini 1987: Susumu Tonegawa 1988: James W. Black / Gertrude B. Elion / George H. Hitchings 1989: J. Michael Bishop / Harold E. Varmus 1990: Joseph Murray / E. Donnall Thomas 1991: Erwin Neher / Bert Sakmann 1992: Edmond Fischer / Edwin G. Krebs 1993: Richard J. Roberts / Phillip Sharp 1994: Alfred G. Gilman / Martin Rodbell 1995: Edward B. Lewis / Christiane Nüsslein-Volhard / Eric F. Wieschaus 1996: Peter C. Doherty / Rolf M. Zinkernagel 1997: Stanley B. Prusiner 1998: Robert F. Furchgott / Louis Ignarro / Ferid Murad 1999: Günter Blobel 2000: Arvid Carlsson / Paul Greengard / Eric Kandel
2001–present	2001: Leland H. Hartwell / Tim Hunt / Paul Nurse 2002: Sydney Brenner / H. Robert Horvitz / John E. Sulston 2003: Paul Lauterbur / Peter Mansfield 2004: Richard Axel / Linda B. Buck 2005: Barry Marshall / Robin Warren 2006: Andrew Fire / Craig Mello 2007: Mario Capecchi / Martin Evans / Oliver Smithies 2008: Harald zur Hausen / Luc Montagnier / Françoise Barré-Sinoussi 2009: Elizabeth Blackburn / Carol W. Greider / Jack W. Szostak 2010: Robert G. Edwards 2011: Bruce Beutler / Jules A. Hoffmann / Ralph M. Steinman (posthumously) 2012: John Gurdon / Shinya Yamanaka 2013: James Rothman / Randy Schekman / Thomas C. Südhof 2014: John O'Keefe / May-Britt Moser / Edvard Moser 2015: William C. Campbell / Satoshi Ōmura / Tu Youyou 2016: Yoshinori Ohsumi 2017: Jeffrey C. Hall / Michael Rosbash / Michael W. Young 2018: James P. Allison / Tasuku Honjo 2019: Gregg L. Semenza / Peter J. Ratcliffe / William Kaelin Jr. 2020: Harvey J. Alter / Michael Houghton / Charles M. Rice 2021: David Julius / Ardem Patapoutian 2022: Svante Pääbo 2023: Katalin Karikó / Drew Weissman 2024: Victor Ambros / Gary Ruvkun

v t e 1962 Nobel Prize laureates
Chemistry	Max Perutz (Great Britain) John Kendrew (Great Britain)
Literature (1962)	John Steinbeck (United States)
Peace	Linus Pauling (United States)
Physics	Lev Landau (Soviet Union)
Physiology or Medicine	Francis Crick (Great Britain) James D. Watson (United States) Maurice Wilkins (Great Britain/Australia)
Nobel Prize recipients 1957 1958 1959 1960 1961 1962 1963 1964 1965 1966 1967

United States National Medal of Science laureates

Behavioral and social science

1960s	1964 Neal Elgar Miller
1980s	1986 Herbert A. Simon 1987 Anne Anastasi George J. Stigler 1988 Milton Friedman
1990s	1990 Leonid Hurwicz Patrick Suppes 1991 George A. Miller 1992 Eleanor J. Gibson 1994 Robert K. Merton 1995 Roger N. Shepard 1996 Paul Samuelson 1997 William K. Estes 1998 William Julius Wilson 1999 Robert M. Solow
2000s	2000 Gary Becker 2003 R. Duncan Luce 2004 Kenneth Arrow 2005 Gordon H. Bower 2008 Michael I. Posner 2009 Mortimer Mishkin
2010s	2011 Anne Treisman 2014 Robert Axelrod 2015 Albert Bandura

Biological sciences

1960s	1963 C. B. van Niel 1964 Theodosius Dobzhansky Marshall W. Nirenberg 1965 Francis P. Rous George G. Simpson Donald D. Van Slyke 1966 Edward F. Knipling Fritz Albert Lipmann William C. Rose Sewall Wright 1967 Kenneth S. Cole Harry F. Harlow Michael Heidelberger Alfred H. Sturtevant 1968 Horace Barker Bernard B. Brodie Detlev W. Bronk Jay Lush Burrhus Frederic Skinner 1969 Robert Huebner Ernst Mayr
1970s	1970 Barbara McClintock Albert B. Sabin 1973 Daniel I. Arnon Earl W. Sutherland Jr. 1974 Britton Chance Erwin Chargaff James V. Neel James Augustine Shannon 1975 Hallowell Davis Paul Gyorgy Sterling B. Hendricks Orville Alvin Vogel 1976 Roger Guillemin Keith Roberts Porter Efraim Racker E. O. Wilson 1979 Robert H. Burris Elizabeth C. Crosby Arthur Kornberg Severo Ochoa Earl Reece Stadtman George Ledyard Stebbins Paul Alfred Weiss
1980s	1981 Philip Handler 1982 Seymour Benzer Glenn W. Burton Mildred Cohn 1983 Howard L. Bachrach Paul Berg Wendell L. Roelofs Berta Scharrer 1986 Stanley Cohen Donald A. Henderson Vernon B. Mountcastle George Emil Palade Joan A. Steitz 1987 Michael E. DeBakey Theodor O. Diener Harry Eagle Har Gobind Khorana Rita Levi-Montalcini 1988 Michael S. Brown Stanley Norman Cohen Joseph L. Goldstein Maurice R. Hilleman Eric R. Kandel Rosalyn Sussman Yalow 1989 Katherine Esau Viktor Hamburger Philip Leder Joshua Lederberg Roger W. Sperry Harland G. Wood
1990s	1990 Baruj Benacerraf Herbert W. Boyer Daniel E. Koshland Jr. Edward B. Lewis David G. Nathan E. Donnall Thomas 1991 Mary Ellen Avery G. Evelyn Hutchinson Elvin A. Kabat Robert W. Kates Salvador Luria Paul A. Marks Folke K. Skoog Paul C. Zamecnik 1992 Maxine Singer Howard Martin Temin 1993 Daniel Nathans Salome G. Waelsch 1994 Thomas Eisner Elizabeth F. Neufeld 1995 Alexander Rich 1996 Ruth Patrick 1997 James Watson Robert A. Weinberg 1998 Bruce Ames Janet Rowley 1999 David Baltimore Jared Diamond Lynn Margulis
2000s	2000 Nancy C. Andreasen Peter H. Raven Carl Woese 2001 Francisco J. Ayala George F. Bass Mario R. Capecchi Ann Graybiel Gene E. Likens Victor A. McKusick Harold Varmus 2002 James E. Darnell Evelyn M. Witkin 2003 J. Michael Bishop Solomon H. Snyder Charles Yanofsky 2004 Norman E. Borlaug Phillip A. Sharp Thomas E. Starzl 2005 Anthony Fauci Torsten N. Wiesel 2006 Rita R. Colwell Nina Fedoroff Lubert Stryer 2007 Robert J. Lefkowitz Bert W. O'Malley 2008 Francis S. Collins Elaine Fuchs J. Craig Venter 2009 Susan L. Lindquist Stanley B. Prusiner
2010s	2010 Ralph L. Brinster Rudolf Jaenisch 2011 Lucy Shapiro Leroy Hood Sallie Chisholm 2012 May Berenbaum Bruce Alberts 2013 Rakesh K. Jain 2014 Stanley Falkow Mary-Claire King Simon Levin

Chemistry

1960s	1964 Roger Adams
1980s	1982 F. Albert Cotton Gilbert Stork 1983 Roald Hoffmann George C. Pimentel Richard N. Zare 1986 Harry B. Gray Yuan Tseh Lee Carl S. Marvel Frank H. Westheimer 1987 William S. Johnson Walter H. Stockmayer Max Tishler 1988 William O. Baker Konrad E. Bloch Elias J. Corey 1989 Richard B. Bernstein Melvin Calvin Rudolph A. Marcus Harden M. McConnell
1990s	1990 Elkan Blout Karl Folkers John D. Roberts 1991 Ronald Breslow Gertrude B. Elion Dudley R. Herschbach Glenn T. Seaborg 1992 Howard E. Simmons Jr. 1993 Donald J. Cram Norman Hackerman 1994 George S. Hammond 1995 Thomas Cech Isabella L. Karle 1996 Norman Davidson 1997 Darleane C. Hoffman Harold S. Johnston 1998 John W. Cahn George M. Whitesides 1999 Stuart A. Rice John Ross Susan Solomon
2000s	2000 John D. Baldeschwieler Ralph F. Hirschmann 2001 Ernest R. Davidson Gábor A. Somorjai 2002 John I. Brauman 2004 Stephen J. Lippard 2005 Tobin J. Marks 2006 Marvin H. Caruthers Peter B. Dervan 2007 Mostafa A. El-Sayed 2008 Joanna Fowler JoAnne Stubbe 2009 Stephen J. Benkovic Marye Anne Fox
2010s	2010 Jacqueline K. Barton Peter J. Stang 2011 Allen J. Bard M. Frederick Hawthorne 2012 Judith P. Klinman Jerrold Meinwald 2013 Geraldine L. Richmond 2014 A. Paul Alivisatos

Engineering sciences

1960s	1962 Theodore von Kármán 1963 Vannevar Bush John Robinson Pierce 1964 Charles S. Draper Othmar H. Ammann 1965 Hugh L. Dryden Clarence L. Johnson Warren K. Lewis 1966 Claude E. Shannon 1967 Edwin H. Land Igor I. Sikorsky 1968 J. Presper Eckert Nathan M. Newmark 1969 Jack St. Clair Kilby
1970s	1970 George E. Mueller 1973 Harold E. Edgerton Richard T. Whitcomb 1974 Rudolf Kompfner Ralph Brazelton Peck Abel Wolman 1975 Manson Benedict William Hayward Pickering Frederick E. Terman Wernher von Braun 1976 Morris Cohen Peter C. Goldmark Erwin Wilhelm Müller 1979 Emmett N. Leith Raymond D. Mindlin Robert N. Noyce Earl R. Parker Simon Ramo
1980s	1982 Edward H. Heinemann Donald L. Katz 1983 Bill Hewlett George Low John G. Trump 1986 Hans Wolfgang Liepmann Tung-Yen Lin Bernard M. Oliver 1987 Robert Byron Bird H. Bolton Seed Ernst Weber 1988 Daniel C. Drucker Willis M. Hawkins George W. Housner 1989 Harry George Drickamer Herbert E. Grier
1990s	1990 Mildred Dresselhaus Nick Holonyak Jr. 1991 George H. Heilmeier Luna B. Leopold H. Guyford Stever 1992 Calvin F. Quate John Roy Whinnery 1993 Alfred Y. Cho 1994 Ray W. Clough 1995 Hermann A. Haus 1996 James L. Flanagan C. Kumar N. Patel 1998 Eli Ruckenstein 1999 Kenneth N. Stevens
2000s	2000 Yuan-Cheng B. Fung 2001 Andreas Acrivos 2002 Leo Beranek 2003 John M. Prausnitz 2004 Edwin N. Lightfoot 2005 Jan D. Achenbach 2006 Robert S. Langer 2007 David J. Wineland 2008 Rudolf E. Kálmán 2009 Amnon Yariv
2010s	2010 Shu Chien 2011 John B. Goodenough 2012 Thomas Kailath

Mathematical, statistical, and computer sciences

1960s	1963 Norbert Wiener 1964 Solomon Lefschetz H. Marston Morse 1965 Oscar Zariski 1966 John Milnor 1967 Paul Cohen 1968 Jerzy Neyman 1969 William Feller
1970s	1970 Richard Brauer 1973 John Tukey 1974 Kurt Gödel 1975 John W. Backus Shiing-Shen Chern George Dantzig 1976 Kurt Otto Friedrichs Hassler Whitney 1979 Joseph L. Doob Donald E. Knuth
1980s	1982 Marshall H. Stone 1983 Herman Goldstine Isadore Singer 1986 Peter Lax Antoni Zygmund 1987 Raoul Bott Michael Freedman 1988 Ralph E. Gomory Joseph B. Keller 1989 Samuel Karlin Saunders Mac Lane Donald C. Spencer
1990s	1990 George F. Carrier Stephen Cole Kleene John McCarthy 1991 Alberto Calderón 1992 Allen Newell 1993 Martin David Kruskal 1994 John Cocke 1995 Louis Nirenberg 1996 Richard Karp Stephen Smale 1997 Shing-Tung Yau 1998 Cathleen Synge Morawetz 1999 Felix Browder Ronald R. Coifman
2000s	2000 John Griggs Thompson Karen Uhlenbeck 2001 Calyampudi R. Rao Elias M. Stein 2002 James G. Glimm 2003 Carl R. de Boor 2004 Dennis P. Sullivan 2005 Bradley Efron 2006 Hyman Bass 2007 Leonard Kleinrock Andrew J. Viterbi 2009 David B. Mumford
2010s	2010 Richard A. Tapia S. R. Srinivasa Varadhan 2011 Solomon W. Golomb Barry Mazur 2012 Alexandre Chorin David Blackwell 2013 Michael Artin

Physical sciences

1960s	1963 Luis W. Alvarez 1964 Julian Schwinger Harold Urey Robert Burns Woodward 1965 John Bardeen Peter Debye Leon M. Lederman William Rubey 1966 Jacob Bjerknes Subrahmanyan Chandrasekhar Henry Eyring John H. Van Vleck Vladimir K. Zworykin 1967 Jesse Beams Francis Birch Gregory Breit Louis Hammett George Kistiakowsky 1968 Paul Bartlett Herbert Friedman Lars Onsager Eugene Wigner 1969 Herbert C. Brown Wolfgang Panofsky
1970s	1970 Robert H. Dicke Allan R. Sandage John C. Slater John A. Wheeler Saul Winstein 1973 Carl Djerassi Maurice Ewing Arie Jan Haagen-Smit Vladimir Haensel Frederick Seitz Robert Rathbun Wilson 1974 Nicolaas Bloembergen Paul Flory William Alfred Fowler Linus Carl Pauling Kenneth Sanborn Pitzer 1975 Hans A. Bethe Joseph O. Hirschfelder Lewis Sarett Edgar Bright Wilson Chien-Shiung Wu 1976 Samuel Goudsmit Herbert S. Gutowsky Frederick Rossini Verner Suomi Henry Taube George Uhlenbeck 1979 Richard P. Feynman Herman Mark Edward M. Purcell John Sinfelt Lyman Spitzer Victor F. Weisskopf
1980s	1982 Philip W. Anderson Yoichiro Nambu Edward Teller Charles H. Townes 1983 E. Margaret Burbidge Maurice Goldhaber Helmut Landsberg Walter Munk Frederick Reines Bruno B. Rossi J. Robert Schrieffer 1986 Solomon J. Buchsbaum H. Richard Crane Herman Feshbach Robert Hofstadter Chen-Ning Yang 1987 Philip Abelson Walter Elsasser Paul C. Lauterbur George Pake James A. Van Allen 1988 D. Allan Bromley Paul Ching-Wu Chu Walter Kohn Norman Foster Ramsey Jr. Jack Steinberger 1989 Arnold O. Beckman Eugene Parker Robert Sharp Henry Stommel
1990s	1990 Allan M. Cormack Edwin M. McMillan Robert Pound Roger Revelle 1991 Arthur L. Schawlow Ed Stone Steven Weinberg 1992 Eugene M. Shoemaker 1993 Val Fitch Vera Rubin 1994 Albert Overhauser Frank Press 1995 Hans Dehmelt Peter Goldreich 1996 Wallace S. Broecker 1997 Marshall Rosenbluth Martin Schwarzschild George Wetherill 1998 Don L. Anderson John N. Bahcall 1999 James Cronin Leo Kadanoff
2000s	2000 Willis E. Lamb Jeremiah P. Ostriker Gilbert F. White 2001 Marvin L. Cohen Raymond Davis Jr. Charles Keeling 2002 Richard Garwin W. Jason Morgan Edward Witten 2003 G. Brent Dalrymple Riccardo Giacconi 2004 Robert N. Clayton 2005 Ralph A. Alpher Lonnie Thompson 2006 Daniel Kleppner 2007 Fay Ajzenberg-Selove Charles P. Slichter 2008 Berni Alder James E. Gunn 2009 Yakir Aharonov Esther M. Conwell Warren M. Washington
2010s	2011 Sidney Drell Sandra Faber Sylvester James Gates 2012 Burton Richter Sean C. Solomon 2014 Shirley Ann Jackson

v t e Copley Medallists (1951–2000)
David Keilin (1951) Paul Dirac (1952) Albert Kluyver (1953) E. T. Whittaker (1954) Ronald Fisher (1955) Patrick Blackett (1956) Howard Florey (1957) John Edensor Littlewood (1958) Macfarlane Burnet (1959) Harold Jeffreys (1960) Hans Krebs (1961) Cyril Norman Hinshelwood (1962) Paul Fildes (1963) Sydney Chapman (1964) Alan Hodgkin (1965) Lawrence Bragg (1966) Bernard Katz (1967) Tadeusz Reichstein (1968) Peter Medawar (1969) Alexander R. Todd (1970) Norman Pirie (1971) Nevill Francis Mott (1972) Andrew Huxley (1973) W. V. D. Hodge (1974) Francis Crick (1975) Dorothy Hodgkin (1976) Frederick Sanger (1977) Robert Burns Woodward (1978) Max Perutz (1979) Derek Barton (1980) Peter D. Mitchell (1981) John Cornforth (1982) Rodney Robert Porter (1983) Subrahmanyan Chandrasekhar (1984) Aaron Klug (1985) Rudolf Peierls (1986) Robin Hill (1987) Michael Atiyah (1988) César Milstein (1989) Abdus Salam (1990) Sydney Brenner (1991) George Porter (1992) James D. Watson (1993) Frederick Charles Frank (1994) Frank Fenner (1995) Alan Cottrell (1996) Hugh Huxley (1997) James Lighthill (1998) John Maynard Smith (1999) Alan Battersby (2000)

Copley Medallists (1951–2000)

v t e Eli Lilly Award in Biological Chemistry
William M. Allen (1935) Harold S. Alcott (1937) Abraham White (1938) George Wald (1939) Eric G. Ball (1940) David Rittenberg (1941) Earl A. Evans, Jr. (1942) Herbert E. Carter (1943) Joseph S. Fruton (1944) Max A. Lauffer (1945) John D. Ferry (1946) Sidney Colowick (1947) Dilworth Woodley (1948) Irving M. Klotz (1949) William Shive (1950) John M. Buchanan (1951) David M. Bonner (1952) Nathan O. Kaplan (1953) Harvey Itano (1954) William F. Neuman (1955) Robert A. Alberty (1956) Harold A. Scheraga (1957) Lester J. Reed (1958) Paul Berg (1959) James Watson (1960) Frederick Crane (1961) Jerard Hurwitz (1962) William P. Jencks (1963) Bruce Ames (1964) Gerald M. Edelman (1965) Phillips W. Robbins (1966) Gordon G. Hammes (1967) Charles C. Richardson (1968) Mario R. Capecchi (1969) Lubert Stryer (1970) David F. Wilson (1971) Bruce M. Alberts (1972) C. Fred Fox (1973) James E. Dahlberg (1974) Mark Ptashne (1975) Joan A. Steitz (1976) Robert G. Roeder (1977) Charles R. Cantor (1978) Christopher T. Walsh (1979) Phillip A. Sharp (1980) Roger D. Kornberg (1981) Harold M. Weintraub (1982) Richard Axel (1983) David V. Goeddel (1984) Gerald M. Rubin (1985) James E. Rothman (1986) Jacqueline K. Barton (1987) Peter Walter (1988) Michael M. Cox (1989) George L. McLendon (1990) Peter G. Schultz (1991) William DeGrado (1992) Stuart L. Schreiber (1993) Peter S. Kim (1994) Jeremy M. Berg (1995) Gregory L. Verdine (1996) Alanna Schepartz (1997) John Kuriyan (1998) Chaitan Khosla (1999) Xiaodong Wang (2000) Jennifer Doudna (2001) Kevan M. Shokat (2002) Andreas Matouschek (2003) Benjamin Cravatt III (2004) Dewey G. McCafferty (2005) Linda Hsieh-Wilson (2006) Anna K. Mapp (2007) Paul J. Hergenrother (2008) Scott K. Silverman (2009) Alice Y. Ting (2010) Nathanael Gray (2011) Christopher J. Chang (2012) Matthew D. Disney (2013) Yi Tang (2014) Minkui Luo (2015) Elizabeth Nolan (2016) Howard Hang (2017) Bradley Pentelute (2018) Neal K. Devaraj (2019) Yimon Aye (2020) Jordan L. Meier (2021)

Eli Lilly Award in Biological Chemistry

v t e History of biology (timeline)
Fields, disciplines	Agricultural science Anatomy Biochemistry Biotechnology Botany Ecology Evolutionary thought Genetics Geology Immunology Medicine Model organisms Molecular biology Molecular evolution Paleontology Phycology Plant systematics RNA biology Zoology (through 1859) Zoology (since 1859)
Theories, concepts	Germ theory of disease Central dogma of molecular biology Darwinism Great chain of being Hierarchy of life Lamarckism One gene–one enzyme hypothesis Protocell RNA world Sequence hypothesis Spontaneous generation
Related	History of science Philosophy of biology Teleology Ethnobotany Eugenics Dysgenics History of the creation-evolution controversy Human Genome Project Humboldtian science Natural history Natural philosophy Natural theology Relationship between religion and science
Category

v t e Time 100: The Most Important People of the 20th Century
Leaders & revolutionaries	David Ben-Gurion Winston Churchill Mahatma Gandhi Mikhail Gorbachev Adolf Hitler Ho Chi Minh Pope John Paul II Ruhollah Khomeini Martin Luther King Jr. Vladimir Lenin Nelson Mandela Mao Zedong Ronald Reagan Eleanor Roosevelt Franklin D. Roosevelt Theodore Roosevelt Margaret Sanger Margaret Thatcher Unknown Tiananmen Square rebel Lech Wałęsa
Artists & entertainers	Louis Armstrong Lucille Ball The Beatles Marlon Brando Coco Chanel Charlie Chaplin Le Corbusier Bob Dylan T. S. Eliot Aretha Franklin Martha Graham Jim Henson James Joyce Pablo Picasso Richard Rodgers and Oscar Hammerstein Bart Simpson Frank Sinatra Steven Spielberg Igor Stravinsky Oprah Winfrey
Builders & titans	Stephen Bechtel Sr. Leo Burnett Willis Carrier Walt Disney Henry Ford Bill Gates Amadeo Giannini Ray Kroc Estée Lauder William Levitt Lucky Luciano Louis B. Mayer Charles E. Merrill Akio Morita Walter Reuther Pete Rozelle David Sarnoff Juan Trippe Sam Walton Thomas J. Watson Jr.
Scientists & thinkers	Leo Baekeland Tim Berners-Lee Rachel Carson Albert Einstein Philo Farnsworth Enrico Fermi Alexander Fleming Sigmund Freud Robert H. Goddard Kurt Gödel Edwin Hubble John Maynard Keynes Leakey family Jean Piaget Jonas Salk William Shockley Alan Turing Francis Crick & James Watson Ludwig Wittgenstein Wright brothers
Heroes & icons	Muhammad Ali The American G.I. Lady Diana Spencer Anne Frank Billy Graham Che Guevara Edmund Hillary & Tenzing Norgay Helen Keller Kennedy family Bruce Lee Charles Lindbergh Harvey Milk Marilyn Monroe Emmeline Pankhurst Rosa Parks Pelé Jackie Robinson Andrei Sakharov Mother Teresa Bill W.

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