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(Redirected from List of ecoregions in Mauritius) Group of islands in the Indian Ocean For the Canadian unincorporated community, see Mascarene, New Brunswick.
Mascarene Islands
Native name: Mascareignes
Topographic map of Mascarene Islands
Geography
LocationIndian Ocean
Coordinates20°12′S 57°45′E / 20.20°S 57.75°E / -20.20; 57.75
Major islandsMauritius, Réunion, Rodrigues
Administration
Mauritius
France
Demographics
Population2,195,087

The Mascarene Islands (English: /mæskəˈriːn/, French: Mascareignes) or Mascarenes or Mascarenhas Archipelago is a group of islands in the Indian Ocean east of Madagascar consisting of islands belonging to the Republic of Mauritius as well as the French department of La Réunion. Their name derives from the Portuguese navigator Pedro Mascarenhas, who first visited them in April 1512. The islands share a common geological origin beneath the Mascarene Plateau known as the Mauritia microcontinent which was a Precambrian microcontinent situated between India and Madagascar until their separation about 70 million years ago. They form a distinct ecoregion with unique biodiversity and endemism of flora and fauna.

Geography

The archipelago comprises three large islands, Mauritius, Réunion, and Rodrigues, plus a number of volcanic remnants in the tropics of the southwestern Indian Ocean, generally between 700 and 1,500 kilometres east of Madagascar. The terrain includes a variety of reefs, atolls, and small islands. They present various topographical and edaphic regions. On the largest islands these gave rise to endemism and unusual biodiversity. The climate is oceanic and tropical.

Mauritius is located 900 km east of Madagascar. It has an area of 1,865 km. The highest point is 828 meters above sea level. Mauritius is the most populous of the Mascarene Islands, with a population of 1,252,964.

Réunion is located 150 km southwest of Mauritius. It is the largest of the islands, with an area of 2512 km. Piton des Neiges (3,069 m), an extinct volcano, is the highest peak on Réunion and in the islands. Piton de la Fournaise is an active volcano on Réunion which erupts frequently.

Rodrigues is located 574 km east of Mauritius. It has an area of 109 km, and reaches 393 meters elevation.

St. Brandon, also known as the Cargados Carajos shoals, is a coral atoll group consisting of a barrier reef, shoals, and low islets. It is the remnant of one -or more- large volcanic islands sitting on the Mauritia (microcontinent) which were submerged by rising tides. Today, the thirty or so islands that comprise St Brandon form part of the Republic of Mauritius. Around seventeen of the uninhabited islands are administered by the 'Outer Islands Development Corporation' of Mauritius (an entity run by the Prime Minister of Mauritius through the Prime Minister's Office) whilst the remaining Thirteen Islands of St. Brandon are managed by the Raphael Fishing Company from three permanently inhabited island bases of Ile Raphael, L'île du Sud (South Island, l'île Boisées) and L'Île Coco. The Raphael Fishing Company was granted these Thirteen Islands of St. Brandon under a 1901 Permanent Lease (999-year lease) which were adjudged as a Permanent Grant by determination of the Privy Council (United Kingdom) in 2008. These Thirteen islands of St. Brandon on Permanent Grant are L'île du Sud (South Island, l'île Boisée), Petit Fou, l'Avocaire, l'île aux Fous, L'île du Gouvernement, Petit Mapou, Grand Mapou, La Baleine, L'Île Coco, Île Raphael, Verronge, l'île aux Bois and Baleines Rocks.

There are several submerged banks or shoals:

Geology

Piton de la Fournaise

The islands are volcanic in origin; Saya de Malha (35 mya) was the first of the Mascarene islands to rise out of the Indian Ocean due to the Réunion hotspot, followed by Nazareth Bank (approximately 2 mya later), Soudan Bank and Cargados Carajos. The youngest islands to form were Mauritius (7–10 mya), the oldest of the existing islands, created along with the undersea Rodrigues ridge. The islands of Rodrigues and Réunion were created in the last two million years. Réunion is the largest of the islands (2,500 km), followed by Mauritius (1,900 km) and Rodrigues (110 km). Eventually, Saya de Malha, Nazareth and Soudan were completely submerged, Cargados Carajos remaining as a coral atoll. The Réunion hotspot was beginning to cool and Rodrigues came out as a small island.

Réunion is home to the highest peaks in the Mascarenes, the shield volcanoes Piton des Neiges (3,069 m) and Piton de la Fournaise (2,525 m). Piton de la Fournaise, on the southeastern corner of Réunion, is one of the most active volcanoes in the world, erupting last in December 2021. Piton de la Petite Rivière Noire (828 m) is the highest peak on Mauritius, and the gentle hills of Rodrigues rise to only 390 m.

The Mascarene Plateau is an undersea plateau that extends approximately 2000 km, from the Seychelles to Réunion. The plateau covers an area of over 115,000 km of shallow water, with depths ranging from 8 to 150 meters, plunging to 4000 m to the abyssal plain at its edges. The southern part of the plateau, including the Saya de Malha Bank, Nazareth Bank, Soudan Banks and Cargados Carajos Shoals (Saint Brandon) (then one large island), was formed by the Réunion hotspot. These were once volcanic islands, much like Mauritius and Réunion, which have now sunk or eroded to below sea level or, in the case of the Cargados Carajos, to low coral islands. The Saya de Malha Bank formed 35 million years ago, and the Nazareth Bank and the Cargados Carajos shoals after that. Limestone banks found on the plateau are the remnants of coral reefs, indicating that the plateau was a succession of islands. Some of the banks may have been islands as recently as 18,000–6,000 years ago, when sea levels were as much as 130 meters lower during the most recent ice age.

History

Pedro Mascarenhas, namesake of the Mascarene Islands, was Viceroy of Portuguese India.
1780 map of Reunion, Mauritius, and Rodrigues
See also: History of Mauritius

The early colonial history of the islands, like that of the Caribbean, featured a succession of takeovers between rival powers: the Portuguese, Dutch, French and British all ruled some or all of the islands.

Around 1507, the explorer Diogo Fernandes Pereira discovered the island group. The area remained under nominal Portuguese rule until Étienne de Flacourt arrived with a French naval squadron and took possession in 1649. From 4 June 1735 to 23 March 1746, a single French Mascarene Islands chartered colony under one gouverneur général (governor general) contained Isle de France (now Mauritius), Île Bourbon (Réunion) and Séchelles (Seychelles). On 14 July 1767 this became a French crown colony, still under one governor general. From 3 February 1803 until 2 September 1810 the French colony of Indes-Orientales, under a capitaine général (captain-general), included Réunion and (nominally) the Seychelles.

Mauritius

VOC - Logo of the Dutch East India Company
Logo of the Dutch East India Company

It is postulated from navigational charts, such as the Cantino planisphere of 1502, that Arab sailors first discovered Mauritius around 975, calling it Dina Arobi (abandoned island). The earliest confirmed discovery on record was in 1507 by Portuguese sailors.

Isle de France (Mauritius)
Isle de France (Mauritius) under the French

The Dutch took physical possession in 1598, establishing a succession of short-lived settlements over a period of about 120 years, before abandoning their efforts in 1710. France took control in 1715, renaming it Isle de France. In 1810, the United Kingdom invaded the island to protect its East India ships coming around the Cape of Good Hope from French pirates. Four years later, at the Treaty of Paris, France ceded Mauritius and its dependencies to the United Kingdom. Under British Mauritius (1810-1968), the island successfully developed as a sugar cane-based plantation economy and colony until independence in 1968.

Rodrigues

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Rodrigues was first discovered by the Arabs but named after Portuguese navigator Diogo Rodrigues. It was under Dutch control in 1601 and settled by the French in 1691. Britain took possession of Rodrigues in 1809. When Mauritius gained independence in 1968, Rodrigues was forcefully joined to it. Rodrigues remains an autonomous region of Mauritius.

Réunion

Réunion was discovered first by the Arabs then by the Portuguese, who named it Santa Apolónia. It was then occupied by the French as part of Mauritius. It was first inhabited by French mutineers who arrived on the island between 1646 and 1669. It was given its current name in 1793. From 1810 to 1815 it was held by the British, before being returned to France. Réunion became an overseas department of France in 1946.

Climate

The climate of the islands is tropical. At low elevations summer temperatures (December to April) average 30 °C. Winter temperatures (May to November) are generally cooler, around 25 °C. Temperatures are cooler in the mountains, averaging 18 °C. During the winter short-lived snow can fall on Réunion's high peaks.

Southeasterly trade winds blow throughout the year. Rainfall is generally higher on the windward sides of the islands. On Mauritius average annual rainfall in the lowlands varies from 1905 mm on the windward southeast coast to 890 mm on the leeward side. Rainfall in the mountains is higher, varying from 2540 mm to 4445 mm annually. Tropical cyclones occur occasionally, bringing heavy rain and high winds, and causing erosion and landslides.

Flora

Foetidia mauritiana
Phyllanthus tenellus (Mascarenes leaf-flower)

The habitats of the Macarenes vary with the islands' size, topography, age, and proximity to Madagascar, the nearest major land mass. As is common among remote islands, the Mascarene fauna and flora display a high degree of endemism; they include over a thousand species of which several hundred are endemic. The Mascarene islands form a distinct ecoregion known as the Mascarene forests. Until Europeans first settled the islands in the sixteenth century the Mascarenes are not known to have harbored any human population, and the islands' wildlife was still flourishing during the early days of settlement. Today much of the natural vegetation is gone, and there are many introduced species on the islands.

In prehistoric times the islands were covered in a diverse range of tropical moist broadleaf forests. The plant communities are not homogeneous, and comprise at least five fairly distinct vegetation zones that reflect variations in altitude and in moisture regime. These include dry lowland forests, semi-dry sclerophyllous forests, lowland rainforests, montane cloud forests, and high-elevation heathlands on Réunion.

Coastal habitats include beach vegetation, coastal wetlands, and swamp forests, grading into lowland rain forest on the windward sides of the islands and lowland dry forests to leeward.

The dry lowland forests are found on the leeward sides of the islands, from sea level to 200 metres in areas with less than 1000 mm of average annual rainfall. Palms are the dominant trees, including species of Latania and Dictyosperma album, along with the palm-like screw-pines (Pandanus spp.).

Semi-dry sclerophyllous forests are found between the coast and 360 metres elevation on Mauritius and Rodrigues, and from 200 to 750 metres elevation on the western slopes of Réunion, where only small patches of such forest now remain. Average annual rainfall in the semi-dry forests ranges from 1000 to 1500 mm. Tree genera and families well-represented in the semi-dry forests include ebony species in the genus Diospyros (Ebenaceae), Pleurostylia (Celastraceae), Foetidia (Lecythidaceae), Olea europaea subsp. cuspidata (Oleaceae), Cossinia pinnata (Sapindaceae) Dombeya (Malvaceae), Terminalia bentzoe (Combretaceae), and Sideroxylon boutonianum, Sideroxylon borbonicum, and various species of Mimusops in family Sapotaceae. Shrubs in the semi-dry forests include several spectacular endemic species of Hibiscus (Malvaceae), Zanthoxylum (Rutaceae), Obetia ficifolia (Urticaceae), and Scolopia heterophylla (Flacourtiaceae).

The lowland rain forests are characterized by dense evergreen forests, composed of diverse species creating a canopy of 30 meters or more in height. They are found on the Mauritius' eastern lowlands from the coast to 800–900 metres elevation, and above 360 m on the western slopes. On Réunion lowland rain forests are found between 750 and 1100 m. They occur in areas with average annual rainfall of 1500 to 6000 mm. Characteristic trees include species of Mimusops and Labourdonnaisia (Sapotaceae), Hernandia mascarenensis (Hernandiaceae), Calophyllum (Clusiaceae), and species of Syzygium, Eugenia, Sideroxylon, and Monimiastrum (Myrtaceae). Characteristic shrubs include species of Gaertnera, Chassalia, Bertiera, and Coffea in family Rubiaceae. Other plants include bamboos like Nastus borbonicus, numerous species of orchids (e.g., Angraecum, Bulbophyllum) and ferns e.g., Asplenium, Hymenophyllum, Trichomanes, Elaphoglossum, and Marattia fraxinea.

The cloud forests are evergreen rain forests found on mountain slopes with high rainfall. They occur on Réunion between 800 and 1900 metres elevation on eastern slopes with an average annual rainfall of 2000–10,000 mm, and between 1100 and 2000 metres on western slopes with an average annual rainfall of 2000–3000 mm. By 2005, relatively intact cloud forests still covered approximately 44,000 ha on Réunion. On Mauritius they are restricted to a small area of Montagne Cocotte in the island's southwest, above 750 metres elevation and with average annual rainfall of 4500–5500 mm. Trees form a dense canopy 6 to 10 metres high. Typical canopy trees include species of Dombeya on Réunion, and species of Monimia and Tambourissa (Monimiaceae) on both islands. There is an understory of small trees and shrubs including species of Psiadia (Asteraceae) and Melicope (Rutaceae). The forests are rich in epiphytes (orchids, ferns, mosses, lichens), emergent tree ferns (Cyathea species), and, originally the palms Acanthophoenix rubra, but poaching has wiped out palms in many areas of Réunion. Three monodominant plant communities are also found in the cloud forests – forests of Acacia heterophylla (Fabaceae) as the canopy tree, which are similar to the Acacia koa forests in Hawaii, thickets dominated by Erica reunionensis (Ericaceae), and hyperhumid forests of screw-pine (Pandanus montanus).

Open subalpine shrubland on Piton de la Fournaise, with Erica reunionensis, Stoebe passerinoides, Acacia heterophylla, and Erica galioides

Subalpine shrub occurs on the volcanoes on Réunion, above the treeline at 1800 to 2000 metres elevation. Average annual rainfall is from 2000 to 6000 mm, and frosts occur regularly in winter. Characteristic shrubs include Erica reunionensis and Erica galioides (Ericaceae), Stoebe passerinoides and species of Hubertia and Psiadia (Asteraceae), and Phylica nitida (Rhamnaceae). Herbaceous endemic species include Heterochaenia rivalsii (Campanulaceae), Eriotrix commersonii (Asteraceae), and Cynoglossum borbonicum (Boraginaceae).

Recent volcanic deposits on Réunion's volcanic summits are covered by sparse grasslands rich in endemic grasses, including Festuca borbonica, Agrostis salaziensis, and Cenchrus afer. along with orchids like Disa borbonica. Ericoid thickets and thickets of the small tree Sophora denudata (Fabaceae) are found on weathered volcanic substrates.

Most of the indigenous Mascarene flora and fauna are thought to have descended originally from Madagascan and African ancestors.

The four largest families present in the Mascarene Islands are Myrtaceae, Rubiaceae, Orchidaceae, and Euphorbiaceae, which comprise between 193 and 223 species each, for a total of 831 species, or 26.9% of the flowering plant flora. Another seven families contain 80 or more species each: Poaceae, Apocynaceae, Cyperaceae, Cunoniaceae, Rutaceae, Araliaceae, and Sapotaceae, representing an additional ca. 660 species in all, 21.3% of the angiosperm flora. Ferns are prominent components of the biotas of the islands, especially in the tropical forest. Most ferns disperse easily via ornithochory of their spores, allowing fairly frequent colonization from Madagascar and exchange among the Mascarene islands.

The Mascarenes are home to many endemic species of Dombeyoideae, the monotypic genus Psiloxylon (Psiloxylon mauritianum), and members of the family Monimiaceae (the genera Monimia and Trochetia, and species of Tambourissa) and Escalloniaceae. Indigenous trees include species of the genera Ocotea, Erythrina, Sideroxylon, and Foetidia. The palm genera Latania, Hyophorbe, Acanthophoenix, Dictyosperma, and Tectiphiala are all endemic to the Mascarene Islands. Other endemic genera include Berenice, Heterochaenia, Nesocodon, Ruizia, and Astiria.

There are 24 species of trees and shrubs in the Mascarenes from subfamily Dombeyoideae, 23 of which are endemic to the islands. The Macarenes' endemic species are polyphyletic and split into nine clades. Trochetia appears monophyletic and more closely related to Eriolaena and Helmiopsis than to Dombeya. All Dombeya taxa are included in a clade together with Ruizia and Astiria, this means that Dombeya is paraphyletic. In terms of breeding systems the Malagasy Dombeyoideae are hermaphroditic, whereas those of the Mascarenes are considered dioecious. The polyphyly of the Mascarene Dombeyoideae suggests that dioecy has been acquired several times. At least five colonization events from Madagascar to the Mascarene archipelago occurred. The evolutionary history of two lineages of Mascarene Domeyoideae seems to be related to adaption to xeric habitats.

The flora of the islands co-evolved with the island's unique fauna over millions of years. The human-caused extinction of several of the Mascarenes' land animals, and the introduction of exotic animals to the islands by humans, has disrupted the reproduction and thriving of various island plants. For example, the Tambalacoque tree (Sideroxylon grandiflorum), often called the dodo tree, is not producing young trees and is threatened with extinction. The extinct dodo, Cylindraspis tortoises, and broad-billed parrots, along with fruit bats, played an important role in dispersing the seeds. Passing through the animals' digestive tract aided in the seeds' germination. However the trees' decline may be mostly due to introduced crab-eating macaques (Macaca fascicularis) destroying unripened seeds.

Fauna

The dodo was a frugivore giant columbiforme of Mauritius Island where there are several Ocotea species. Painted by Jacob Hoefnagel, c. 1602

Much of the Mascarenes' native fauna has become endangered or extinct since the human settlement of the islands in the 17th century. Settlers cleared most of the forests for agriculture and grazing, and introduced many exotic species, including pigs, rats, cats, monkeys, and mongooses.

Birds

Many of the Mascarene birds evolved into flightless forms, the most famous of which was the extinct flightless pigeon, the dodo of Mauritius. Thirteen additional species of birds became extinct, including the Réunion flightless ibis, broad-billed parrot, red rail, Rodrigues rail, and Rodrigues solitaire.

Contemporary sources state that the dodo used gizzard stones and could swallow Ocotea fruits. Seed distribution of at least some Ocotea species is performed by frugivorous birds and there exist a few reports of "solitaires" from the Mascarenes without mention of which island these came from, and the term was also used for other species with "solitary" habits, such as the Réunion blue swamphen and the Réunion sacred ibis. At one point it was even believed that Réunion was the home of not only a white dodo, but also a white solitaire. In 1786, sub-fossil bones were discovered in a cave which confirmed Leguat's descriptions, but at this time no living residents of Rodrigues remembered having seen living birds. The star constellation Turdus Solitarius was named after this bird.

Today 17 endemic bird species survive on the islands. Two species – the Mascarene paradise flycatcher (Terpsiphone bourbonnensis) and Mascarene swiftlet (Aerodramus francicus) – inhabit both Mauritius and Réunion. Eight species are endemic to Mauritius – the Mauritius grey white-eye, (Zosterops mauritianus), Mauritius cuckooshrike (Lalage typica), Mauritius kestrel (Falco punctatus), Mauritius fody (Foudia rubra), Mauritius bulbul (Hypsipetes olivaceus), Mauritius parakeet (Psittacula eques), Mauritius olive white-eye (Zosterops chloronothos), and pink pigeon (Nesoenas mayeri). The Réunion grey white-eye (Zosterops borbonicus), Réunion cuckooshrike (Lalage newtoni), Réunion stonechat (Saxicola tectes), Réunion olive white-eye (Zosterops olivaceus), and Réunion bulbul (Hypsipetes borbonicus) are endemic to Réunion. The Rodrigues warbler (Acrocephalus rodericanus) and Rodrigues fody (Foudia flavicans) are found only on Rodrigues.

Reptiles

Mauritius, Réunion, and Rodrigues were also once each home to one or more species of giant tortoises, now extinct, which comprised the genus Cylindraspis. There are 13 living endemic reptile species, including a number of species of day geckoes (genus Phelsuma).

Mammals

The islands' only native mammals are bats, including the Mauritian flying fox (Pteropus niger), which is endemic to Mauritius and Réunion, and the Rodrigues flying fox (Pteropus rodricensis) found only on Rodrigues. The lesser Mascarene flying fox (Pteropus subniger) is extinct.

Freshwater

The three largest islands have many freshwater streams which descend from the mountains to the sea. The upper reaches, particularly on Réunion, are fast-flowing and steep with numerous waterfalls. Tamarin Falls or Sept Cascades (293 m) on Mauritius is the highest waterfall on Mauritius, while Cascade Trou de Fer (725 m, ranks 23rd worldwide) on Réunion is the highest of the Mascarenes.

The Grand River South East (34 km) is the longest river on Mauritius, followed by Rivière du Poste (23 km), Grand River North West (22 km), Rivière La Chaux (22 km), and Rivière des Créoles (20 km). Mauritius has two natural crater lakes, Grand Bassin and Bassin Blanc.

There are about twenty species of fish which spend at least part of their lives in freshwater. The islands have five endemic freshwater fish species, all gobies (Gobiiformes) – Hypseleotris cyprinoides, Cotylopus acutipinnis, Glossogobius kokius, Gobius commersonii, and Oxyurichthys guibei. Cotylopus acutipinnis, which is endemic to Réunion, hatches in the sea before migrating to the rainforest streams where it lives its adult life.

Marine

The Mascarene Islands are surrounded by approximately 750 square km of coral reef. Rodrigues has nearly continuous fringing reefs bounding an extensive lagoon, 7 to 20 km wide, with deep channels. Mauritius is also surrounded by a fringing reef. In contrast, Réunion has very short stretches of narrow fringing reef along the western and southwestern coasts only. The islets of the Cargados Carajos Shoals, which have a very depauperate terrestrial biota owing to being so low-lying and swamped during cyclones, are bound to the east by an extensive arc of fringing reef, which accounts for ~30% of the reefs of the Mascarene Islands. Lagoon reefs and reef flats are dominated by scleractinian corals such as branching and tabular Acropora, Porites massives, foliaceous Montipora and Pavona, and sand consolidated with beds of seagrass such Halophila (Hydrocharitaceae). Among coral reef fishes, wrasses (Labridae), damselfish (Pomacentridae), carnivorous groupers (Serranidae), and surgeonfishes (Acanthuridae) have many species.

Wildlife, threats, and preservation

The Saint Brandon Conservation Trust at Corporate Council on Africa in May 2024 in Dallas, Texas

Mauritius has one of the highest population densities in the world. All of the islands have experienced significant losses of habitats. Many of the surviving endemic species are still threatened with extinction and have little protection. Less than 40 percent of Réunion is covered with natural vegetation, only about 5 percent of Mauritius, and almost none of Rodrigues. On Réunion, forest has been cleared for agriculture and then overtaken by introduced plants. Mauritius was largely converted to sugar cane, tea, and conifer plantations. On Rodrigues, shifting cultivation has caused the damage.

Launch of Saint Brandon Conservation Trust

On 8 May 2024, Saint Brandon Conservation Trust, an independent Mauritian NGO, was internationally launched at the Corporate Council on Africa in Dallas, Texas, to protect St. Brandon from pollution and to protect surviving flora and fauna in the Cargados Carajos.

Protected areas

Protected areas constitute 40.6% of the Mascarene Islands' land area. Terrestrial protected areas on Mauritius include Black River Gorges National Park, Bras d'Eau National Park, Perrier Nature Reserve, Corps de Garde Nature Reserve, Le Pouce Nature Reserve, Cabinet Nature Reserve, Gouly Pere Nature Reserve, and Bois Sec Nature Reserve. There are 32 designated protected areas on Réunion, including Réunion National Park (1055.15 km), which cover 63.13% of the island.

See also

References

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Sources

  • Quammen, David, (1996) The Song of the Dodo. Touchstone, New York.
  • Diamond, Jared, (1984) "Historic extinctions: A rosetta stone for understanding prehistoric extinctions". In: P. Martin and R. Klein (eds.) (1984) Quaternary Extinctions: A prehistoric revolution. University of Arizona Press, Tucson.

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French overseas empire
Former
Former French colonies in Africa and the Indian Ocean
North Africa
West Africa
Equatorial Africa
Comoros
Former French colonies in the Americas
French North America
French Caribbean
Equinoctial France
Former French colonies in Asia and Oceania
French India
Indochinese Union
Mandate for Syria
and the Lebanon
Oceania
Present
Overseas France
Inhabited territories
Overseas regions
Overseas collectivities
Sui generis collectivity
Uninhabited territories
North Pacific Ocean
Overseas territory (French Southern and Antarctic Lands)
Scattered Islands in the Indian Ocean
Portuguese Empire
North Africa

15th century

1415–1640 Ceuta
1458–1550 Alcácer Ceguer (El Qsar es Seghir)
1471–1550 Arzila (Asilah)
1471–1662 Tangier
1485–1550 Mazagan (El Jadida)
1487–16th century Ouadane
1488–1541 Safim (Safi)
1489 Graciosa

16th century

1505–1541 Santa Cruz do Cabo de Gué (Agadir)
1506–1525 Mogador (Essaouira)
1506–1525 Aguz (Souira Guedima)
1506–1769 Mazagan (El Jadida)
1513–1541 Azamor (Azemmour)
1515–1541 São João da Mamora (Mehdya)
1577–1589 Arzila (Asilah)

Anachronous map of the Portuguese Empire (1415-1999)
Sub-Saharan Africa

15th century

1455–1633 Arguim
1462–1975 Cape Verde
1470–1975 São Tomé
1471–1975 Príncipe
1474–1778 Annobón
1478–1778 Fernando Poo (Bioko)
1482–1637 Elmina (São Jorge da Mina)
1482–1642 Portuguese Gold Coast
1498–1540 Mascarene Islands

16th century

1500–1630 Malindi
1501–1975 Portuguese Mozambique
1502–1659 Saint Helena
1503–1698 Zanzibar
1505–1512 Quíloa (Kilwa)
1506–1511 Socotra
1508–1547 Madagascar
1557–1578 Accra
1575–1975 Portuguese Angola
1588–1974 Cacheu
1593–1698 Mombassa (Mombasa)

17th century

1645–1888 Ziguinchor
1680–1961 São João Baptista de Ajudá, Benin
1687–1974 Bissau

18th century

1728–1729 Mombassa (Mombasa)
1753–1975 Portuguese São Tomé and Príncipe

19th century

1879–1974 Portuguese Guinea
1885–1974 Portuguese Congo

Middle East

16th century

1506–1615 Gamru (Bandar Abbas)
1507–1643 Sohar
1515–1622 Hormuz (Ormus)
1515–1648 Quriyat
1515–? Qalhat
1515–1650 Muscat
1515?–? Barka
1515–1633? Julfar (Ras al-Khaimah)
1521–1602 Bahrain (Muharraq • Manama)
1521–1529? Qatif
1521?–1551? Tarut Island
1550–1551 Qatif
1588–1648 Matrah

17th century

1620–? Khor Fakkan
1621?–? As Sib
1621–1622 Qeshm
1623–? Khasab
1623–? Libedia
1624–? Kalba
1624–? Madha
1624–1648 Dibba Al-Hisn
1624?–? Bandar-e Kong

South Asia

15th century

1498–1545 Laccadive Islands
(Lakshadweep)

16th century
Portuguese India

 • 1500–1663 Cochim (Kochi)
 • 1501–1663 Cannanore (Kannur)
 • 1502–1658
 1659–1661
Quilon
(Coulão / Kollam)
 • 1502–1661 Pallipuram (Cochin de Cima)
 • 1507–1657 Negapatam (Nagapatnam)
 • 1510–1961 Goa
 • 1512–1525
 1750
Calicut
(Kozhikode)
 • 1518–1619 Portuguese Paliacate outpost (Pulicat)
 • 1521–1740 Chaul
  (Portuguese India)
 • 1523–1662 Mylapore
 • 1528–1666 Chittagong
(Porto Grande De Bengala)
 • 1531–1571 Chaul
 • 1531–1571 Chalé
 • 1534–1601 Salsette Island
 • 1534–1661 Bombay (Mumbai)
 • 1535 Ponnani
 • 1535–1739 Baçaím (Vasai-Virar)
 • 1536–1662 Cranganore (Kodungallur)
 • 1540–1612 Surat
 • 1548–1658 Tuticorin (Thoothukudi)
 • 1559–1961 Daman and Diu
 • 1568–1659 Mangalore
  (Portuguese India)
 • 1579–1632Hugli
 • 1598–1610Masulipatnam (Machilipatnam)
1518–1521 Maldives
1518–1658 Portuguese Ceylon (Sri Lanka)
1558–1573 Maldives

17th century
Portuguese India

 • 1687–1749 Mylapore

18th century
Portuguese India

 • 1779–1954 Dadra and Nagar Haveli

East Asia and Oceania

16th century

1511–1641 Portuguese Malacca
1512–1621 Maluku
 • 1522–1575  Ternate
 • 1576–1605  Ambon
 • 1578–1650  Tidore
1512–1665 Makassar
1515–1859 Larantuka
1557–1999 Macau
1580–1586 Nagasaki

17th century

1642–1975 Portuguese Timor (East Timor)

19th century
Portuguese Macau

 • 1864–1999 Coloane
 • 1851–1999 Taipa
 • 1890–1999 Ilha Verde

20th century
Portuguese Macau

 • 1938–1941 Lapa and Montanha (Hengqin)

  • 1975 is the year of East Timor's Declaration of Independence and subsequent invasion by Indonesia. In 2002, East Timor's independence was fully recognized.
North America and North Atlantic

15th century

1420 Madeira
1432 Azores

16th century

1500–1579? Terra Nova (Newfoundland)
1500–1579? Labrador
1516–1579? Nova Scotia

South America and Caribbean

16th century

1500–1822 Brazil
 • 1534–1549  Captaincy Colonies of Brazil
 • 1549–1572  Brazil
 • 1572–1578  Bahia
 • 1572–1578  Rio de Janeiro
 • 1578–1607  Brazil
 • 1621–1815  Brazil
1536–1620 Barbados

17th century

1621–1751 Maranhão
1680–1777 Nova Colónia do Sacramento

18th century

1751–1772 Grão-Pará and Maranhão
1772–1775 Grão-Pará and Rio Negro
1772–1775 Maranhão and Piauí

19th century

1808–1822 Cisplatina (Uruguay)
1809–1817 Portuguese Guiana (Amapá)
1822 Upper Peru (Bolivia)

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