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List of pelycosaurs

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This list of pelycosaurs is an attempt to create a comprehensive listing of all genera that have ever been included in the synapsida excluding therapsida and purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomina dubia), or were not formally published (nomina nuda), as well as junior synonyms of more established names, and genera that are no longer considered pelycosaurs.

The list currently contains 83 generic names.

Naming conventions and terminology

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

  • Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
  • Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
  • Nomen oblitum (Latin for "forgotten name"): A name that has not been used in the scientific community for more than fifty years after its original proposal.
  • Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
  • Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

List

Genus Authors Year Status Age Location Notes

Aerosaurus

Romer

1937

Valid.

Late Carboniferous - Lower Permian

United States of America

Angelosaurus

Olson
Beerbower

1953

Valid.

Upper Permian

United States of America

Anglosaurus

Bruner

1991

Sic

N/A

N/A

Incorrect spelling for Angelosaurus

Apsisaurus

Laurin

1991

Valid.

Lower Permian

United States of America

Formerly assigned as a "eosuchian" diapsid.

Archaeobelus

Cope

1877

Syn.

N/A

N/A

Synonym of Clepsydrops

Archaeothyris

Reisz

1972

Valid.

Upper Carboniferous

United States of America

Archaeovenator

Reisz
Dilkes

2003

Valid.

Upper Carboniferous

United States of America

Arribasaurus

Williston

1914

Syn.

N/A

N/A

Synonym of Ophiacodon

Baldwinonus

Romer
Price

1940

Valid.

Late Carboniferous - Early Permian

United States of America

Basicranodon

Vaughn

1958

Syn.

Lower Permian

United States of America

Synonym of Mycterosaurus

Bathyglyptus

Case

1911

Syn.

N/A

N/A

Synonym of Dimetrodon

Bathygnathus

Leidy

1854

Valid.

Lower Permian

United States of America

Brachycnemius

Williston

1911

Syn.

N/A

N/A

Synonym of Edaphosaurus

Callibrachion

Boule
Glangeau

1893

Syn.

N/A

N/A

Synonym of Haptodus

Casea

Williston

1910

Valid.

Lower Permian

France and United States of America

Caseoides

Olson
Beerbower

1953

Valid.

Lower Permian

United States of America

Caseopsis

Olson

1962

Valid.

Upper Permian

United States of America

Clepsydrops

Cope

1875

Valid.

Late Carboniferous

United States of America

Colobomycter

Vaughn

1958

Valid.

Lower Permian

United States of America

It is a protorothyridid eureptile

Cotylorhynchus

Stovall

1937

Valid.

Lower Permian

United States of America

Cryptovenator

Fröbisch
Schoch
Müller
Schindler
Schweiss

2011

Valid.

Late Carboniferous

Germany

Ctenorhachis

Hook
Hotton

1991

Valid.

Upper Permian

United States of America

Ctenospondylus

Romer

1936

Valid.

Lower Permian

United States

Cutleria

Lewis
Vaughn

1965

Valid.

Early Permian

United States of America

Delorhynchus

Fox

1962

Valid.

Lower Permian

United States of America

Originally descript as a pelycosaur, but probably a parareptile

Datheosaurus

Schroeder

1905

Syn.

N/A

N/A

Synonym of Haptodus

Dimacrodon

Olson
Beerbower

1953

Valid?

Lower Permian

United States

Originally descript as an anomodont therapsid by Olson & Beerbower (1953) and Olson (1962), it lacks any diagnostic features of anomodonts and rather represent pelycosaur-grade synapsids

Dimetrodon

Cope

1878

Valid.

Early Permian - Middle Permian

United States of America and Germany

Diopaeus

Cope

1892

Syn.

N/A

N/A

Synonym of Ophiacodon

Echinerpeton

Reisz

1972

Valid.

Upper Carboniferous

Canada

Edaphosaurus

Cope

1882

Valid.

Late Carboniferous - Lower Permian

Czech Republic and United States of America

Elcabrosaurus

Case

1907

Syn.

N/A

N/A

Synonym of Sphenacodon

Elliotsmithia

Broom

1937

Valid.

Middle-Upper Permian

South Africa

Embolophorus

Cope

1878

Syn.

N/A

N/A

Synonym of Dimetrodon

Ennatosaurus

Efremov

1956

Valid.

Lower Permian

Russia

Eothyris

Romer

1937

Valid.

Lower Permian

United States of America

Eumatthevia

Broom

1930

Syn.

N/A

N/A

Synonym of Mycterosaurus

Euromycter

Reisz
Maddin
Fröbisch
Falconnet

2011

Valid.

Early-Late Permian

France

Glaucosaurus

Williston

1915

Valid.

Lower Permian

United States of America

Haptodus

Gaudry

1866

Valid.

Late Carboniferous - Lower Permian

United States of America and Europe

Heleosaurus

Broom

1907

Valid.

Upper Permian

South Africa

Ianthasaurus

Reisz
Berman

1986

Valid.

Late Carboniferous

United States of America

Ianthodon

Kissel
Reisz

2004

Valid.

Upper Carboniferous

United States of America

Ianthosaurus

Reisz
Berman

1986

Sic

N/A

N/A

Incorrect spelling of Ianthasaurus

Lupeosaurus

Romer

1937

Valid.

Lower Permian

United States of America

Macromerion

Fritsch

1879

Valid.

Upper Carboniferous

Germany

Mesenosaurus

Efremov

1938

Valid.

Lower Permian

Russia

Mesonosaurus

Ivakhnenko
Kurzanov

1979

Sic

N/A

N/A

Incorrect spelling of Mesenosaurus

Milosaurus

DeMar

1970

Valid.

Upper Carboniferous

United States of America

Mycterosaurus

Williston

1915

Valid.

Lower Permian

United States of America

Naosaurus

Cope

1886

Syn.

N/A

N/A

Synonym of Edaphosaurus

Neosaurus

Nopsca

1923

Valid.

Lower Permian

France

Nitosaurus

Romer

1937

Valid.

Lower Permian

United States of America

Oedaleops

Langston

1965

Valid.

Lower Permian

United States of America

Ophiacodon

Marsh

1878

Valid.

Late Carboniferous-Lower Permian

United States of America and England

Oromycter

Reisz

2005

Valid.

Lower Permian

United States of America

Oxyodon

Huene

1908

Preocc.

N/A

N/A

Preoccupied by fish genus Oxyodon Brauer, 1906. Synonym of Sphenacodon

Palaeohatteria

Credner

1888

Valid.

Early Permian

Germany

Palaeosphenodon

Baur

1889

Syn.

N/A

N/A

Synonym of Palaeohatteria

Pantelosaurus

von Huene

1925

Valid.

Early Permian

Germany

Phreatophasma

Efremov

1954

Valid.

Middle Permian

Russia

Poecilospondylus

Case

1910

Syn.

N/A

N/A

Synonym of Varanosaurus

Poliosaurus

Case

1907

Syn.

N/A

N/A

Synonym of Ophiacodon

Protoclepsydrops

Carroll

1964

Valid.

Upper Carboniferous

Canada

Pyozia

Anderson
Reisz

2004

Valid.

Middle Permian

Russia

Ruthenosaurus

Reisz
Maddin
Fröbisch
Falconnet

2011

Valid.

Early-Late Permian

France

Ruthiromia

Eberth
Brinkman

1983

Valid.

Lower Permian

United States of America

Scoliomus

Williston
Case

1915

Syn.

Lower Permian

United States of America

Junior synonym of Sphenacodon

Secodontosaurus

Romer

1936

Valid.

Lower Permian

United States of America

Sphenacodon

Marsh

1878

Valid.

Late Carboniferous-Early Permian

United States of America and England

Stereophallodon

Romer

1937

Valid.

Late Carboniferous-Early Permian

United States of America

Stereorhachis

Gaudry

1880

Valid.

Late Carboniferous

France

Theropleura

Cope

1878

Syn.

N/A

N/A

Synonym of Ophiacodon (in partium) and Dimetrodon (in partium)

Therosaurus

von Huene

1925

Syn.

N/A

N/A

Synonym of Ophiacodon

Thrausmosaurus

Fox

1962

Valid.

Lower Permian

United States of America

The type species, T. serratidens, was declared to be a nomen dubium

Trichasaurus

Williston

1913

Valid.

Lower Permian

United States of America

Replacement name for Trispondylus Williston, 1910

Trispondylus

Williston

1910

Preocc.

N/A

N/A

Preoccupied by phenacodontid Trispondylus Cope, 1884

Varanodon

Olson

1965

Valid.

Lower Permian

United States of America

Varanops

Williston

1911

Valid.

Lower Permian

United States of America

Varanosaurus

Broili

1904

Valid.

Lower Permian

United States of America

Xyrospondylus

Reisz
Heaton
Pynn

1982

Valid.

Upper Carboniferous

United States of America

Watongia

Olson

1974

Valid.

Middle Permian

United States of America

Winfeldia

Romer

1925

Syn.

N/A

N/A

Synonym of Ophiacodon

Archaeovenator
Casea
Cotylorhynchus
Ctenospondylus
Dimetrodon
Edaphosaurus
Ennatosaurus
Haptodus
Ianthasaurus
Mycterosaurus
Oedaleops
Ophiacodon
Secodontosaurus
Sphenacodon
Varanodon

See also

References

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  2. ^ Romer, A. S. (1937). "New genera and species of pelycosaurian reptiles". Proceedings of the New England Zoological Club. XVI: 90–96.
  3. ^ Olson, E. C. & Barghusen, H. (1962). "Permian vertebrates from Oklahoma and Texas". Oklahoma Geological Survey. 59: 1–20.
  4. ^ Bruner, J. C. (1991). "A catalogue of Type Specimens of Fossil Vertebrates in the Field Museum of Natural History. Classes Amphibia, Reptilia, Aves and ichnites". Geology. 22: 1–68. doi:10.5962/bhl.title.3392.
  5. ^ Reisz, R.R.; Laurin, M. & Marjanovic, D. (2010). "Apsisaurs witteri from the lower Permian of Texas: yet another small varanopid synapsid, not a diapsid". Journal of Vertebrate Paleontology. 30 (5): 1628–1631. doi:10.1080/02724634.2010.501441. S2CID 129835335.
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  10. Reisz, R. R.; Wilson, H. & Scott, D. (1997). "Varanopseid synapsid skeletal elements from Richards Spur, a Lower Permian fissure fill near Fort Sill". Oklahoma Geology Notes. 57: 160–170.
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