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Litokoala Temporal range: early-Middle Miocene
Artist's reconstruction
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Infraclass:	Marsupialia
Order:	Diprotodontia
Family:	Phascolarctidae
Genus:	†Litokoala Stirton, Tedford & Woodburne, 1967.
Species
L. dicktedfordi Pledge, 2010 L. garyjohnstoni Louys et al., 2007 L. kutjamarpensis (type species) Stirton et al., 1967 L. kanunkaensis Springer, 1987 L. thurmerae Pledge, 2010 L. dicksmithi Black, Louys & Price 2013

Litokoala is an extinct genus of marsupials, and along with Nimiokoala, is closely related to the modern koala. The three genera may have diverged at an earlier date, although the drying of the continent and the expansion of Eucalyptus forests towards the late Miocene may have delayed the evolution of cranial features unique to the modern genera. This indicates that either fossil genus could be an ancestor of the modern genus, or the modern genus has a common ancestor to both. More material needs collection to improve their taxonomical relationships.

The genus lived about 10–16 million years ago in the middle Miocene Riversleigh of Queensland. This area is described as a rainforest habitat at time of sediment deposition. It had a different diet to the modern species, with the dental symphysis unfused, indicating a diet that was properly varied in nature, unlike the specialised nature of Phascolarctos. The size is estimated to be only half of the modern genus. Cranial adaptations are intermediate between the extant common brushtail possum and koala, with minor divergence from either.

This genus and Nimiokoala are similar in most anatomical features so far as is known, except Litokoala possessed a superficial messateric process, while Nimiokoala had "more marked basiooccipital-basisphenoid flexion and a more extensive posterior attachment of the pterygoid", which make these features basal in their taxonomical position in relation to the Phascolarctos. The basiocranial (back of skull) features are similar to Phascolarctos, while anterior (facial) features exhibit similarities with the genus Trichosurus. Only partial fragments are known, with only the posterior section of the zygomatic process known from the L. kutjamarpensis skull.

References

1. ^ Stirton, R. A.; Tedford, Richard H.; Woodburne, Michael O. (1967). "A new Tertiary formation and fauna from the Tirari Desert, South Australia". Records of the South Australian Museum. 15 (3): 427–462.
2. ^ N. S. Pledge. 2010. A new koala (Marsupialia : Phascolarctidae) from the late Oligocene Etadunna Formation, Lake Eyre Basin, South Australia. Australian Mammalogy 32:79-86.
3. Louys, J., K. Black, M. Archer, S. J. Hand, and H. Godthelp. 2007. Descriptions of koala fossils from the Miocene of Riversleigh, northwestern Queensland and implications for Litokoala. Alcheringa 31:99–110.
4. Karen H. Black, Julien Louys & Gilbert J. Price (2013). «Understanding morphological variation in the extant koala as a framework for identification of species boundaries in extinct koalas (Phascolarctidae; Marsupialia)». Journal of Systematic Palaeontology (published 14 May 2013) doi:10.1080/14772019.2013.768304
5. (Louys et al., 2009, p. 989)

v t e Vombatiformes
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Diprotodontia
†Thylacoleonidae Enigmaleo Lekaneleo Microleo Thylacoleo Wakaleo Phascolarctidae †Invictokoala †Koobor †Litokoala †Madakoala †Nimiokoala †Perikoala Phascolarctos †Priscakoala Vombatomorphia †Thylacoleonidae? †Ilariidae Ilaria Kuterintja †Wynyardiidae Muramura Namilamadeta Wynyardia Vombatoidea †Marada? †Mukupirna Vombatidae Lasiorhinus †Nimbavombatus †Phascolonus †Ramsayia †Rhizophascolonus †Sedophascolomys Vombatus †Warendja †Diprotodontoidea Palorchestidae Palorchestes Propalorchestes Diprotodontidae Alkwertatherium Ambulator Diprotodon Euowenia Euryzygoma Hulitherium Kolopsis Kolopsoides Maokopia Meniscolophus Neohelos Ngapakaldia Nimbadon Nototherium Pitikantia Plaisiodon Pyramios Raemeotherium Silvabestius Sthenomerus Zygomaturus

• Miocene mammals of Australia
• Prehistoric vombatiforms
• Miocene marsupials
• Koalas
• Prehistoric marsupial genera
• Riversleigh fauna
• Fossil taxa described in 1967