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Tyrannosaurus is one of the most iconic dinosaurs and is known from numerous specimens, some of which have individually acquired notability due to their scientific significance and media coverage.

Specimen data

Specimen number	Name	Completeness	Discovery	Museum	Museum city	Discoverer	Formation	Location	Notes
AMNH 3982	Manospondylus	< 1%	1892	American Museum of Natural History	New York City	Edward Cope	Hell Creek Formation	Faith, South Dakota	Described as Manospondylus gigas
BMNH R7994	Dynamosaurus	13%	1900	Natural History Museum, London	London	Barnum Brown	Lance Formation	Seven Mile Creek, Wyoming	Originally AMNH 5866 Described as Dynamosaurus imperiosus, analysis of the specimen suggests a large individual similar to Sue
CM 9380	Holotype	10%	1902	Carnegie Museum of Natural History	Pittsburgh	Barnum Brown Richard Lull	Hell Creek Formation	Montana	Originally AMNH 973
AMNH 5027		45%	1908	American Museum of Natural History	New York City	Barnum Brown	Hell Creek Formation	Montana	The base skeleton that was used as the iconic symbol for the famous logo of the Jurassic Park film series.
CMNH 7541	Nanotyrannus		1942	Cleveland Museum of Natural History	Cleveland	David Dunkle	Hell Creek Formation	Montana	Described as Gorgosaurus lancensis and later given its own genus, Nanotyrannus. Now considered a juvenile specimen of Tyrannosaurus rex by most researchers.
LACM 23844		25%	1966	Natural History Museum of Los Angeles County	Los Angeles	Harley Garbani	Hell Creek Formation	Montana
RTMP 1981.6.1	"Black Beauty"	28%	1980	Royal Tyrrell Museum of Palaeontology	Drumheller	Jeff Baker	Willow Creek Formation	Crowsnest Pass, Alberta	One of the westernmost specimens, found at ${\displaystyle 114^{\circ }}$ W longitude. Noted for black colouration taken on during fossilisation.
Private	"Stan"	65%	1987		Hill City	Stan Sacrison	Hell Creek Formation	Buffalo, South Dakota	One of the best-known specimens, with casts present in numerous institutions. Sold to a private owner in 2020 for US$31 million, formerly in the Black Hills Institute as BHI 3033.
MOR 555	"Wankel"	80-85%	1988	Museum of the Rockies	Bozeman	Kathy Wankel	Hell Creek Formation	Montana	Also known as The Nation's Rex. Specimen on loan to the Smithsonian Institution, with a cast on display at the Museum of the Rockies.
MOR 1125	"B-rex/Bob"	37%	2000	Museum of the Rockies	Bozeman	Bob Harmon	Hell Creek Formation	Montana	Confirmed as female, due to the presence of medullary bone.
MOR 1126	"C-rex/Celeste"	9%	2000	Museum of the Rockies	Bozeman	Celeste Horner	Hell Creek Formation	Montana	One of the largest specimens,although size estimation is difficult due to its fragmentary nature.
MOR 1128	"G-rex/Greg"	8%	2001	Museum of the Rockies	Bozeman	Greg Wilson	Hell Creek Formation	Montana
MOR 1152	"F-rex/Frank"	8%?	2001	Museum of the Rockies	Bozeman	Frank Stewart	Hell Creek Formation	Montana
FMNH PR2081	"Sue"	90%	1990	Field Museum of Natural History	Chicago	Sue Hendrickson	Hell Creek Formation	Faith, South Dakota	The largest known specimen at the time of discovery as well as one of the most complete.
RSM 2523.8	"Scotty"	70–75%	1991	Royal Saskatchewan Museum	Eastend	Tim Tokaryk Robert Gebhardt John Storer	Frenchman Formation	Saskatchewan	Proposed to be largest known specimen, scientific consensus not yet reached. Exceeds "Sue" in 84.6% of the published bone measurements. Has been estimated to be larger than "Sue" in two published studies.
PTRM 4667		10-20%	1992	Pioneer Trails Regional Museum	Bowman	Dean Pearson	Hell Creek Formation	Bowman County, North Dakota	Considered the 14th T-rex specimen to be found. The first in-situ t-rex to be known from the state of North Dakota. Discovered and collected by amateurs and shows tooth marks on bones, indicating scavenging.
TCM 2001.90.1	"Bucky"	34%	1998	The Children's Museum of Indianapolis	Indianapolis	Bucky Derflinger	Hell Creek Formation	Faith, South Dakota
BHI 6248	"E. D. Cope"	10%	1999	Black Hills Institute	South Dakota	Bucky Derfilinger	Hell Creek Formation	Faith, South Dakota
BMRP 2002.4.1	"Jane"	50%	2001	Burpee Museum of Natural History	Rockford	Carol Tuck William Harrison	Hell Creek Formation	Montana	An adolescent specimen, approximately 14 years old. Considered by a minority of paleontologists to be a separate taxon from Tyrannosaurus.
LACM 150167	"Thomas"	70%	2003	Natural History Museum of Los Angeles County	Los Angeles	Luis M. Chiappe	Hell Creek Formation	Montana
BHI 6437	"Bloody Mary"	98%	2006	North Carolina Museum of Natural Sciences	Raleigh	Clayton Phipps, Mark Eatman, & Chad O’Connor	Hell Creek Formation	Montana	Currently the most complete specimen known to science,an adolescent preserved entangled with a Triceratops. Like Jane, a minority of paleontologists consider it a different taxon.
MB.R.91216	"Tristan"	57%	2010	Natural History Museum, Berlin	Berlin	Craig Pfister	Hell Creek Formation	Carter County, Montana	Inventory number attributed by the Natural History Museum, Berlin, although the specimen belongs to a private collector.
Private	"Baby Bob"	20%	2013		Wichita	Robert (Bob) Detrich	Hell Creek Formation	Jordan, Montana	A juvenile or baby specimen. The discoverers have stated its age to be 4 years based on histology. An attempt was made to sell the specimen on eBay.
RGM 792.000	"Trix"	75–80%	2013	Naturalis Biodiversity Center	Leiden	Naturalis Biodiversity Center/Black Hills Institute	Hell Creek Formation	Montana	One of the largest known specimens that is also well preserved. Has been stated to be the oldest known specimen but this has not been confirmed.
Private	"Titus"	20%	2014		Nottingham	Craig Pfister	Hell Creek Formation	Montana	The specimen belongs to a private collector, but 3D-printed replicas of the bones are accessioned in the Nottingham Natural History Museum collection under the number NCMG 2021-7
UWBM 99000	"Tufts-Love"	30%	2016	Burke Museum of Natural History and Culture	Seattle	Jason Love Luke Tufts	Hell Creek Formation	Montana
AWMM-IL 2022.9	"Peter"	47%	2018	Auckland War Memorial Museum	Auckland	Dick Wills	Lance Formation	Wyoming
AWMM-IL2022.21	"Barbara"	44.7%	2005	Auckland War Memorial Museum	Auckland	Bob Harmon	Hell Creek Formation	Montana
NMMNH P-3698	Tyrannosaurus mcraeensis		1983	New Mexico Museum of Natural History and Science	Albuquerque	Donald Staton Joe LaPoint	Hall Lake Formation	New Mexico	The holotype of T. mcraeensis, dating to around 5–7 million years before T. rex

Manospondylus: AMNH 3982

The first-named fossil specimen which can be attributed to Tyrannosaurus rex consists of two partial vertebrae (one of which has been lost) found by Edward Drinker Cope in 1892. Cope believed that they belonged to an "agathaumid" (ceratopsid) dinosaur, and named them Manospondylus gigas, meaning "giant porous vertebra" in reference to the numerous openings for blood vessels he found in the bone. The M. gigas remains were later identified as those of a theropod rather than a ceratopsid, and H.F. Osborn recognized the similarity between M. gigas and Tyrannosaurus rex as early as 1917. However, due to the fragmentary nature of the Manospondylus vertebrae, Osborn did not synonymize the two genera.

Dynamosaurus: BMNH R7994

The holotype of Tyrannosaurus rex, a partial skull and skeleton originally called AMNH 973 (AMNH stands for American Museum of Natural History), was discovered in the U.S. state of Montana in 1902 and excavated over the next three years. Another specimen (AMNH 5866), found in Wyoming in 1900, was described in the same paper under the name Dynamosaurus imperiosus. At the time of their initial description and naming, these specimens had not been fully prepared and the type specimen of T. rex had not even been fully recovered. In 1906, after further preparation and examination, Henry Fairfield Osborn recognized both skeletons as belonging to the same species. Because the name Tyrannosaurus rex had appeared just one page earlier than Dynamosaurus in Osborn's 1905 work, it was considered the older name and has been used since. Had it not been for page order, Dynamosaurus would have become the official name.

Holotype: CM 9380

CM 9380 is the type specimen used to describe Tyrannosaurus rex. Fragments of (then) AMNH 973 were first found in 1902 by Barnum Brown, assistant curator of the American Museum of Natural History and a famous paleontologist in his own right. He forwarded news of it to Osborn; it would be three years before they found the rest of it. In 1905 when the type was described by Osborn, previous knowledge of dinosaur predators at the time were based on Jurassic carnosaurs, so the short fore-arms of the Tyrannosaurus were treated with extreme caution, with suspicion that bones of a smaller theropod had become jumbled with the remains of the bigger fossil. Following the 1941 entry of the United States into World War II, the holotype was sold to the Carnegie Museum of Natural History in Pittsburgh for protection against possible bombing raids. The specimen, now labeled CM 9380, is still mounted in Pittsburgh, at first with the tail acting as a tripod in the old-fashioned kangaroo pose. It has since received a modernization of its posture (mounted by Phil Fraley and crew) and can now be found balancing with tail outstretched. Along with a more lifelike posture, the specimen also now includes a composite reconstruction of the skull by Michael Holland. It has been reconstructed in recent years, it measured an estimated 11.9 meters in length and an estimated weight of 7.4–14.6 metric tonnes, 9.1 metric tonnes being the average estimate in that study, although most earlier studies have suggested lower weight figures.

AMNH 5027

With a length of 12.1-12.2 meters, AMNH 5027 was discovered and excavated in 1908 by Barnum Brown in Montana, and described by Osborn in 1912 and 1916. At the time of discovery, a complete cervical (neck vertebrae) series for Tyrannosaurus was not previously known, so it was this specimen that brought the short, stocky tyrannosaur neck to light. Compared to later specimens (BMNH R7994 and FMNH PR2081, for instance) the cervical series of AMNH 5027 is much more gracile, so with later discoveries the distinction between tyrannosaurid necks and the necks of carnosaurs became more obvious. This specimen also provided the first complete skull of Tyrannosaurus rex. In total, Brown found five partial Tyrannosaurus skeletons. The skeleton of this specimen was used as the iconic symbol for the Jurassic Park film series.

Osborn planned to mount the similarly sized AMNH 5027 and AMNH 973 together in dynamic poses. Designed by E.S. Christman, the scene was to depict a rearing Tyrannosaurus (AMNH 5027) snapping at another cowering one (AMNH 973), as they fought over the remains of a hadrosaur, described at the time as Trachodon. However, technical difficulties prevented the mount from being executed. One obvious problem was that the Cretaceous Dinosaur Hall was too small to accommodate this dramatic display, and AMNH 5027 was already mounted by itself as the central attraction of the hall. The forearms of Tyrannosaurus were not well documented and the hands were unknown, so for the sake of the display, the forearms of AMNH 5027 were given three fingers, based on the forelimbs of Allosaurus (the more allosaur-like arms were replaced several years later when better fossils of tyrannosaurid arms were found).

The mount retained a rearing pose similar to the initial proposal. By the 1980s it was generally accepted that such a pose would have been anatomically impossible in life, and the skeleton was re-mounted in a more accurate, horizontal pose during a renovation of the museum's dinosaur halls in the early 1990s. The mount can still be seen on display on the fourth floor of the American Museum. The American Museum of Natural History features AMNH 5027 in its famed Hall of Saurischian Dinosaurs to this day.

Nanotyrannus: CMNH 7541

A small but nearly complete skull of Nanotyrannus lancensis, frequently considered to be a juvenile T. rex, was recovered from Montana in 1942. This skull, Cleveland Museum of Natural History (CMNH) 7541, measures 60 centimeters (2.0 ft) in length and was originally classified as a species of Gorgosaurus (G. lancensis) by Charles W. Gilmore in 1946. In 1988, the specimen was re-described by Robert T. Bakker, Phil Currie, and Michael Williams, then the curator of paleontology at the Cleveland Museum of Natural History, where the original specimen was housed and is now on display. Their initial research indicated that the skull bones were fused, and that it therefore represented an adult specimen. In light of this, Bakker and colleagues assigned the skull to a new genus, named Nanotyrannus for its apparently small adult size. The specimen is estimated to have been around 5.2 meters (17 ft) long when it died. However, a detailed analysis of the specimen by Thomas Carr in 1999 showed that the specimen was, in fact, a juvenile, leading Carr and many other paleontologists to consider it a juvenile specimen of T. rex. The current classification of CMNH 7541 is not universal, with some research suggesting the specimen belongs to a taxon distinct from Tyrannosaurus.

LACM 23844

In 1966, a crew working for the Natural History Museum of Los Angeles County under the direction of Harley Garbani discovered another T. rex (LACM 23844) which included most of the skull of a very large, mature animal. When it was put on display in Los Angeles, LACM 23844 was the largest T. rex skull on exhibit anywhere.

"Black Beauty": RTMP 81.6.1

"Black Beauty" (specimen number RTMP 81.6.1) is a well-preserved fossil of Tyrannosaurus rex. The nickname stems from the apparent shiny dark color of the fossil bones, which occurred during fossilization by the presence of minerals in the surrounding rock; it was the first Tyrannosaurus rex specimen to receive a nickname, beginning a trend that continues with most major T. rex finds. Black Beauty was found in 1980 by a high school student, Jeff Baker, while on a fishing trip with a friend in the region of the Crowsnest Pass, Alberta. A large bone was found in the riverbank and shown to their teacher. Soon afterward, the Royal Tyrrell Museum was contacted, and excavation of the sandstone matrix surrounding the fossils began in 1982. The dig site where the fossil was found is located at approximately ${\displaystyle 114^{\circ }~{\textrm {W}}}$ near the confluence of the Crowsnest and Willow Rivers, and consisted of rock belonging to the Willow Creek Formation. The specimen is housed in the Royal Tyrrell Museum in Drumheller, Alberta, Canada.

In 2009, a paper by Jack Horner and colleagues illustrated the concept of parasitic infections in dinosaurs by analysing the lesions found on the cranial bones of Black Beauty. The specimen has been used to study comparative morphology between tyrannosaurids and Tyrannosaurus individuals.

Replicas of Black Beauty have been shown in some exhibitions and museums, including both simple skull montages and complete skeletons. Casts are on display in museums around the world.

"Stan": BHI 3033

Stan is the nickname given to a fossil about 11.78 m (38 ft) long found in Hell Creek Formation, South Dakota, close to Buffalo in 1987 by Stan Sacrison, who also discovered the Tyrannosaurus specimen nicknamed "Duffy". The original fossils are now housed at Black Hills Institute of Geological Research, Inc. center. It is a well known specimen, and one of the most complete, with 199 bones recovered. About 30 casts of the original fossil have been sold worldwide, each for a price of about $100,000. Stan's skeleton was auctioned for $31.8 million in a 2020 Christie's New York sale, making it a record-breaking dinosaur sale, with the buyer eventually being revealed as the under construction Natural History Museum Abu Dhabi in the Saadiyat Cultural District.

Like many other fossils of Tyrannosaurus rex, the skeleton of Stan shows many broken and healed bones. These include broken ribs and damages in the skull. One of the most prominent injuries are in the neck and the skull. A piece of bone is missing at the rear, and the skull also bears a hole 1 inch wide, probably made by another Tyrannosaurus. Also, two of the cervical vertebrae are fused, and another has additional bone growth. This could have been caused by another Tyrannosaurus bite. The bite marks are healed, indicating that Stan survived the wounds. Stan could also have been infected by Trichomonas-like parasites.

"Wankel Rex": MOR 555

In 1988, local rancher Kathy Wankel discovered another Tyrannosaurus rex in Hell Creek sediments on an island in the Charles M. Russell National Wildlife Refuge of Montana. This specimen was excavated by a team from the Museum of the Rockies led by paleontologist Jack Horner, with assistance from the U.S. Army Corps of Engineers. The specimen, given the number MOR 555 but informally called the "Wankel rex," includes approximately 80-85 percent of the skeleton, including the skull, as well as what at the time was the first complete T. rex forelimb. It has an estimated length of around 11.6 meters (38 ft) and a weight between 5.8 metric tons (6.4 short tons) and 10.8 metric tons (11.9 short tons) in newer figures. It is estimated that the "Wankel rex" was 18 years old when it died, an adult but not completely grown. The "Wankel rex" was also one of the first fossil dinosaur skeletons studied to see if biological molecules still existed within the fossilized bones. Doctoral candidate Mary Schweitzer found heme, a biological form of iron that makes up hemoglobin (the red pigment in blood).

It was long on exhibit at the Museum of the Rockies in Bozeman, Montana. In June 2013, the Corps loaned the specimen to the National Museum of Natural History, Smithsonian Institution museum in Washington, D.C., for 50 years. (The Museum of the Rockies continues to display a cast reconstruction of the skull by Michael Holland). The specimen went on temporary display on National Fossil Day, 16 October 2013, and was exhibited until the museum's dinosaur hall exhibit closed for renovation in the spring of 2014. The skeleton, named "The Nation's T. rex" became the centerpiece of the dinosaur hall when it reopened in 2019. Casts of MOR 555 are on display at the National Museum of Scotland, the Australian Fossil and Mineral Museum, and the University of California Museum of Paleontology. A bronze cast of the specimen, known as "Big Mike", stands outside the Museum of the Rockies.

In 2022, Gregory S. Paul and colleagues argued that the Wankel rex was not actually a T. rex, but rather the holotype for a new species: Tyrannosaurus regina. This was heavily criticized by several other leading paleontologists, including Stephen Brusatte, Thomas Carr, Thomas Holtz, David Hone, Jingmai O'Connor, and Lindsay Zanno when they were approached by various media outlets for comment. Their criticism was subsequently published in a technical paper.

"Sue": FMNH PR2081

Susan Hendrickson of the Black Hills Institute discovered the best-preserved Tyrannosaurus currently known, in the Hell Creek Formation near Faith, South Dakota, on 12 August 1990. About 90% of the skeleton was recovered, allowing the first complete description of a Tyrannosaurus rex skeleton. This specimen, named "Sue" in honor of its discoverer, soon became embroiled in a legal battle over its ownership. The owner of the land the fossil was found, Maurice Williams, as well the Sioux Tribe he belonged to, claimed ownership, the Institute had considered itself to have. In 1997, the suit was settled in favor of Williams and the fossil was returned to Williams' ownership. Williams quickly offered up "Sue" for auction by Sotheby's in New York, where it was sold to the Field Museum of Natural History in Chicago for US$8.4 million—the highest price ever paid for a fossil before being surpassed by Stan.

Sue has a length of 12.3–12.4 meters (40.4–40.7 ft), stands 3.66–3.96 m (12–13 ft) tall at the hips, and according to the most recent studies estimated to have weighed between 8.4 and 14 metric tons when alive. It has been hypothesised that Sue's impressive size may have been achieved due to a prolonged ontogenic development, since it is the third oldest Tyrannosaurus known. Sue's age at the time of death was estimated by Peter Mackovicky and the University of Florida to be 28 years old, over 6–10 years older than most big Tyrannosaurus specimens, like MOR 555, AMNH 5027 or BHI 3033. The only known specimen of T. rex that is older than Sue is Trix.

Preparation of "Sue" (FMNH PR2081) was completed at the Field Museum and the skeleton was placed on exhibit on 17 May 2000.

"Montana's T. rex": MOR980

Montana's T. rex (also known as "Peck's rex", "Peckrex", "Rigby's rex" and Tyrannosaurus "imperator") is the nickname given to a fossil specimen found in Montana in 1997. The discovery was made by Louis E. Tremblay on 4 July 1997 working under the supervision of J. Keith Rigby Jr. who led the excavation and bone preparation.

The skeleton of Montana's T. rex includes a relatively complete skull with jaws, multiple vertebrae of the back and tail, a well preserved gastralium, and hipbone with complete ischium and pubis. The left hindleg is relatively complete with a 1.2-meter-long (3.9 ft) femur, missing only some toe bones. The forelimbs include the scapula and furcula, both humeri and right hand phalanges, as well as metacarpal III. Montana's T. rex has been the subject of research regarding parasitic infections in dinosaurs. The forelimbs of Montana's T. rex have also been studied as they show evidence of use. This evidence includes the construction of metacarpal III, as well as repeated fractures in the furcula—possibly caused by heavy loads or pressure (Carpenter and Lipkin, 2005).

The fossils of Montana's T. rex are exhibited at Museum of the Rockies as part of a full skeletal mount completed with cast elements replacing the missing bones. This mount was installed after the Wankel Rex (now nicknamed The Nation's T. rex) was loaned to the Smithsonian to occupy a central part in the museum's dinosaur hall, featuring a dynamic mount showing the apex predator devouring a Triceratops carcass. It has been stated by Pete Makovicky, the Chicago museum's lead curator of dinosaurs, that this specimen is in the same size range as "Sue" and "Scotty".

"Bucky": TCM 2001.90.1

Bucky is a fossil of a juvenile specimen on display at The Children's Museum of Indianapolis in Indianapolis, Indiana. It is the first juvenile Tyrannosaurus ever placed on permanent exhibit in a museum. The dinosaur remains were found in 1998 in the Hell Creek Formation near the town of Faith, South Dakota. The skeleton, transported by water, ended up in a low shallow valley along with bones from an Edmontosaurus and Triceratops. It was discovered by rancher and cowboy Bucky Derflinger. Bucky was well preserved and easily prepared by the Black Hills Institute in South Dakota. Part of The Children's Museum of Indianapolis Dinosphere exhibit, Bucky is displayed along with Stan, an adult Tyrannosaurus, in a hunting scene. Both dinosaurs are attacking a Triceratops specimen known as "Kelsey".

Bucky is one of the few dinosaur fossils found with a furcula; Bucky's furcula was the first one found for the genus Tyrannosaurus. Bucky also has a nearly complete set of gastralia, or belly ribs, and an ulna, or lower arm bone. As of now, 101 bones, or about 34% of Bucky's skeleton, has been discovered and verified. Bucky is the sixth-most complete Tyrannosaurus rex out of more than 40 that have been discovered.

"E. D. Cope": BHI 6248

E. D. Cope (named after the paleontologist of the same name) is a Tyrannosaurus specimen discovered in South Dakota by Bucky Derflinger in 1999 at the same site as AMNH 3982. Excavations of this 10% complete skeleton began in 2000. The known material includes a partial skull, several vertebrae, and ribs. A very wide femur with a length of 1300 mm and a circumference of 630 mm is also known.

"Jane": BMRP 2002.4.1

Jane is a fossil specimen of small tyrannosaurid dinosaur, officially known as BMRP 2002.4.1, discovered in the Hell Creek Formation in southern Montana in 2001. Despite having a typically female name, Jane's sex is unknown—the specimen was named after Burpee Museum benefactor Jane Solem. The specimen was found in the summer of 2001 by Carol Tuck and Bill Harrison on an expedition led by Burpee Museum curator Michael Henderson. After four years of preparation, Jane was put on display at Rockford, Illinois' Burpee Museum of Natural History as the centerpiece of an exhibit called "Jane: Diary of a Dinosaur."

The Jane specimen has been central to the debate regarding the validity of the proposed tyrannosaurid genus Nanotyrannus. However, the Jane material has yet to be properly studied and described by scientists. Although Larson (2013) saw Jane as more identical to CMNH 7541 and LACM 28471 than to adult T. rex in having a higher tooth count, large pneumatic foramen on the center of the quadratojugal, T-shaped postorbital, and fused shoulder blade and pelvis, Yun (2015) concurred with the opinion of most workers that Nanotyrannus is a juvenile T. rex, noting that a juvenile specimen of Tarbosaurus described by Tsuihiji et al. (2011) also has a T-shaped postorbital. Paleontologists who support the theory that Jane represents a juvenile believe the tyrannosaur was approximately 11 years old at its time of death, and its fully restored skeleton measured 6.5 meters (21 ft) long, a bit more than half as long as the largest-known complete T. rex specimen, nicknamed "Sue," which measures 12.4 m (41 ft) long. According to Hutchinson et al. (2011), the weight of the Jane specimen in life was probably between 639 kg (1,409 lb) and 1,269 kg (2,798 lb), 954 kg (2,103 lb) being the average estimate.

"B-rex": MOR 1125

This specimen was found in the lower portion of the Hell Creek Formation near Fort Peck Lake in the Charles M. Russell National Wildlife Refuge in Garfield County, Montana. Its discoverer was Bob Harmon, a fossil preparator for the Museum of the Rockies, and was nicknamed the "B-rex" (or "Bob-rex") in honor of Harmon. The specimen was discovered in 2000, and excavated by MOR from 2001 to 2003. Although only 37 percent of the skeleton was present, this included almost all of the skull (although the skull was nearly completely disarticulated). The specimen also includes several cervical, dorsal, sacral, and caudal vertebrae; several chevrons; some cervical and dorsal ribs; left scapula and coracoid; the furcula; the left ulna; both femora, tibiae, and ulnae; the right calcaneum; right astragalus; and a number of pes phalanges.

In the March 2005 Science magazine, Mary Higby Schweitzer of North Carolina State University and colleagues announced the recovery of soft tissue from the marrow cavity of a fossilized femur belong to B-Rex. Flexible, bifurcating blood vessels and fibrous but elastic bone matrix tissue were recognized. In addition, microstructures resembling blood cells were found inside the matrix and vessels. The structures bear resemblance to ostrich blood cells and vessels. However, since an unknown process distinct from normal fossilization seems to have preserved the material, the researchers are being careful not to claim that it is original material from the dinosaur. Paleontologist Thomas Kaye of the University of Washington in Seattle hypothesized that the soft-tissue is permineralized biofilm created by bacteria while digesting and breaking down the original specimen. He has discovered this to be true in many specimens from the same area. In 2016, it was finally confirmed by Mary Higby Schweitzer and Lindsay Zanno et al that the soft tissue was medullary bone tissue, like that in modern birds when they are readying to lay eggs. This confirmed the identity of the Tyrannosaurus MOR 1125 as a female.

"Samson"

A T. rex specimen was discovered on private land in Harding County, South Dakota, once in 1981 by Michael and Dee Zimmerschied, and again on 4 October 1992 (Alan and Robert Detrich re-discovered Samson after it was originally found and deemed by paleontologists that several bones had washed in and there was nothing left). It was shortly after that when Fred Nuss and Candace Nuss of Nuss Fossils with the Detrich brothers found the most complete and undistorted Tyrannosaurus rex skull ever discovered. Following the sale of "Sue," another Tyrannosaurus rex skeleton was, the specimen was put up for auction on eBay in 2000 under the name of "Z-rex", with an asking price of over US$8 million. It failed to sell online but was purchased for an undisclosed price in 2001 by British millionaire Graham Ferguson Lacey, who renamed the skeleton "Samson" after the Biblical figure of the same name. It was prepared by the Carnegie Museum starting in May 2004. After preparation was complete in March 2006, the specimen was returned to Lacey. It, along with some other dinosaur skeletons, was sold again at auction on 3 October 2009.

Samson measured 11.9 m (39 ft), only slightly shorter than Sue.

"Baby Bob"

On 7 July 2013, fossil hunter Robert Detrich of Wichita, Kansas, unearthed the remains of what is believed to be a 4-year-old Tyrannosaurus rex. Detrich unearthed the fossil dubbed "Baby Bob" in a fossil-rich area near the Eastern Montana town of Jordan. Its femur measures about 25 inches, and if all the preliminary data pans out, that would make it among the smallest T. rex specimens ever found. Baby Bob has been fully excavated, although it will take another year to clean. Detrich said the skull, which is about 75 percent complete, and most of the major skeletal elements were found strewn across a flood plain, although very few vertebra and ribs were found.

"Scotty": RSM P2523.8

"Scotty", cataloged as RSM P2523.8, was discovered in Saskatchewan, Canada in 1991. Since its discovery and extensive subsequent study, "Scotty" has been referred to as the largest T. rex ever discovered in the world, the largest of any dinosaur discovered in Canada, and as one of the oldest and most complete fossils of its kind at more than 70% bulk. "Scotty" resides at the Royal Saskatchewan Museum's T. rex Discovery Centre in Eastend, SK, Canada. In May 2019, a second mount was erected at the Royal Saskatchewan Museum in Regina, where the exhibit reflects the recent discoveries about the fossil.

"Scotty" was discovered by Robert Gebhardt, a high school principal from Eastend, SK who accompanied palaeontologists from the Royal Saskatchewan Museum on a prospective expedition into the Frenchman Formation in southwestern Saskatchewan on 16 August 1991. It wasn't until June 1994 that the Royal Saskatchewan Museum was able to begin the excavation, which was led and overseen by the Museum's Ron Borden, as well as resident paleontologists Tim Tokaryk and John Storer who were with Gebhardt when he uncovered the first fossils. The bones were deeply packed in dense, iron-laden sandstone, which took more than twenty years for the team to fully remove, excavate, and assemble the majority of the skeleton, with additional trips being made to the site to retrieve smaller bones and teeth. The entire process of excavating the skeleton was also slowed down by its considerable size.

"Scotty" is reported to be 13.1 metres (43 feet) long and weighed an estimated 8.8 tonnes (8,800 kg). Despite it not being a complete fossil, paleontologists were able to create the estimation for the weight and length through measurements of important weight bearing bones such as the femur, hip, and shoulder bones that have all been measured to be larger and thicker with "Scotty" than the corresponding bones with "Sue". Going from the latest study "Scotty" exceeds "Sue" in 84.6% of the published measurements. While the reported measurements and weight for "Scotty" are larger than those of "Sue", some scientists posit that the two fossils are too close in size to officially declare "Scotty" the largest.

Like other T. rex fossils, "Scotty" shows signs of trichomoniasis, a parasitic infection in the jaw that left visible holes in the bone and was unique to this specific species of dinosaur. Additionally, a broken and healed rib on its right side, broken tail vertebra, as well as a hole near the eye socket are possibly the result of another T. rex attack. Other abnormalities, such as impacted teeth, suggest that "Scotty" was not only bitten, but also bit other animals.

"Tristan"

Commercial paleontologist Craig Pfister discovered the specimen in the lower Hell Creek Formation in Carter County, Montana, in 2010. Its excavation and preservation lasted four years. It was later sold to Danish-born investment banker Niels Nielsen, who loaned the specimen to the Museum für Naturkunde in Berlin, Germany, for research and exhibition. It has been on display at Museum für Naturkunde between 2015 and 2020, moving to the Natural History Museum of Denmark for one year, and expected back in 2021. Nielsen and his friend Jens Jensen named the specimen Tristan-Otto (short: Tristan) for their sons. The Museum für Naturkunde Berlin lists it under specimen number MB.R.91216. Several European museums have Tyrannosaurus casts (replicas) or parts, but Tristan is one of only two original skeletons on display in the continent (the other is "Trix" in the Netherlands). The matte-black fossilised skeleton is about 12 m (39 ft) long and 3.4 m (11 ft) tall at the hips. Tristan is among the most complete known Tyrannosaurus skeletons: It was re-assembled from about 300 separate parts, 170 of which are original (including 98% of the skull and all the teeth), the rest reproductions. It is estimated to have died when about 20 years old and it was in poor health, having several bone fractures, bite marks to the skull and signs of disease in the jaw. The disease present in Tristan's jaw was suggested to be a case of tumefactive osteomyelitis.

"Thomas"

From 2003 to 2005, Thomas was excavated by NHMLA paleontologists in southeastern Montana. At 17 years old, 34 ft (10 m) long and nearly 7,000 lb (3,200 kg), it is estimated to be a 70% complete specimen. Thomas is mounted in a "growth series" with the youngest-known Tyrannosaurus rex fossil, a two-year-old, 11-foot-long (3.4 m) specimen, and a 13-year-old, 20-foot-long (6.1 m) juvenile specimen.

This fossil is one of the geologically youngest T. rex specimens known, discovered very near the Cretaceous–Tertiary boundary.

"Victoria"

Victoria is a specimen found near Faith, South Dakota in 2013. Victoria is estimated to be around 12 ft tall and 40 ft long, and she is thought to have died in her subadult stage, between 15 and 25 years of age. Victoria has also been the subject of a traveling exhibition being displayed in places such as the Arizona Science Center. Her cause of death is unknown; however, she was believed to have been bitten in the lower jaw by another Tyrannosaurus. The bite may have become infected, spreading and leading to sepsis.

"Ivan"

Ivan is a 65% complete T. rex displayed at the Museum of World Treasures in Wichita, Kansas. The specimen has the most complete tail of any T. rex, only missing around 3 vertebrae. Ivan is around 40 ft long and 12 feet high.

"Trix": RGM 792.000

In 2013, a team of paleontologists from the Naturalis Biodiversity Center (Leiden, Netherlands) traveled to Montana where they discovered and unearthed a large and remarkably complete Tyrannosaurus rex specimen that lived 67 million years ago. Black Hills institute collaborated with the team in the excavation. The bones were cleaned and assembled in a mount at Black Hills Institute's installations, with the help of both Chicago's Field Museum of Natural History and the Naturalis Museum in Leiden. Chicago's Field museum sent digital models of their famous specimen, FMNH PR 2081 (Sue) to complete the cast and Naturalis museum replicated the bones using 3D-printing technology.

The specimen was named Trix after former Netherlands' Queen Beatrix. At arrival in the Netherlands, it started touring on public display in an itinerant exhibition titled T. rex in Town. The first exhibit spanned from 10 September 2016 to 5 June 2017 and was set at the only room of the Naturalis museum open to public at the time (the 17th-century building known as Pesthuis), due to the fact that the museum was undergoing restoration. When the Netherlands exhibition ended, it continued travelling through other European countries in 2017, 2018 and 2019. As of August 2019, Trix was returned to display at the Naturalis museum where it is installed in a special room that was under construction during Trix's European tour.

According to Peter Larson (director of Black Hills Institute), Trix is among the most complete Tyrannosaurus found. Between 75% and 80% of its skeletal volume was recovered. They are thought to have been at least 30 years old at death.

"Titus"

"Titus" is the name given to an obsidian-black skeleton of a Tyrannosaurus rex discovered in Montana's Hell Creek Formation in 2014 and excavated in 2018. It is 20% complete, and was named after the protagonist in Shakespeare's Titus Andronicus. Exhibited in the Nottingham Natural History Museum for 13 months beginning July 2021, it was during that time only the second specimen of Tyrannosaurus to be on exhibit in United Kingdom, the other being the type of the junior synonym Dynamosaurus imperosus which has the jaw on display in the Natural History Museum, London. External bone inspection has revealed injuries to Titus' right tibia (possibly a claw or bite wound); a deformed toe on the right foot; and a bitten and healed tail. The bite wound near the end of the tail indicates a possible attack by another Tyrannosaurus rex.

The remains of "Titus" were discovered in September 2014 by commercial paleontologist Craig Pfister near Ekalaka, Carter County, Montana. Excavation of the specimen began in 2018, and took 18 months. The bones of "Titus" were shipped to conservationist Nigel Larkin in the UK, who constructed the mount using a cast of the Tyrannosaurus specimen Stan to supplement the known bones of "Titus", after scanning the bones using photogrammetry to create digital models that were 3D printed for use in the exhibition, alongside the display of the real fossil skeleton, and which remained at the museum after the end of the exhibition. For the exhibit at the Nottingham Natural History Museum at Wollaton Hall, Titus was reconstructed in a walking pose.

"Tufts-Love": UWBM 99000

In 2016 Greg Wilson, David DeMar, and a paleontology team from the Burke Museum of Natural History and Culture, the University of Washington, and the Dig Field school excavated the partial remains of a Tyrannosaurus rex from Montana. The partial skeleton was found by two Burke Museum volunteers, Jason Love and Luke Tufts, and was named the "Tufts-Love" rex. Paleontologists at the Burke Museum believe that the Tufts-Love rex was around 15 years old when it died. The skull is of average size for an adult T. rex. The specimen was found in Late Cretaceous deposits and it is estimated to be 66.3 million years old. The Tufts-Love rex is undergoing preparation by Michael Holland and his team at the Burke Museum. The skeleton is estimated to be 30% complete, but it includes a complete (all of the bones of the skull and jaws are preserved) and mostly articulated skull. Holland describes the skull as minimally distorted and in an "exquisite" state of preservation.

"Peter": AWMM-IL 2022.9

Peter is the nickname given to a specimen on loan to the Auckland War Memorial Museum by an anonymous owner, currently on display alongside “Barbara” until the end of 2023. The specimen is estimated to be 66.8 million years old and almost of adult size. He was recovered from Niobrara County, Wyoming.

“Peter” is one of only four incredibly rare and visually stunning obisidian black colored tyrannosaurus rex.

He was likely killed by exocannibalism as entire sections of bone were damaged, and some were split open by huge crushing bite-forces. The nature of the crushing on the femur and tibia, along with size of the bite marks, indicates that these bones were bitten through by another Tyrannosaurus rex. There is also a set of smaller, parallel tooth marks nearby on the shaft that are not attributable to an adult T. rex. Explanations for this behavior range from response to over-crowded populations, limited food supply, sexual dominance, or even play.

"Barbara": AWMM-IL 2022.21

Barbara is the nickname given to a specimen on loan to the Auckland War Memorial Museum by the same anonymous owner as "Peter", currently on display alongside him until the end of 2023. The pair will be the first adult male and female T. rex to be displayed together. She is one of a few specimens believed to be pregnant.

"Barbara's" circumstances are particularly rare, more so when taken into account that she suffered and survived long after a debilitating foot injury. While she was no longer able to capture her prey, it is suggested that she got by with the help of a mate or cohort feeding her, as the injury would have rendered her immobile for upwards of 6 months. This is vaguely supported by trackway evidence that has been used to imply tyrannosaur group hunting. It is doubtful that "Barbara" ever successfully hunted again as a predator unless its prey was nearby.

"Bloody Mary": BHI 6437

"Bloody Mary" (as named by Pete Larson) is a nickname for a nearly complete tyrannosaurid (generally considered an adolescent T. rex or a "Nanotyrannus") at the North Carolina Museum of Natural Sciences. This specimen, considered the most complete of any Tyrannosaurus specimen at more than 98% preservation, was discovered in Montana in 2006. Following years of unsuccessful attempts to sell it to museums or auction it off, the NCMNS started negotiations in 2016, which were prolonged due to a legal battle over the rights to the fossil, which was resolved in 2020. The fossils were officially displayed in 2024.

The Dueling Dinosaurs T. rex is an adolescent that was preserved entangled with a Triceratops; their combined fossil is referred to as the "Dueling Dinosaurs". Given the injuries present on both fossils, it has been theorized that both died while fighting one another. The specimen is presently undergoing study. Important biological data is likely preserved within the specimen, including body outlines, skin impressions, soft tissues, injuries, stomach contents, and even original proteins.

Tyrannosaurus mcraeensis: NMMNH P-3698

The remains of a tyrannosaur were discovered in 1983 in the Campanian-early Maastrichtian Hall Lake Formation in New Mexico by Donald Staton and Joe LaPoint. Reposited at the New Mexico Museum of Natural History and Science, the fossil material (NMMNH P-3698) consists of skull and lower jaw bones, in addition to isolated teeth and chevrons. Some of the bones were briefly mentioned in 1984 as belonging to T. rex, and were described in 1986. In 2024, Sebastian G. Dalman and colleagues described this specimen as the holotype of a new Tyrannosaurus species, T. mcraeensis. This species differs from T. rex in having smaller postorbital crests, a proportionately longer and shallower lower jaw with a less prominent chin suggestive of a weaker bite, and more laterally compressed teeth.

See also

• Timeline of tyrannosaur research

References

1. Larkin, N.R., Dey, S., Smith, A.S. and Evans, R. 2022. 21st Century Rex: maximising access to a privately owned Tyrannosaurus rex skeleton in the digital age. Geological Curator, 11(6), 341–354.
2. ^ Dalman, Sebastian G.; Loewen, Mark A.; Pyron, R. Alexander; Jasinski, Steven E.; Malinzak, D. Edward; Lucas, Spencer G.; Fiorillo, Anthony R.; Currie, Philip J.; Longrich, Nicholas R. (11 January 2024). "A giant tyrannosaur from the Campanian–Maastrichtian of southern North America and the evolution of tyrannosaurid gigantism". Scientific Reports. 13 (1): 22124. doi:10.1038/s41598-023-47011-0. ISSN 2045-2322. PMC 10784284. PMID 38212342.
3. Quinlan, Elizibeth D.; Derstler, Kraig; Miller, Mercedes M. (2007). "Anatomy and function of digit III of the Tyrannosaurus rex manus". Geological Society of America Annual Meeting — Abstracts with Programs: 77. Archived from the original on 24 February 2008. Retrieved 28 May 2012.
4. Osborn, H. F. (1917). "Skeletal adaptations of Ornitholestes, Struthiomimus, Tyrannosaurus". Bulletin of the American Museum of Natural History. 35 (43). New York City: American Museum of Natural History: 733–771. hdl:2246/1334.
5. ^ Osborn, H.F. (1905). "Tyrannosaurus and other Cretaceous carnivorous dinosaurs". Bulletin of the American Museum of Natural History. 21 (3): 259–265. doi:10.5281/zenodo.1038151. hdl:2246/1464.
6. Osborn, Henry Fairfield; Brown, Barnum (1906). "Tyrannosaurus, Upper Cretaceous carnivorous dinosaur". Bulletin of the American Museum of Natural History. 22. hdl:2246/1473.
7. Brown, B. (1915). "Tyrannosaurus, a Cretaceous carnivorous dinosaur, the largest flesh-eater that ever lived". Scientific American. 63 (15): 322–323. doi:10.1038/scientificamerican10091915-322.
8. Norell, M. A.; Gaffney, E. S.; Dingus, L. (1995). Discovering Dinosaurs in the American Museum of Natural History. New York: Knopf. p. 117. ISBN 0-679-43386-4.
9. Makovicky, Peter J.; Allen, Vivian; Molnar, Julia; Bates, Karl T.; Hutchinson, John R. (12 October 2011). "A Computational Analysis of Limb and Body Dimensions in Tyrannosaurus rex with Implications for Locomotion, Ontogeny, and Growth". PLOS ONE. 6 (10): e26037. Bibcode:2011PLoSO...626037H. doi:10.1371/journal.pone.0026037. PMC 3192160. PMID 22022500.
10. Kenneth Carpenter & Philip J. Currie (1990). "Variation in Tyrannosaurus rex". Dinosaur Systematics. Cambridge University Press. p. 143. ISBN 0-521-43810-1.
11. Osborn, H.F. (1913). "Tyrannosaurus: restoration and model of the skeleton". Bulletin of the American Museum of Natural History. 32: 91–92. hdl:2246/1735.
12. Gilmore, C. W. (1946). "A new carnivorous dinosaur from the Lance Formation of Montana". Smithsonian Miscellaneous Collections. 106: 1–19.
13. Bakker, R.T.; Williams, M.; Currie, P.J. (1988). "Nanotyrannus, a new genus of pygmy tyrannosaur, from the latest Cretaceous of Montana". Hunteria. 1: 1–30.
14. Carr, T.D. (1999). "Craniofacial ontogeny in Tyrannosauridae (Dinosauria, Coelurosauria)". Journal of Vertebrate Paleontology. 19 (3): 497–520. Bibcode:1999JVPal..19..497C. doi:10.1080/02724634.1999.10011161. S2CID 83744433.
15. Tsuihiji, T.; Watabe, M.; Tsogtbaatar, K.; Tsubamoto, T.; Barsbold, R.; Suzuki, S.; Lee, A.H.; Ridgely, R.C.; Kawahara, Y.; Witmer, L.M. (2011). "Cranial osteology of a juvenile specimen of Tarbosaurus bataar from the Nemegt Formation (Upper Cretaceous) of Bugin Tsav, Mongolia". Journal of Vertebrate Paleontology. 31 (3): 497–517. Bibcode:2011JVPal..31..497T. doi:10.1080/02724634.2011.557116. S2CID 15369707.
16. Longrich, Nicholas R.; Saitta, Evan T. (1 March 2024). "Taxonomic Status of Nanotyrannus lancensis (Dinosauria: Tyrannosauroidea)—A Distinct Taxon of Small-Bodied Tyrannosaur". Fossil Studies. 2 (1): 1–65. doi:10.3390/fossils2010001. eISSN 2813-6284.
17. ^ Horner, J.R. & Lessem, D. 1993. The Complete T. rex New York: Simon & Schuster. 238pp.
18. ^ Larson, Peter H. (2008). "One Hundred Years of Tyrannosaurus Rex: The Skeletons.". In Larson, Neal L. (ed.). Tyrannosaurus rex the Tyrant King. pp. 16–17.
19. Edgar, Blake. "Discovery Channel's homepage". Dsc.discovery.com. Archived from the original on 10 October 2012. Retrieved 29 October 2012.
20. "www.unearthingtrex.com". Unearthingtrex.com. 26 June 2002. Archived from the original on 17 July 2012. Retrieved 29 October 2012.
21. "www.uleth.ca". Uleth.ca. Archived from the original on 23 February 2012. Retrieved 11 January 2012.
22. Wolff, ED; Salisbury, SW; Horner, JR; Varricchio, DJ (2009). "Common Avian Infection Plagued the Tyrant Dinosaurs". PLOS ONE. 4 (9): e7288. Bibcode:2009PLoSO...4.7288W. doi:10.1371/journal.pone.0007288. PMC 2748709. PMID 19789646.
23. Many sources says that "Stan" is the second most complete specimen, except for "Sue" (FMNH PR 2081). 65% of his total skeleton was found "Yesterday.sg: SUE, STAN & Friends were in town…". Archived from the original on 1 October 2011. Retrieved 1 October 2011."Stan (The University of Manchester)". Archived from the original on 18 September 2010. Retrieved 18 September 2010.
24. The Manchester Museum's homepage: "Stan the Tyrannosaurus rex"
25. Sign On San Diego"New Mexico museum has funds for half a T. rex" Archived 16 July 2011 at the Wayback Machine, 26 February 2006.
26. Get Cynical: "T-Rex Replica Skeletons for Sale", 21 July 2008.
27. Small, Zachary (7 October 2020). "T. Rex Skeleton Brings $31.8 Million at Christie's Auction". The New York Times.
28. "Stan the T. rex found! World's most expensive fossil finds home in a new museum". National Geographic. 23 March 2022. Archived from the original on 23 March 2022.
29. New Mexico Museum of Natural History and Science homepage: "T. REX ATTACK!" Archived 5 June 2010 at the Wayback Machine
30. Wolff, Ewan D. S.; Salisbury, Steven W.; Horner, John R.; Varricchio, David J. (30 September 2009). "Common Avian Infection Plagued the Tyrant Dinosaurs". PLOS ONE. 4 (9): e7288. Bibcode:2009PLoSO...4.7288W. doi:10.1371/journal.pone.0007288. PMC 2748709. PMID 19789646.
31. Institution, Smithsonian. "Smithsonian Welcomes "The Nation's T. rex" to Washington, D.C." Smithsonian Institution. Retrieved 30 March 2024.
32. ^ Hutchinson, J. R.; Bates, K. T.; Molnar, J.; Allen, V.; Makovicky, P. J. (2011). "A Computational Analysis of Limb and Body Dimensions in Tyrannosaurus rex with Implications for Locomotion, Ontogeny, and Growth". PLOS ONE. 6 (10): e26037. Bibcode:2011PLoSO...626037H. doi:10.1371/journal.pone.0026037. PMC 3192160. PMID 22022500.
33. ^ Schontzler, Gail. "Montana T. rex Heading to Smithsonian." Bozeman Daily Chronicle. 28 June 2013. Accessed 28 June 2013.
34. Wu, Katherine, J. (17 July 2018). "Homecoming King: The Nation's T. rex Returns to the Smithsonian". Smithsonian Magazine.{{cite web}}: CS1 maint: multiple names: authors list (link)
35. du Lac, J. Freedom "Tyrannosaurus rex Gets Long-Term Lease at Smithsonian's Natural History Museum." Washington Post. 27 June 2013. Accessed 28 June 2013.
36. "T. rex arrives at National Museum of Scotland". Archived from the original on 20 September 2011. Retrieved 29 July 2011.
37. ^ Paul, Gregory S.; Persons IV, W. Scott; van Raalte, Jay (2022). "The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus Tyrannosaurus". Evolutionary Biology. 49 (2): 156–179. Bibcode:2022EvBio..49..156P. doi:10.1007/s11692-022-09561-5. S2CID 247200214.
38. Elbein, Asher (28 February 2022). "They Want to Break T. Rex Into 3 Species. Other Paleontologists Aren't Pleased". The New York Times. Archived from the original on 1 March 2022. Retrieved 1 March 2022.
39. Hunt, Katie (1 March 2022). "Tyrannosaurus rex may have been misunderstood". CNN.
40. Greshko, Michael (1 March 2022). "Call to split T. rex into 3 species sparks fierce debate". National Geographic. Archived from the original on 1 March 2022.
41. Carr, Thomas D.; Napoli, James G.; Brusatte, Stephen L.; Holtz, Thomas R.; Hone, David W. E.; Williamson, Thomas E.; Zanno, Lindsay E. (1 September 2022). "Insufficient Evidence for Multiple Species of Tyrannosaurus in the Latest Cretaceous of North America: A Comment on "The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus Tyrannosaurus"". Evolutionary Biology. 49 (3): 327–341. Bibcode:2022EvBio..49..327C. doi:10.1007/s11692-022-09573-1. ISSN 1934-2845.
42. The Field Museum. "Unearthing the Secrets of Sue: Educator Guide" (PDF) (Second ed.). pp. 3, 52. Archived from the original (PDF) on 3 October 2018. Retrieved 9 July 2016.
43. Brochu, C.R. (2003). "Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull". Memoirs of the Society of Vertebrate Paleontology. 7: 1–138. doi:10.2307/3889334. JSTOR 3889334.
44. Holtz, T. R. (2011). "Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2011 Appendix" (PDF). Retrieved 13 January 2012.
45. "Sue Fact Sheet" (PDF). Sue at the Field Museum. Field Museum of Natural History. Archived from the original (PDF) on 18 August 2016.
46. "How well do you know SUE?". Field Museum of Natural History. 11 August 2016. Retrieved 31 December 2018.
47. "Sue the T. Rex". Field Museum. 5 February 2018. Retrieved 20 July 2018.
48. Hartman, Scott (7 July 2013). "Mass estimates: North vs South redux". Scott Hartman's Skeletal Drawing.com. Retrieved 24 August 2013.
49. kcook (30 June 2014). "SUE Educator Guide". Field Museum. Retrieved 23 March 2019.
50. ^ "T. rex is coming to Naturalis" (PDF). Archived from the original (PDF) on 15 December 2016. Retrieved 12 September 2016.
51. ^ Koen Stein, paleontologist at the Free University of Brussels (Vrije Universiteit Brussel), "Oldest T. rex Trix on view in Leiden, VUB researchers established the dinosaur's age", VUB Today, 23 January 2017 Archived 22 October 2019 at the Wayback Machine
52. "The Story of a Dinosaur Named Sue" Archived 27 December 2014 at the Wayback Machine by Neal Larson. Black Hills Institute of Geological Research, Inc. 19 May 2000. Retrieved 16 July 2006.
53. "Sue at the Field Museum" Archived 4 April 2014 at the Wayback Machine Field Museum of Natural History. 9 September 2005. Retrieved 16 July 2006.
54. "article-niche.com". Article-niche.com. 4 March 2009. Retrieved 29 October 2012.
55. "Parasite may have felled a mighty T. rex | Paleontology". Science News. Retrieved 29 October 2012.
56. "dml.cmnh.org". Dml.cmnh.org. 13 June 2005. Archived from the original on 13 November 2007. Retrieved 29 October 2012.
57. "New exhibit at Museum of the Rockies features real fossilized T. Rex".
58. "What's the Army doing with dinosaurs? Redux". www.army.mil. 13 April 2015. Retrieved 23 March 2019.
59. "New exhibit at Museum of the Rockies features real fossilized T. Rex".
60. "Scotty vs. Sue: Is the Canadian T. Rex really bigger than Chicago's? The Field Museum disputes new study". Chicago Tribune. 29 March 2019.
61. ^ Crosslin, Rick; Mary Fortney; Dinosphere Exhibit Development Team (2004). "Resource Materials: Bucky-Tyrannosaurus rex". Dinosphere: A 3–5 Unit of Study.
62. ^ Larson, P.L.; Carpenter, K. (2008). Tyrannosaurus Rex, the Tyrant King. Life of the past. Indiana University Press. ISBN 978-0-253-35087-9. Retrieved 22 June 2020.
63. Larson, Neal L.; Peter Larson; Kenneth Carpenter (2008). "One Hundred Years of Tyrannosaurus Rex: The Skeletons". Tyrannosaurus Rex, the Tyrant King: 43–44.
64. "Meet the Dinos of Dinosphere | The Children's Museum of Indianapolis". www.childrensmuseum.org. Retrieved 24 January 2021.
65. Crosslin, Rick; Mary Fortney; Dinosphere Exhibit Development Team (2004). "Tyrannosaurus rex Attack Scene-What will be the outcome?". Dinosphere: A 3–5 Unit of Study: 88.
66. "The Origin of Birds". The Macro Library. University of California. Archived from the original on 15 June 2012. Retrieved 5 April 2011.
67. Larson, Peter. The First Recognized Furcula For Tyrannosaurus Rex. Black Hills Institute of Geological Research.
68. "NIU teams up with Burpee Museum to bring world's top dinosaur hunters to Rockford". Northern Illinois University. 7 September 2007. Archived from the original on 5 August 2012. Retrieved 9 September 2007.
69. News Release: NIU teams up with Burpee Museum to bring world's top dinosaur hunters to Rockford Archived 5 August 2012 at archive.today
70. Larson, P. L. 2013. The case for Nanotyrannus in J. M. Parrish, R. A. Molnar, P. J. Currie., and E. B. Koppelhus (eds.) Tyrannosaurid Paleobiology, University of Indiana Press, Bloomington and Indianapolis, pp. 15–53.
71. Yun, C (2015). "Evidence points out that "Nanotyrannus" is a juvenile Tyrannosaurus rex". PeerJ Preprints. 3: e1052. doi:10.7287/peerj.preprints.852v1.
72. Schweitzer, M.H.; Wittmeyer, J.L.; Horner, J.R.; Toporski, J.B. (2005). "Soft Tissue Vessels and Cellular Preservation in Tyrannosaurus rex". Science. 307 (5717): 1952–1955. Bibcode:2005Sci...307.1952S. doi:10.1126/science.1108397. PMID 15790853. S2CID 30456613.
73. Fox, Maggie (30 July 2008). "Scientists question dinosaur soft tissue find". Reuters. Archived from the original on 22 July 2012. Retrieved 30 July 2008.
74. Schweitzer, Mary Higby; Zheng, Wenxia; Zanno, Lindsay; Werning, Sarah; Sugiyama, Toshie (15 March 2016). "Chemistry supports the identification of gender-specific reproductive tissue in Tyrannosaurus rex". Scientific Reports. 6 (1): 23099. Bibcode:2016NatSR...623099S. doi:10.1038/srep23099. ISSN 2045-2322. PMC 4791554. PMID 26975806.
75. ^ "Carnegie Museum digs into controversial, but promising T-rex skull" by Byron Spice. Pittsburgh Post-Gazette. 12 May 2004. Retrieved 16 July 2006.
76. "It's "Bone Voyage" for a famous fossil" by Dan Lagiovane. Carnegie Museum of Natural History . 2 March 2006. Retrieved 30 December 2010.
77. "Bonhams : The T. rex known as Samson - One of the Most Complete Tyrannosaurus rex Specimens in Existence". www.bonhams.com. Retrieved 23 March 2019.
78. "New Arrivals – T-Rex & Nanotyrannus Teeth From South Dakota". FossilEra. Retrieved 23 March 2019.
79. ^ Persons, W. Scott; Currie, Philip J.; Erickson, Gregory M. (April 2020). "An Older and Exceptionally Large Adult Specimen of Tyrannosaurus rex". The Anatomical Record. 303 (4): 656–672. doi:10.1002/ar.24118. PMID 30897281. S2CID 85448862.
80. ^ "The Cretaceous « T.rex Discovery Centre". royalsaskmuseum.ca. Archived from the original on 11 April 2019. Retrieved 11 April 2019.
81. ^ "World's Largest T. Rex Skeleton nicknamed "Scotty" Discovered in Canada". The Vintage News. 25 March 2019. Retrieved 12 April 2019.
82. Ashley Strickland (26 March 2019). "Meet 'Scotty,' the largest Tyrannosaurus rex ever discovered". CNN. Retrieved 11 April 2019.
83. ^ Holson, Laura M. (28 March 2019). "'Scotty' the T. Rex Is the Heaviest Ever Found, Scientists Say". The New York Times. ISSN 0362-4331. Retrieved 11 April 2019.
84. Johnson, Steve (29 March 2019). "Scotty vs. Sue: Is the Canadian T. rex really bigger than Chicago's? The Field Museum disputes new study". Chicago Tribune. Retrieved 12 April 2019.
85. ^ Francis, Jennifer (22 March 2019). "Sask. T-rex Scotty is officially biggest ever discovered". cbc.ca.
86. Article about Scotty on Osel.cz (in Czech)
87. https://docs.google.com/spreadsheets/d/1xbuGjf_PaBcJNKJSu4sEixEqZzEc-wyvfrg1iTnCSno/edit#gid=0
88. Choi, Charles Q. (29 September 2009). "Mighty T. rex Killed by Lowly Parasite, Study Suggests". Live Science. Retrieved 11 April 2019.
89. ^ Cookson, Clive (25 September 2015). "How to buy a Tyrannosaurus Rex". Financial Times. Retrieved 1 January 2016.
90. ^ "Tristan – Berlin bares teeth". Museum für Naturkunde. Archived from the original on 1 April 2019. Retrieved 18 January 2019.
91. ^ Gennies, Sidney (17 December 2015). "T-Rex Tristan: Entzauberung eines Mythos". Der Tagesspiegel. Retrieved 1 January 2016.
92. "Nach drei Jahren verlässt T. rex Tristan Otto Berlin". Berliner Morgenpost. 26 November 2018. Archived from the original on 3 April 2019. Retrieved 18 January 2019.
93. Back in 2021 Back in 2021 - Tristan, accessed 13 February 2020
94. Karberg, Sascha (13 July 2015). "Schädel eines T. Rex in Berlin eingetroffen". Der Tagesspiegel. Retrieved 1 January 2016.
95. "tristan-otto". Museum für Naturkunde. Retrieved 1 January 2016.
96. ^ Okke, Tom (17 December 2015). "Dansk velhaver investerer i ægte Tyrannosaurus rex". Børsen. Retrieved 18 January 2019.
97. Ring, Caroline (25 July 2015). "Der T. rex von Berlin". Spektrum.de. Retrieved 18 January 2019.
98. "Famous T. Rex had a bone infection, new medical scans reveal". Live Science. December 2021.
99. "Dinosaur Hall". Natural History Museum of Los Angeles County. 15 December 2010. Retrieved 4 June 2018.
100. "Collection Highlights". dinosaurs.nhm.org. Retrieved 4 June 2018.
101. ^ "Victoria, the 66-million-year-old Tyrannosaurus rex, will have her exhibit extended at the Arizona Science Center". KNXV. 16 October 2020. Retrieved 27 June 2021.
102. "Meet Victoria, one of the most complete T. rex fossils in the world". CNN. 12 September 2019.
103. "Victoria, the 66-million-year-old Tyrannosaurus rex, will have her exhibit extended at the Arizona Science Center". ABC15. 16 October 2020.
104. "Victoria: The T. rex | Houston Museum Of Natural Science". www.hmns.org. Retrieved 1 July 2021.
105. Reidl, Matt (15 February 2018). "Wichita gets to keep its T. rex skeleton in Old Town". The Wichita Eagle.
106. "Unit". September 2016. Archived from the original on 4 November 2016. Retrieved 12 September 2016.
107. "Trix the T Rex makes her mark on Leiden in new exhibition". DutchNews.nl. 6 September 2019. Retrieved 6 September 2019.
108. http://www.thefossilforum.com/uploads/monthly_2016_09/10.thumb.jpg.592a065fd0a51cd2c23fa6d1a2063c29.jpg
109. Johnson, Steve (24 August 2016). "Don't worry, Sue – Trix is just passing through". Chicago Tribune. Retrieved 23 March 2019.
110. ^ Burnham, David A.; Rothschild, Bruce M.; Nudds, John R. (2021). "A Paleontological Study of a New Specimen of Tyrannosaurus rex Named "Titus" Nottingham Natural History Museum Specimen #NGMG 2022-7" (PDF). Wollaton Hall. Archived from the original (PDF) on 8 March 2022. Retrieved 30 June 2022.
111. "Titus T. rex Scientific Report: Discussion". Wollaton Hall. 2 February 2022. Retrieved 25 March 2022.
112. Pinchess, Lynette (1 July 2021). "Real T. Rex descends on Nottingham in a world exclusive". NottinghamshireLive. Retrieved 25 March 2022.
113. Ingram, Simon (12 May 2021). "'Titus' the T. rex is coming to the UK this summer. Here's why it's a big deal". National Geographic. Retrieved 30 June 2022.
114. "Mounting the skeleton of 'Titus' the Tyrannosaurus rex". Natural-History-Conservation.com. Retrieved 30 June 2022.
115. "Burke Museum team discovers a T. rex". Burke Museum. 17 August 2016. Retrieved 23 March 2019.
116. "Tyrannosaurus rex: A gigantic beast". Burke Museum. 28 June 2018. Retrieved 23 March 2019.
117. Basagre, Bernadette (1 April 2022). "Peter the T Rex: Auckland Museum hosts world debut of dinosaur bones found in 2018". Stuff. Retrieved 30 May 2022.
118. Burnham, David A.; Rothschild, Bruce M.; Nudds, John R. (2022). Peter the T. rex, a Paleontological Study on a New Specimen of Tyrannosaurus Rex Named 'Peter', from the Lance Formation (Maastrichtian) of Wyoming specimen number AWMM-IL 2-22.9 (PDF) (Report). Auckland War Memorial Museum. Retrieved 25 June 2022.
119. Burnham, M.; Mclain, M.; Nelson, D.R.; Snyder, K. (2018). "Tyrannosaur cannibalism: a case of a tooth-traced tyrannosaurid bone in the Lance Formation (Maastrichtian), Wyoming". PALAIOS. 33 (4): 164–173. Bibcode:2018Palai..33..164M. doi:10.2110/palo.2017.076. S2CID 134871802.
120. "Peter Scientific Report" (PDF). 2022. Retrieved 4 December 2022.
121. Anderson, Ryan (24 November 2022). "Skeleton of pregnant T-Rex Barbara joins Auckland Museums growing collection". Stuff. Retrieved 4 December 2022.
122. ^ Burnham, David A.; Rothschild, Bruce M.; Nudds, John R. (2022). Barbara the T. rex, a Paleontological Study on a New Specimen of Tyrannosaurus Rex Named 'Barbara', from the Hell Creek Formation (Maastrichtian) of Montana specimen number AWMM-IL2022.21 (PDF) (Report). Auckland War Memorial Museum. Retrieved 4 December 2022.
123. accpaleo (11 March 2018). "Nanotyrannus: Up Hell Creek Without a Paddle". A Central Coast Paleontologist. Retrieved 7 June 2024.
124. Keats, Jonathon. "Montana's Dueling Dinosaurs To Fetch Up To $9 Million At Bonhams Auction". Forbes. Retrieved 18 November 2020.
125. "Dueling Dinosaurs | North Carolina Museum of Natural Sciences". Dueling Dinosaurs | North Carolina Museum of Natural Sciences. Retrieved 10 November 2022.
126. "North Carolina Museum of Natural Sciences to receive the "Dueling Dinosaurs" | Programs and Events Calendar". Retrieved 18 November 2020.
127. "'Dueling Dinosaurs' fossil, hidden from science for 14 years, could finally reveal its secrets". Science. 17 November 2020. Archived from the original on 17 November 2020. Retrieved 18 November 2020.
128. Lozinsky, Richard P.; Hunt, Adrian P.; Wolberg, Donald L.; Lucas, Spencer G. (1984). "Late Cretaceous (Lancian) dinosaurs from the McRae Formation, Sierra County, New Mexico" (PDF). New Mexico Geology. 6 (4): 72–77. doi:10.58799/NMG-v6n4.72. ISSN 0196-948X. S2CID 237011797.
129. Gillette, David D.; Wolberg, Donald L.; Hunt, Adrian P. (1986). "Tyrannosaurus rex from the McRae Formation (Lancian, Upper Cretaceous), Elephant Butte reservoir, Sierra County, New Mexico" (PDF). New Mexico Geological Society Guidebook. 37: 235–238. doi:10.56577/FFC-37.235. S2CID 251985284.

External links

• Listing of Tyrannosaurus rex specimens at The Theropod Database.
• The Black Hills Institute's article on Stan Archived 12 October 2007 at the Wayback Machine
• AMNH Article on the First Tyrannosaur specimens
• Comparison of known skeletal material and sizes for five tyrannosaur specimens at Skeletal Drawing.

• Collection of the Carnegie Museum of Natural History
• Tyrannosaurus
• Lists of dinosaur specimens
• Collection of the Smithsonian Institution
• Cretaceous fossil record