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Nannippus Temporal range: late Miocene- early Pleistocene
Jaw of Nannippus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Perissodactyla
Family:	Equidae
Subfamily:	Equinae
Tribe:	†Hipparionini
Genus:	†Nannippus Matthew, 1926

Nannippus is an extinct genus of three-toed horse endemic to North America during the Miocene through Pleistocene, about 13.3—1.8 million years ago (Mya), living around 11.5 million years. This ancient species of three-toed horse grew up to 3.5 feet (1.1 meters) and weighed between 165 pounds to 199 pounds, which was around the same size as a domestic sheep.

Nannippus lived as far south as central Mexico (N. peninsulatus) to as far north as Canada (N. lenticularis), to California in the west, and North Carolina (N. lenticularis) and Florida (N. peninsulatus) in the east. Unlike its relatives, Nannippus, as well as all other three-toed horses endemic to North America, were browsers and mixed feeders that had diets consisting of C3 plants.

Species

• N. aztecus Mooser, 1968. Widespread in Florida and also found in Texas, Oklahoma, and Chihuahua, became extinct 11.2—5.7 Mya.
• N. beckensis Dalquest and Donovan, 1973 found in Texas only and became extinct about 3.4 Mya.
• N. lenticularis (Cope, 1893) was found in Alberta, Canada, Texas, North Carolina, Alabama, Nebraska, and Kansas, becoming extinct about 13 Mya.
• N. minor (Sellards, 1916) lived in Georgia during the Pliocene.
• N. montezumae (Leidy, 1882)
• N. morgani Hulbert, 1993 is restricted to Florida and appears to have become extinct about 8.6 Mya.
• N. peninsulatus (Cope, 1893) was found in Florida, Texas, Oklahoma, New Mexico, Arizona, and Mexico, becoming extinct about 3.3 Mya.
• N. parvulus (Marsh, 1868)
• N. westoni Simpson, 1930 is restricted to Florida and became extinct about 9.1—8.7 Mya.

References

1. MacFadden, Bruce J. (1984). "Systematics and phylogeny of Hipparion, Neohipparion, Nannippus, and Cormohipparion (Mammalia, Equidae) from the Miocene and Pliocene of the New World. Bulletin of the AMNH; v. 179, article 1". hdl:2246/997. {{cite journal}}: Cite journal requires |journal= (help)
2. Johnston, C. Stuart (1938). "The Skull of Nannippus gratus (Leidy) from the Lower Pliocene of Texas". The American Midland Naturalist. 19 (1): 245–248. doi:10.2307/2420435. ISSN 0003-0031. JSTOR 2420435.
3. Hulbert, Richard C. (1993). "Late Miocene Nannippus (Mammalia: Perissodactyla) from Florida, with a Description of the Smallest Hipparionine Horse". Journal of Vertebrate Paleontology. 13 (3): 350–366. Bibcode:1993JVPal..13..350H. doi:10.1080/02724634.1993.10011515. ISSN 0272-4634. JSTOR 4523517.
4. "Nannippus". prehistoric-fauna.com. Retrieved 2021-06-23.
5. Tudge, Colin (April 3, 1994). "Science: Why the four-toed horse lost the race: If evolution is random, why has the family Equidae grown steadily larger and faster? The answer lies in the gut, says Colin Tudge". Independent.co.uk. Archived from the original on 2022-06-18. Retrieved August 10, 2021.
6. MacFadden, Bruce J.; Solounias, Nikos; Cerling, Thure E. (1999-02-05). "Ancient Diets, Ecology, and Extinction of 5-Million-Year-Old Horses from Florida". Science. 283 (5403): 824–827. doi:10.1126/science.283.5403.824. ISSN 0036-8075. PMID 9933161.
7. E. H. Sellards. 1916. Fossil vertebrates from Florida: A new Miocene fauna; new Pliocene species; the Pleistocene fauna. Florida State Geological Survey Annual Report 8:79-119
8. Mooser, O. (1968). "Fossil Equidae from the Middle Pliocene of the Central Plateau of Mexico". The Southwestern Naturalist. 13 (1): 1–12. doi:10.2307/3668809. ISSN 0038-4909. JSTOR 3668809.
9. W. W. Dalquest and T. J. Donovan (1973). "A new three-toed horse (Nannippus) from the late Pliocene of Scurry County, Texas". Journal of Paleontology. 47 (1): 34–45.
10. Dalquest, Walter W.; Donovan, Terrence J. (1973). "A New Three-Toed Horse (Nannippus) from the Late Pliocene of Scurry County, Texas". Journal of Paleontology. 47 (1): 34–45. ISSN 0022-3360. JSTOR 1302864.
11. Voorhies, M. R. (1974). "The Pliocene Horse Nannippus Minor in Georgia: Geologic Implications". Tulane Studies in Geology and Paleontology. 11 (2).
12. R. C. Hulbert. 1993. Journal of Vertebrate Paleontology
13. Hulbert, Richard C. Jr. (1993-09-23). "Late Miocene Nannippus (Mammalia: Perissodactyla) from Florida, with a description of the smallest hipparionine horse". Journal of Vertebrate Paleontology. 13 (3): 350–366. Bibcode:1993JVPal..13..350H. doi:10.1080/02724634.1993.10011515. ISSN 0272-4634.
14. G. G. Simpson. 1930. Bulletin of the American Museum of Natural History 59(3)

• Paleobiology database: Nannippus

v t e Perissodactyla
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Perissodactyla
Perissodactyla Incertae sedis †Ghazijhippus †Mesolambdolophus †Olbitherium †Anthracobunia Radinskya †Anthracobunidae Anthracobune Hsanotherium Jozaria Obergfellia †Cambaytheriidae Cambaytherium Kalitherium Nakusia Perissobune †Brontotheriidae See Brontotheriidae Equoidea †Cymbalophus †Hallensia †Pliolophus †Systemodon Equidae See Equidae †Palaeotheriidae †Pachynolophinae Anchilophus Bepitherium Eurohippus Hyracotherium Lijiangia Lophiohippus Lophiotherium Metanchilophus Orolophus Pachynolophus Paranchilophus Propalaeotherium Qianohippus †Palaeotheriinae Cantabrotherium Iberolophus Idiodontherium Leptolophus Mekodontherium Metaplagiolophus Palaeotherium Paraplagiolophus Plagiolophus Pseudopalaeotherium †Hyopsodontidae? Aletodon Chacomylus Dorraletes Haplaletes Haplomylus Hyopsodus Litomylus Louisina Monshyus Paratricuspiodon Paschatherium Phenacodaptes Tricuspiodon †Phenacodontidae? Tetraclaenodon †Meniscotheriinae Ectocion Meniscotherium †Phenacodontinae Almogaver Copecion Eodesmatodon Lophocion Phenacodus Tapiromorpha See below↓
Tapiromorpha Incertae sedis †Cambaylophus †Skopaiolophus †Isectolophidae Chowliia Gandheralophus Karagalax Meridiolophus Orientolophus Sastrilophus †Homogalaxinae Homogalax †Isectolophinae Isectolophus Schizolophodon †Ancylopoda See Ancylopoda Ceratomorpha Tapiroidea †Deperetellidae Bahinolophus Deperetella Irenolophus Teleolophus †Helaletidae Colodon Dilophodon Helaletes Heptodon Heteraletes Paracolodon Plesiocolopirus Thuliadanta? Vastanolophus Tapiridae †Eotapirus †Miotapirus †Nexuotapirus †Paratapirus †Plesiotapirus †Protapirus †Tapiravus Tapirus Rhinocerotoidea See below↓
Rhinocerotoidea Rhinocerotoidea sensu lato †Breviodon †Fouchia †Minchenoletes †Siamolophus †Triplopus †Yimengia †Indolophidae Indolophus Rhinocerotoidea sensu stricto †Selenaletes †Hyrachyidae Hyrachyus †Eggysodontidae Allacerops Eggysodon Guangnanodon Proeggysodon †Hyracodontidae Ardynia Ephyrachyus Epitriplophus Hyracodon Prohyracodon Rhodopagus Triplopides †Amynodontidae Amynodontopsis Armania Cadurcotherium Caenolophus Huananodon Hypsamynodon Procadurcodon Rostriamynodon Teilhardia †Amynodontinae Amynodon †Cadurcodontini Cadurcodon Lushiamynodon Sharamynodon Sianodon †Metamynodontini Gigantamynodon Megalamynodon Metamynodon Paramynodon Zaisanamynodon †Paraceratheriidae †Forstercooperiinae Forstercooperia Gobicerops Pappaceras †Paraceratheriinae Aralotherium Dzungarotherium Juxia Paraceratherium Urtinotherium Rhinocerotidae See below↓
Rhinocerotidae Rhinocerotidae †Amphicaenopus †Epiaceratherium †Guixia †Teletaceras †Uintaceras †Woodoceras †Aceratheriinae Aceratherium Acerorhinus Alicornops Aphelops Chilotheridium Chilotherium Floridaceras Galushaceras Hoploaceratherium Mesaceratherium Peraceras Persiatherium Plesiaceratherium Ronzotherium Shansirhinus Sinorhinus Subchilotherium †Elasmotheriinae Gulfoceras Victoriaceros †Diceratheriini Diceratherium Penetrigonias Skinneroceras Subhyracodon Trigonias †Elasmotheriini Bugtirhinus Caementodon Elasmotherium Hispanotherium Iranotherium Kenyatherium Meninatherium Menoceras Ninxiatherium Ningxiatherium Ougandatherium Parelasmotherium Procoelodonta Sinotherium Tongxinotherium Rhinocerotinae †Dromoceratherium †Lartetotherium †Molassitherium †Protaceratherium †Teleoceratini Aprotodon Brachydiceratherium Brachypotherium Diaceratherium Prosantorhinus Shennongtherium Teleoceras Rhinocerotina Dicerorhinini †Coelodonta Dicerorhinus †Dihoplus †Pliorhinus †Stephanorhinus Dicerotini Ceratotherium Diceros †Miodiceros †Paradiceros Rhinocerotini †Gaindatherium †Nesorhinus Rhinoceros †Rusingaceros

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