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Notopalaeognathae Temporal range: Paleocene – Holocene, 60–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Greater rhea (Rhea americana)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Infraclass:	Palaeognathae
Clade:	Notopalaeognathae Yuri et al., 2013
Clades
Rheiformes (rheas) Novaeratitae (kiwis, cassowaries, emus and †elephant birds) Dinocrypturi Sangster et al., 2022 Tinamiformes (tinamous) †Dinornithiformes (moas)

Notopalaeognathae is a clade that contains the order Rheiformes (rheas), the clade Novaeratitae (which includes the cassowaries and emus, the kiwis, and the extinct elephant birds), and the clade Dinocrypturi (comprising the tinamous and the extinct moas). Notopalaeognathae was named by Yuri et al. (2013) and defined in the PhyloCode by Sangster et al. (2022) as "the least inclusive crown clade containing Rhea americana, Tinamus major, and Apteryx australis". The exact relationships of this group, including its recently extinct members, have only recently been uncovered. The two lineages endemic to New Zealand, the kiwis and the extinct moas, are not each other's closest relatives: the moas are most closely related to the Neotropical tinamous, and the kiwis are sister to the extinct elephant birds of Madagascar, with kiwis and elephant birds together sister to the cassowaries and emu of New Guinea and Australia. The South American rheas are either sister to all other notopalaeognaths or sister to Novaeratitae. The sister group to Notopalaeognathae is Struthionidae (the ostrich family).

References

1. "Notopalaeognathae". paleobiodb.org. Retrieved 2021-09-25.
2. Van Tuinen M. (2009) Birds (Aves). In The Timetree of Life, Hedges SB, Kumar S (eds). Oxford: Oxford University Press; 409–411.
3. ^ Yuri, T.; et al. (2013). "Parsimony and model-based analyses of indels in avian nuclear genes reveal congruent and incongruent phylogenetic signals". Biology. 2 (1). MDPI: 419–44. doi:10.3390/biology2010419. PMC 4009869. PMID 24832669.
4. Sangster, George; Braun, Edward L.; Johansson, Ulf S.; Kimball, Rebecca T.; Mayr, Gerald; Suh, Alexander (2022-01-01). "Phylogenetic definitions for 25 higher-level clade names of birds" (PDF). Avian Research. 13: 100027. Bibcode:2022AvRes..1300027S. doi:10.1016/j.avrs.2022.100027. ISSN 2053-7166.
5. Phillips MJ, Gibb GC, Crimp EA, Penny D (January 2010). "Tinamous and moa flock together: mitochondrial genome sequence analysis reveals independent losses of flight among ratites". Systematic Biology. 59 (1): 90–107. doi:10.1093/sysbio/syp079. PMID 20525622.
6. Allentoft, M. E.; Rawlence, N. J. (2012-01-20). "Moa's Ark or volant ghosts of Gondwana? Insights from nineteen years of ancient DNA research on the extinct moa (Aves: Dinornithiformes) of New Zealand". Annals of Anatomy - Anatomischer Anzeiger. 194 (1): 36–51. doi:10.1016/j.aanat.2011.04.002. PMID 21596537.
7. ^ Mitchell, K. J.; Llamas, B.; Soubrier, J.; Rawlence, N. J.; Worthy, T. H.; Wood, J.; Lee, M. S. Y.; Cooper, A. (2014-05-23). "Ancient DNA reveals elephant birds and kiwis are sister taxa and clarifies ratite bird evolution". Science. 344 (6186): 898–900. Bibcode:2014Sci...344..898M. doi:10.1126/science.1251981. hdl:2328/35953. PMID 24855267. S2CID 206555952.
8. Baker, A. J.; Haddrath, O.; McPherson, J. D.; Cloutier, A. (2014). "Genomic Support for a Moa-Tinamou Clade and Adaptive Morphological Convergence in Flightless Ratites". Molecular Biology and Evolution. 31 (7): 1686–1696. doi:10.1093/molbev/msu153. PMID 24825849.
9. Hackett, S.J.; et al. (2008). "A Phylogenomic Study of Birds Reveals Their Evolutionary History". Science. 320 (5884): 1763–8. Bibcode:2008Sci...320.1763H. doi:10.1126/science.1157704. PMID 18583609. S2CID 6472805.

v t e Palaeognath genera
Kingdom: Animalia Phylum: Chordata Class: Aves Avemetatarsalia see Avemetatarsalia Theropoda see Theropoda Maniraptora see Maniraptora Palaeognathae see below↓
Palaeognathae Palaeognathae †Asteriornis? †Eremopezus? †Limenavis? †Lithornithidae Calciavis Fissuravis Lithornis Paracathartes Pseudocrypturus Struthioniformes †Remiornis? †Geranoididae? Eogeranoides Galligeranoides? Geranodornis Palaeophasianus Paragrus †Palaeotididae? Palaeotis Galligeranoides? †Eogruidae? Eogrus Sonogrus †Ergilornithidae? Amphipelargus Ergilornis Sinoergilornis Urmiornis Struthionidae †Orientornis †Pachystruthio? Struthio Notopalaeognathae see below↓
Notopalaeognathae Rheiformes †Opisthodactylidae Diogenornis? Opisthodactylus Rheidae †Heterorhea †Hinasuri †Protorhea Rhea †Dinornithiformes Dinornis Megalapteryx Emeidae Anomalopteryx Emeus Euryapteryx Pachyornis Tinamidae †Miniothura †Querandiornis †Roveretornis Tinaminae Crypturellus Nothocercus Tinamus Nothurinae Eudromia Nothoprocta Nothura Rhynchotus Taoniscus Tinamotis Novaeratitae Apterygidae Apteryx †Proapteryx †Aepyornithidae Aepyornis Mullerornis Casuariiformes †Diogenornis? Casuariidae Casuarius Dromaius †Emuarius
Ootaxa Diamantornis Incognitoolithus Medioolithus Namornis Psammornis Tsondabornis
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