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Microcleidus Temporal range: Toarcian ~182–175 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Fossil M. homalospondylus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Superorder:	†Sauropterygia
Order:	†Plesiosauria
Family:	†Microcleididae
Genus:	†Microcleidus Watson, 1909
Type species
Plesiosaurus homalospondylus (Owen, 1865)
Other species
M. macropterus (Seeley, 1865) M. tournemirensis (Sciau et al., 1990) M. melusinae Vincent et al., 2019
Synonyms
M. macropteus Plesiosaurus macropteus Seeley, 1865 Eretmosaurus macropterus Benton and Spencer, 1995 M. tournemirensis Plesiosaurus tournemirensis Sciau et al., 1990 Occitanosaurus tournemirensis Bardet et al., 1999

Microcleidus is an extinct genus of sauropterygian reptile belonging to the Plesiosauroidea. The species has 40 neck vertebrae and a short tail of 28 vertebrae. Fossils of the genus have been found in France, the Posidonia Shale in Germany and Luxembourg, and the Alum Shale Formation of England.

Description

The type species, M. homalospondylus, was the largest, measuring 5.1 m (17 ft) long and weighing 650 kg (1,430 lb). Other species were smaller: M. tournemirensis was about 4 m (13 ft) long and weighed 300 kg (660 lb), and M. melusinae was about 3 m (9.8 ft) long and weighed 120 kg (260 lb).

Classification

Species include: Microcleidus homalospondylus (Owen 1865) and Microcleidus macropterus (Seeley 1865).

Occitanosaurus tournemirensis (originally "Plesiosaurus" tournemirensis), was named by Sciau et al. in 1990, based on a nearly complete skeleton of an animal approximately 4 meters (13 ft) long. It was later found to be a species of Microcleidus.

The following cladogram follows an analysis by Ketchum & Benson, 2011.

See also

• List of plesiosaur genera
• Timeline of plesiosaur research

References

1. ^ Ketchum HF, Benson RBJ. Global interrelationships of Plesiosaur (Reptilia, Sauropterygia) and the pivotal role of taxon sampling in determining the outcome of phylogenetic analyses. Biological Reviews
2. Valentin Fischer; Nikolay G. Zverkov; Maxim S. Arkhangelsky; Ilya M. Stenshin; Ivan V. Blagovetshensky; Gleb N. Uspensky (2020). "A new elasmosaurid plesiosaurian from the Early Cretaceous of Russia marks an early attempt at neck elongation". Zoological Journal of the Linnean Society. 192 (4): 1167–1194. doi:10.1093/zoolinnean/zlaa103. hdl:2268/251614.Supplementary Information
3. Paul, Gregory S. (2022). The Princeton Field Guide to Mesozoic Sea Reptiles. Princeton University Press. pp. 108–109. doi:10.1515/9780691241456. ISBN 9780691193809. S2CID 251553177.
4. Hilary F. Ketchum; Roger B. J. Benson (2011). "A new pliosaurid (Sauropterygia, Plesiosauria) from the Oxford Clay Formation (Middle Jurassic, Callovian) of England: evidence for a gracile, longirostrine grade of Early-Middle Jurassic pliosaurids". Special Papers in Palaeontology. 86: 109–129.

Bibliography

• Bardet, Nathalie; Fernandez, Marta; Garcia-Ramos, Jose Carlos; Superbiola, Xabier Pereda; Pinuela, Laura; Ruiz-Omenaca, Jose Ignacio; and Vincent, Peggy (2008). "A juvenile plesiosaur from the Pliensbackian (Lower Jurassic) of Asturias, Spain". Journal of Vertebrate Paleontology 28(1): 258–263
• Plesiosaur.com 13 Oct 2006
• Palæos.com 13 Oct 2006

v t e Sauropterygia
Kingdom: Animalia Phylum: Chordata Class: Sauropsida Subclass: Diapsida Clade: Neodiapsida Sauropsida see Sauropsida Sauropterygia see below↓
Sauropterygia Sauropterygia Alexeyisaurus Atopodentatus Palatodonta? Pomolispondylus? Hanosaurus? Helveticosauridae? Eusaurosphargis? Helveticosaurus Saurosphargidae Hemilopas? Largocephalosaurus Prosaurosphargis Saurosphargis Sinosaurosphargis Stauromatodon? Placodontia Paraplacodus Pararcus Placodus Cyamodontoidea Cyamodus Glyphoderma Henodus Macroplacus Parahenodus Placochelys Protenodontosaurus Psephochelys Psephoderma Psephosauriscus Psephosaurus Sinocyamodus Eosauropterygia see below↓
Eosauropterygia Eosauropterygia Corosaurus Cymatosaurus Pachypleurosauria Anarosaurus Chusaurus Dactylosaurus Dawazisaurus? Diandongosaurus Dianmeisaurus Dianopachysaurus Hanosaurus? Honghesaurus Keichousaurus Luopingosaurus Majiashanosaurus Neusticosaurus Odoiporosaurus Prosantosaurus Panzhousaurus Qianxisaurus Serpianosaurus Wumengosaurus Nothosauroidea Brevicaudosaurus Hispaniasaurus? Sanchiaosaurus Simosauridae Paludidraco Simosaurus Nothosauridae Ceresiosaurus Germanosaurus Lariosaurus Metanothosaurus Nothosaurus Proneusticosaurus Silvestrosaurus Pistosauroidea Augustasaurus Bobosaurus Chinchenia Kwangsisaurus Pistosaurus Tanaisosaurus Wangosaurus Yunguisaurus Plesiosauria see below↓
Plesiosauria Plesiosauria Anningasaura Bathyspondylus Bobosaurus? Lindwurmia Sinopliosaurus Termatosaurus? Rhomaleosauridae Anningasaura? Archaeonectrus Atychodracon Avalonnectes Bishanopliosaurus? Borealonectes Eurycleidus Lindwurmia? Macroplata Maresaurus Meyerasaurus Rhomaleosaurus Sthenarosaurus Stratesaurus Thaumatodracon Yuzhoupliosaurus? Pliosauridae Anguanax Arminisaurus Attenborosaurus Cryonectes Hauffiosaurus Marmornectes Pachycostasaurus Rhaeticosaurus Thalassiodracon Thalassophonea Eardasaurus Gallardosaurus Liopleurodon Lorrainosaurus Megalneusaurus UANL-FCT-R2? Peloneustes "Pliosaurus" andrewsi Pliosaurus Simolestes Brachaucheninae Acostasaurus Brachauchenius Eiectus Kronosaurus Luskhan Makhaira Megacephalosaurus Monquirasaurus Polyptychodon Sachicasaurus Stenorhynchosaurus Plesiosauroidea Eoplesiosaurus Eretmosaurus Franconiasaurus Leurospondylus Plesiopharos Plesiopterys Plesiosaurus Westphaliasaurus Microcleididae Hydrorion Lusonectes Microcleidus Occitanosaurus Seeleyosaurus Cryptoclidia see below↓
Cryptoclidia Cryptoclididae Opallionectes Colymbosaurinae Abyssosaurus Colymbosaurus Djupedalia Pantosaurus Spitrasaurus Cryptoclidinae Cryptoclidus Kimmerosaurus Ophthalmothule Tatenectes Muraenosaurinae Muraenosaurus Picrocleidus Tricleidus? Vinialesaurus Xenopsaria Leptocleidia Hastanectes? Nichollssaura? Leptocleididae Brancasaurus Hastanectes? Leptocleidus Nichollssaura? Umoonasaurus? Vectocleidus Polycotylidae Edgarosaurus Manemergus Mauriciosaurus Thililua Palmulasaurinae Pahasapasaurus? Palmulasaurus Occultonectia Plesiopleurodon Sulcusuchus Polycotylinae Eopolycotylus Georgiasaurus Polycotylus Scalamagnus Serpentisuchops Trinacromerum Dolichorhynchia Dolichorhynchops Martinectes Unktaheela Elasmosauridae Callawayasaurus Cimoliasaurus Eromangasaurus Goniosaurus Jucha Lagenanectes Leivanectes Mauisaurus Ogmodirus Orophosaurus Plesioelasmosaurus Scanisaurus Wapuskanectes Woolungasaurus Euelasmosaurida Cardiocorax Libonectes Thalassomedon Elasmosaurinae Albertonectes Elasmosaurus Fluvionectes Hydrotherosaurus? Nakonanectes Styxosaurus Terminonatator Weddellonectia Aphrosaurus Chubutinectes Futabasaurus Fresnosaurus? Hydrotherosaurus? Kawanectes Marambionectes Morenosaurus Tuarangisaurus Vegasaurus Zarafasaura? Aristonectinae Alexandronectes Aristonectes Kaiwhekea Morturneria Wunyelfia

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