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Raeticodactylus

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(Redirected from Raeticodactylus filisurensis) Genus of raeticodactylid pterosaur from the Late Triassic

Raeticodactylus
Temporal range: Late Triassic, 213–209 Ma PreꞒ O S D C P T J K Pg N
Fossil in Bündner Naturmuseum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Order: Pterosauria
Family: Raeticodactylidae
Genus: Raeticodactylus
Stecher, 2008
Species: R. filisurensis
Binomial name
Raeticodactylus filisurensis
Stecher, 2008

Raeticodactylus is a genus of non-pterodactyloid pterosaur from the late Norian-early Rhaetian-age Upper Triassic lower Kössen Formation (about 213-209 million years ago) of the central Austroalpine of Grisons, Switzerland. It is known from holotype BNM 14524, a single disarticulated partial skeleton including an almost complete skull, found in August 2005. This genus was named and described in 2008 by its discoverer Rico Stecher; the type species is Raeticodactylus filisurensis. The specific name refers to Filisur.

Description

Life restoration

Raeticodactylus had a tall thin bony crest running along the midline of the front of the upper jaw, and a keel on the lower jaw; however, it does not seem to be closely related to Austriadactylus, the only other crested Triassic pterosaur named by the time Raeticodactylus was described. The teeth at the front of the upper jaw, in the premaxillae, were fanglike, whereas the teeth in the upper cheeks (the maxillae) had three, four, or five cusps, similar to those of Eudimorphodon. Raeticodactylus had a wingspan of about 135 centimeters (53 in), and may have been a piscivore, potentially feeding by skimming the water. However, skim-feeding has since been disproven in pterosaurs, and the related Caviramus appears to have been an omnivore.

Classification

A 2009 study by Fabio Dalla Vecchia, which analyzed the relationships within the family Campylognathoididae, found that Raeticodactylus is very closely related to Caviramus. Caviramus is known only from a partial jaw, exhibiting the same type of teeth and unique cavities in the jaw bone, and both specimens come from the same time and place (the Alplihorn Member of the Swiss Kössen Formation). Dalla Vecchia concluded that the two belong to the same genus, and possibly the same species, if the subtle differences (such as size and the presence of a crest in the Raeticodactylus specimen) are not due to sex or age.

A study by Brian Andres et al. in 2014 concluded that Raeticodactylus was the sister species of Caviramus within the family Raeticodactylidae. The following phylogenetic analysis follows the topology of Andres et al. (2014).

Eopterosauria

Preondactylus buffarinii

Austriadactylus cristatus

Peteinosaurus zambellii

Eudimorphodontoidea
Raeticodactylidae

Raeticodactylus filisurensis

Caviramus schesaplanensis

Eudimorphodontidae

Arcticodactylus cromptonellus

Carniadactylus rosenfeldi

Eudimorphodon ranzii

In 2020 however, a study upheld by Matthew G. Baron about early pterosaur interrelationships found Raeticodactylus to group with Carniadactylus, Caviramus, and the clade Austriadraconidae, which in turn were within a larger clade called Caviramidae.

See also

References

  1. ^ Stecher, Rico (2008). "A new Triassic pterosaur from Switzerland (Central Austroalpine, Grisons), Raeticodactylus filisurensis gen. et sp. nov" (PDF). Swiss Journal of Geosciences. 101: 185–201. doi:10.1007/s00015-008-1252-6. S2CID 128980861. Online First
  2. Müller R.T., Ezcurra M.D., Garcia M.S., Agnolín F.L., Stocker M.R., Novas F.E., Soares M.B., Kellner A.W.A. & Nesbitt S.J. (2023). ”New reptile shows dinosaurs and pterosaurs evolved among diverse precursors”. Nature 620(7974): p. 589–594. doi:10.1038/s41586-023-06359-z
  3. Wilton, Mark P. (2013). Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press. ISBN 0691150613.
  4. Dalla Vecchia, F.M. (2009). "Anatomy and systematics of the pterosaur Carniadactylus (gen. n.) rosenfeldi (Dalla Vecchia, 1995)". Rivista Italiana de Paleontologia e Stratigrafia. 115 (2): 159–188.
  5. Andres, B.; Clark, J.; Xu, X., 2014, "The Earliest Pterodactyloid and the Origin of the Group", Current Biology, doi:10.1016/j.cub.2014.03.030
  6. Matthew G. Baron (2020). "Testing pterosaur ingroup relationships through broader sampling of avemetatarsalian taxa and characters and a range of phylogenetic analysis techniques". PeerJ. 8: e9604. doi:10.7717/peerj.9604. PMC 7512134. PMID 33005485.
Pterosauria
Avemetatarsalia
Pterosauria
    • see below↓
Pterosauria
Pterosauria
Preondactylia
Caviramidae?
Austriadraconidae
Eopterosauria
Eudimorphodontoidea
Raeticodactylidae
Eudimorphodontidae
Eudimorphodontinae
Zambellisauria?
Macronychoptera
Dimorphodontidae
Lonchognatha?
Novialoidea
Campylognathoididae
Breviquartossa
Rhamphorhynchidae
Rhamphorhynchinae
Rhamphorhynchae
Angustinaripterini
Rhamphorhynchini
Digibrevisauria?
Scaphognathidae?
Pterodactylomorpha
    • see below↓
Campylognathoides liasicus

Scaphognathus crassirostris

Dorygnathus banthensis
Pterodactylomorpha
Pterodactylomorpha
Monofenestrata
Darwinoptera
Wukongopteridae
Wukongopterinae
Pterodactyliformes
Caelidracones
Anurognathidae
Anurognathinae
Batrachognathinae
Pterodactyloidea
Lophocratia
    • see below↓
Jeholopterus ninchengensis Kryptodrakon progenitor
Lophocratia
Archaeopterodactyloidea
Germanodactylidae
Euctenochasmatia
Ctenochasmatoidea
Gallodactylidae
Aurorazhdarchia
Aurorazhdarchidae
Ctenochasmatidae
Moganopterinae?
Gnathosaurinae
Ctenochasmatinae
Pterodaustrini
Eupterodactyloidea
Ornithocheiroidea
    • see below↓
Pterodactylus antiquus Plataleorhynchus streptorophorodon
Ornithocheiroidea
Ornithocheiroidea
Tapejaroidea
Dsungaripteridae
Dsungaripterinae
Azhdarchoidea
Neoazhdarchia
Tapejaromorpha
Thalassodromidae?
Tapejariformes
Caupedactylia?
Tapejaridae
Sinopterinae
Tapejarinae
Tapejarini
Caiuajarina
Azhdarchomorpha
Alanqidae?
Neopterodactyloidea
Chaoyangopteridae
Chaoyangopterinae
Azhdarchiformes
Azhdarchidae
Azhdarchinae
Quetzalcoatlinae
Pteranodontoidea
    • see below↓
Bakonydraco galaczi

Tupandactylus imperator

Quetzalcoatlus
Pteranodontoidea
Pteranodontoidea
Pteranodontia
Pteranodontidae
Nyctosauromorpha
Aponyctosauria
Nyctosauridae
Ornithocheiromorpha
Lonchodectidae
Lanceodontia
Lonchodraconidae
Istiodactyliformes
Mimodactylidae
Istiodactylidae
Istiodactylinae
Ornithocheiriformes
Boreopteridae
Ornithocheirae
Ornithocheiridae
Ornithocheirinae
Targaryendraconia?
Cimoliopteridae
Targaryendraconidae
Anhangueria
Hamipteridae?
Anhangueridae
Tropeognathinae?
Coloborhynchinae?
Anhanguerinae
Pteranodon longiceps

Nyctosaurus gracilis

Ludodactylus sibbicki
Taxon identifiers
Raeticodactylus filisurensis
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