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Saurichthyiformes Temporal range: Changhsingian-Aalenian PreꞒ Ꞓ O S D C P T J K Pg N
Life restoration of Saurichthys
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	†Saurichthyiformes Aldinger, 1937
Families
†Yelangichthyidae Wu et al., 2013 †Saurichthyidae Owen, 1860

Saurichthyiformes is an extinct order of ray-finned fish which existed in Asia, Africa, Australia, Europe and North America, during the late Permian to early Middle Jurassic. Saurichthyiiformes comprise two families, Saurichthyidae and Yelangichthyidae. Yelangichthyidae is monotypic, containing only the genus Yelangichthys. The gar or needlefish-like Saurichthyidae is primarily known from the genus Saurichthys. Additionally, the subgenera Saurorhynchus Costasaurichthys, Eosaurichthys, Lepidosaurichthys, and Sinosaurichthys are frequently used to group species, and are sometimes considered separate genera. Species are known from both marine end freshwater deposits. They had their highest diversity during the Early and Middle Triassic. Their phylogenetic position is uncertain, while they have often been considered members of Chondrostei, and thus related to living sturgeons and paddlefish, phylogenetic analysis of well-preserved remains has considered this relationship equivocal. They may actually belong to the stem-group of Actinopterygii, and thus not closely related to any living group of ray-finned fish.

Systematics

• Order †Saurichthyiformes Aldinger, 1937
  • Family †Saurichthyidae Owen 1860
    • Genus †Saurichthys Agassiz, 1834
      • †Saurichthys apicalis Agassiz 1834 (type species)
      • †Saurichthys breviabdominalis Maxwell et al. 2015
      • †Saurichthys curionii (Bellotti 1857)
      • †Saurichthys daubreei Firtion 1934
      • †Saurichthys dawaziensis Wu et al. 2009
      • †Saurichthys dianneae Maxwell et al. 2016
      • †Saurichthys dongusensis Minich 1992
      • †Saurichthys eximius Minikh 1982
      • †Saurichthys gigas (Woodward 1890), non von Quenstedt 1858
      • †Saurichthys gracilis (Woodward 1890)
      • †Saurichthys grignae Tintori 2013
      • †Saurichthys gypsophilus Reis 1892
      • †Saurichthys hamiltoni Stensiö 1925
      • †Saurichthys hoffmanni Werneburg et al. 2014
      • †Saurichthys huanshenensis Chou & Liu 1957
      • †Saurichthys irregularis Oertle 1928
      • †Saurichthys latifrons Eck 1865
      • †Saurichthys lepidosteoides Frech 1903
      • †Saurichthys macrocephalus (Deecke 1889)
      • †Saurichthys majiashanensis Tintori et al. 2014
      • †Saurichthys minimahleri Werneburg et al. 2014
      • †Saurichthys nepalensis Beltan & Janvier 1978
      • †Saurichthys obrutchevi Minich 1981
      • †Saurichthys orientalis Sytchevskaya 1999
      • †Saurichthys parvidens Wade 1935
      • †Saurichthys piveteaui Beltan 1968
      • †Saurichthys proximus Minich 1981
      • †Saurichthys rieppeli Maxwell et al. 2015
      • †Saurichthys sceltrichensis Renesto et al. 2021
      • †Saurichthys seefeldensis Kner 1867
      • †Saurichthys spinosa Wu et al. 2018
      • †Saurichthys stensioi Lehman 1952
      • †Saurichthys sui Fang & Wu 2022
      • †Saurichthys tenuirostris Münster 1839
      • †Saurichthys tertius Minich 1982
      • †Saurichthys ultimus Minikh 1992
      • †Saurichthys vjuschkovi Minikh 1983
      • †Saurichthys yangjuanensis Wu et al. 2015
      • †Saurichthys yunnanensis Zhang, Zhou, Lu & Bai 2010
    • Subgenus †Costasaurichthys Tintori, 2013
      • †Saurichthys (Costasaurichthys) costasquamosus Rieppel 1985 (type species)
      • †Saurichthys (Costasaurichthys) paucitrichus Rieppel 1992
    • Subgenus †Eosaurichthys Liu & Wei, 1988
      • †Saurichthys (Eosaurichthys) chaoi (Liu and Wei 1988) (type species)
      • †Saurichthys (Eosaurichthys) madagascariensis Piveteau 1945
    • Subgenus †Lepidosaurichthys Tintori, 2013
      • †Saurichthys (Lepidosaurichthys) dayi (Raymond 1925) (type species)
      • †Saurichthys (Lepidosaurichthys) elongatus Stensiö 1925
      • †Saurichthys (Lepidosaurichthys) ornatus Stensiö 1925
      • †Saurichthys (Lepidosaurichthys) toxolepis Mutter et al. 2008
      • †Saurichthys (Lepidosaurichthys) wimani (Woodward 1912)
    • Subgenus †Sinosaurichthys Wu et al., 2010
      • †Saurichthys (Sinosaurichthys) longipectoralis Wu et al. 2009 (type species)
      • †Saurichthys (Sinosaurichthys) longimedialis Wu et al. 2009
      • †Saurichthys (Sinosaurichthys) minuta Wu et al. 2009
    • Genus †Saurorhynchus Reis, 1892
      • †Saurichthys (Saurorhynchus) calcaratus Griffith 1977
      • †Saurichthys (Saurorhynchus) deperditus (Costa 1862)
      • †Saurichthys (Saurorhynchus) striolatus Bronn 1858
      • †Saurorhynchus acutus (Agassiz 1844) (type species)
      • †Saurorhynchus anningae Maxwell & Stumpf 2017
      • †Saurorhynchus brevirostris (Woodward 1895)
      • †Saurorhynchus hauffi Maxwell & Stumpf 2017
  • Family †Yelangichthyidae Wu et al., 2013
    • Genus †Yelangichthys Wu et al., 2013
      • †Yelangichthys macrocephalus Wu et al., 2013 (type species)

Phylogeny

Modified from Maxwell et al. (also see Renesto et al., Argyriou et al. and Gardiner et al.)

Timeline of genera

References

1. ^ Wu Feixiang; Sun Yuanlin; Xu Guanghui; Hao Weicheng; Jiang Dayong; Sun Zuoyu (2010). "New saurichthyid fishes (Actinopterygii) from the Middle Triassic (Pelsonian, Anisian) of southwestern China" (PDF). Acta Palaeontologica Polonica. 56. doi:10.4202/app.2010.0007. S2CID 54877125.
2. Stensiö, E. (1925). "Triassic Fishes from Spitzebergen, Part II". Kungliga Svenska Vetenskapsakademiens Handlinga. 2: 1–126.
3. Cartanyà, J., ed. (1999). An overview of the Middle Triassic actinopterygians from Alcover, Mont-ral and El Pinetell (Catalonia, Spain). In: G. Arratia and H. P. Schultze (eds.) Mesozoic Fishes 2—Systematics and Fossil Record: Verlag Dr. F. Pfeil, München. pp. 535–551.{{cite book}}: CS1 maint: location (link)
4. Maxwell, E.E. (2016): First Middle Jurassic record of Saurichthyidae (Actinopterygii). Paläontologische Zeitschrift 90:287-291 https://doi.org/10.1007/s12542-015-0281-5
5. Romano C., Kogan I., Jenks J., Jerjen I., Brinkmann W. (2012). "Saurichthys and other fossil fishes from the late Smithian (Early Triassic) of Bear Lake County (Idaho, USA), with a discussion of saurichthyid palaeogeography and evolution" (PDF). Bulletin of Geosciences. 87: 543–570. doi:10.3140/bull.geosci.1337.{{cite journal}}: CS1 maint: multiple names: authors list (link)
6. ^ Argyriou, Thodoris; Giles, Sam; Friedman, Matt; Romano, Carlo; Kogan, Ilja; Sánchez-Villagra, Marcelo R. (December 2018). "Internal cranial anatomy of Early Triassic species of †Saurichthys (Actinopterygii: †Saurichthyiformes): implications for the phylogenetic placement of †saurichthyiforms". BMC Evolutionary Biology. 18 (1): 161. Bibcode:2018BMCEE..18..161A. doi:10.1186/s12862-018-1264-4. ISSN 1471-2148. PMC 6211452. PMID 30382811.
7. Haaramo, Mikko (2007). "†Saurichthyiformes". Mikko's Phylogeny Archive. Retrieved 30 December 2016.
8. Nelson, Joseph S.; Grande, Terry C.; Wilson, Mark V. H. (2016). Fishes of the World (5th ed.). John Wiley & Sons. ISBN 9781118342336.
9. van der Laan, Richard (2016). "Family-group names of fossil fishes" (PDF). Retrieved 2023-12-06.
10. ^ Discovery of two new species of primitive fishes
11. Silvio Renesto; Fabio Magnani; Rudolf Stockar (2021). "A new species of Saurichthys (Actinopterygii: Saurichtydae) from the Middle Triassic of Monte San Giorgio". Rivista Italiana di Paleontologia e Stratigrafia. 127 (1): 49–71. doi:10.13130/2039-4942/15143.
12. Fei-Xiang Wu; Yuan-Lin Sun; Geng-Yu Fang (2018). "A new species of Saurichthys from the Middle Triassic (Anisian) of southwestern China". Vertebrata PalAsiatica. 56 (4): 273–294. doi:10.19615/j.cnki.1000-3118.171023.
13. Fang, G., Wu, F.X. (2022). "The predatory fish Saurichthys reflects a complex underwater ecosystem of the Late Triassic Junggar Basin, Xinjiang, China". Historical Biology. 35 (8): 1–11. doi:10.1080/08912963.2022.2098023. S2CID 250567176.{{cite journal}}: CS1 maint: multiple names: authors list (link)
14. Fossilworks: Saurichthys Agassiz 1843
15. ^ Tintori, Andrea (2013). "A new species of Saurichthys (Actinopterygii) from the Middle Triassic (early Ladinian) of the northern Grigna Mountain (Lombardy, Italy)". Rivista Italiana di Paleontologia e Stratigrafia. 119 (3): 287–302. doi:10.13130/2039-4942/6041.
16. Kogan , Romano (2016). "Redescription of Saurichthys madagascariensis Piveteau, 1945 (Actinopterygii, Early Triassic), with implications for the early saurichthyid morphotype". Journal of Vertebrate Paleontology. 36 (4): e1151886. Bibcode:2016JVPal..36E1886K. doi:10.1080/02724634.2016.1151886. S2CID 87234436.
17. Kogan, Ilja; Romano, Carlo (2016). "A new postcranium of Saurichthys from the Early Triassic of Spitsbergen" (PDF). Freiberger Forschungshefte C (Paläontologie, Stratigraphie, Fazies 23). 550: 205–221. ISBN 9783860125526.
18. Kogan, I. (2016): Acidorhynchus Stensiö, 1925 or Saurorhynchus Reis, 1892: how to call the Jurassic saurichthyid? Neues Jahrbuch für Geologie und Paläontologie-Abhandlungen 279:123-126 https://doi.org/10.1127/njgpa/2016/0545
19. ^ Ilja Kogan; Andrea Tintori; Martin Licht (2020). "Locomotor function of scales and axial skeleton in Middle-Late Triassic species of Saurichthys (Actinopterygii". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2): 475–498. doi:10.13130/2039-4942/13551. S2CID 244984857.
20. Maxwell, E.E.; Romano, C. & Wu, F.-X. (2021). "Regional disparity in the axial skeleton of Saurichthyidae and implications for axial regionalization in non-teleostean actinopterygians". Journal of Zoology. 315: 29–41. doi:10.1111/jzo.12878. ISSN 0952-8369.
21. Silvio Renesto; Fabio Magnani; Rudolf Stockar (2021). "A new species of Saurichthys (Actinopterygii: Saurichtydae) from the Middle Triassic of Monte San Giorgio". Rivista Italiana di Paleontologia e Stratigrafia. 127 (1): 49–71. doi:10.13130/2039-4942/15143.
22. Gardiner, B.G.; Schaeffer, B. & Masserie, J.A. (2005). "A review of the lower actinopterygian phylogeny". Zoological Journal of the Linnean Society. 144 (4): 511–525. doi:10.1111/j.1096-3642.2005.00181.x.

Additional references

• Sepkoski, Jack (2002). "A compendium of fossil marine animal genera". Bulletins of American Paleontology. 364: 560. Retrieved 2011-05-17.

v t e Actinopterygian genera
Kingdom: Animalia Phylum: Chordata Subphylum: Vertebrata Infraphylum: Gnathostomata Superclass: Osteichthyes Gnathostomata see Gnathostomata Actinopterygii see below↓
Actinopterygii Actinopterygii †Austelliscus †Cheirolepis †Cuneognathus †Dialipina? †Kentuckia †Krasnoyarichthys †Ligulalepis? †Limnomis †Meemannia †Moythomasia †Naxilepis? †Orvikuina? †Osorioichthys †Palaeoneiros †Pickeringius †Raynerius †Stegotrachelus †Tegeolepis †Terenolepis? †Howqualepididae Donnrosenia Howqualepis †Mimiidae Gogosardina Mimipiscis Post-Devonian taxa see below↓
Post-Devonian taxa Post-Devonian taxa †Adroichthys †Aetheretmon †Agecephalichthys †Amblypterina †Ambodipia †Aneurolepis †Angatubichthys †Apateolepis †Arratiaichthys †Asarotus †Australichthys †Avonichthys †Beagiascus †Belichthys †Bethesdaichthys †Birgeria †Bishofia †Bluefieldius †Blourugia †Boreolepis †Brachydegma †Brachypareion †Brazilichthys †Burguklia †Carbonilepis †Caruichthys †Centrolepis †Charleuxia †Chichia †Coccocephalus †Concentrilepis †Cornuboniscus †Cosmolepis †Cosmoptychius †Cretolepis †Cylindrichthys †Daqingshaniscus †Diplospondichthys †Disichthys †Dmitrevichthys †Drydenius †Duwaichthys †Dwykia †Elpisopholis †Ganolepis †Gardinerichthys †Gardinerpiscis †Gondwanichthys †Grandfieldia †Guaymayenia †Guntherichthys †Gyrolepidoides †Huanghelepis †Hyllingea †Illiniichthys †Inichthys †Irajapintoseidon †Itararichthys †Kalops †Kansasiella †Karaunguria †Kazanichthys †Kelliaichthys †Kenderlichthys †Khantausia †Kompasia †Kutaichthys †Lambeia †Lanarkichthys †Lawrenciella †Leptogenichthys †Letovichthys †Lineagruan †Luederia †Mansfieldiscus †Megapteriscus †Melanecta †Mesembrioniscus †Mesonichthys †Mesopoma †Ministrella †Monesedeiphus †Muensterichthys †Myriolepis †Nematoptychius †Neuburgella †Notacmon †Novogonatodus †Nozamichthys †Oxypteriscus †Pacorichthys †Palaeobergia †Paphosiscus †Paradrydenius †Paralogoniscus †Peleichthys †Piratata †Phanerorhynchus †Phanerosteon †Praolepis †Protamblyptera †Pseudohaplolepis †Psilichthys †Pteroniscus †Rastrolepis †Reticulolepis †Roslerichthys †Rubidus †Samarichthys †Santosichthys †Sceletophorus †Schizolepis †Senekichthys †Sinkiangichthys †Sinoniscus †Sixtelia †Sphaerolepis †Spinofacia †Strepheoschema †Sundayichthys †Tanypterichthys †Tanyrhinichthys †Tienshaniscus †Trachelacanthus †Trawdenia †Turgoniscus †Ufalepis †Urosthenes †Usclasichthys? †Uydenia †Uydenichthys †Varialepis †Wayaobulepis †Westlepis †Whiteichthys †Willomorichthys †Woodichthys †Xingshikous †Yogoniscus †Yaomoshania †Zhongweilepis †Acrolepidae Acrolepis Acropholis Challaia Kasanilepis Namaichthys Plegmolepis Progyrolepis? Watsonichthys †Aeduellidae Aeduella Bourbonnella Decazella Neslovicella Platysella Puertollanichthys Schafeevus Spinarichthys Westollia †Aesopichthyidae Aesopichthys Proceramala †Amblypteridae Amblypterus Daphnaechelus Korutichthys? Lawnia Paramblypterus Tchekardichthys Tholonosteon Tholonotus †Bobasatraniiformes Dorypterus Bobasatraniidae Bobasatrania Ebenaqua Ecrinesomus? †Canobiidae Canobius Schaefferius †Discordichthyidae Discordichthys Geryonichthys Mamulichthys Mutovinia †Eigiliidae Eigilia Strelnia †Elonichthyidae Alilepis Elonichthys Palaeostrugia? Meisenheimichthys Rhabdolepis? †Eurynotiformes Mesolepis Paramesolepis Proteurynotus Wardichthys Amphicentridae Amphicentrum Cheirodopsis Eurynotus Styracopteridae Benedenius Fouldenia Styracopterus †Eurynotoidiformes Adzvalepis Alvinichthys Eurynotoides Isadia Kichkassia Lapkosubia Minicholepis Vologdinia †Gonatodidae Gonatodus Pseudogonatodus Usolia †Guildayichthyidae Discoserra Guildayichthys †Haplolepidae Andrewsolepis Blairolepis Braccohaplolepis Haplolepis Millerolepis Parahaplolepis Protohaplolepis Pyritocephalus †Igornichthyidae Commentrya Igornella Igornichthys Setlikia †Palaeoniscidae Atherstonia Cteniolepidotrichia Dicellopyge Ferganiscus Gyrolepis Palaeoniscinotus Palaeoniscum Palaeothrissum Shuniscus Triassodus Weixiniscus †Platysomidae Aestuarichthys Dorsolepis Kargalichthys Ningxiaplatysomus Paranaichthys Platysomus Schaefferichthys Sinoplatysomus Tompoichthys †Ptycholepidae Acrorhabdus Ardoreosomus Boreosomus Chungkingichthys Ptycholepis Yuchoulepis †Pygopteridae Briveichthys Progyrolepis? Pygopterus Zaborichthys †Rhadinichthyidae Cycloptychius Cyranorhis Mentzichthys Pteronisculus? Rhadinichthys Wendyichthys †Saurichthyiformes Yelangichthys Saurichthyidae Costasaurichthys Eosaurichthys Lepidosaurichthys Saurichthys Saurorhynchus Sinosaurichthys †Tarrasiidae Paratarrasius Tarrasius †Turseoidae Pteronisculus? Turfania? Turseodus †Uighuroniscidae Indaginilepis Uighuroniscus Cladistia see Cladistia includes bichirs Actinopteri Chondrostei see Chondrostei includes sturgeons and paddlefish Neopterygii see Neopterygii

• Saurichthyiformes
• Prehistoric ray-finned fish orders
• Jurassic bony fish
• Permian bony fish
• Triassic bony fish
• Lopingian first appearances
• Early Jurassic extinctions