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Scotch argus
Male
Underside
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Family:	Nymphalidae
Genus:	Erebia
Species:	E. aethiops
Binomial name
Erebia aethiops (Esper, 1777)
Synonyms
List Papilio ocellatus Goeze, 1779 Papilio blandina Fabricius, 1787 Erebia medusa Borkhausen, 1788 Papilio medea Hubner, 1799-1800 Erebia alcyone Stewart, 1817 Erebia neoridas Freyer, 1833 Erebia flavescens Tutt, 1896 Erebia obsoleta Tutt, 1896 Erebia ochracea Tutt, 1896 Erebia freyeri Oberthur, 1909 Erebia fritschi Oberthur, 1909 Erebia cosinida Fruhstorfer, 1917 Erebia depupillata Steiner, 1918 Erebia magna Rocci, 1923 Erebia biocellata Hartig, 1924 Erebia reducta Hartig, 1924 Erebia albofasciata Osthelder, 1925 Erebia perfumosa Dannehl, 1925 Erebia purpurea Sibille, 1927 Erebia infasciata Warren, 1936 Erebia pupillifer a Kolar, 1938 Erebia amplevittata Holtz, 1939 Erebia inocellata Owen, 1952 Erebia binotata Popescu-Gorj, 1955 Erebia demarginata Popescu-Gorj, 1955 Erebia enucleata Popescu-Gorj, 1955 Erebia mesorubria Popescu-Gorj, 1955 Erebia jigodini Popescu-Gorj, 1955 Erebia magdalena Krzywicki, 1963

The Scotch argus (Erebia aethiops) is a butterfly of the family Nymphalidae. In spite of its English name argus, it is not a close relation of the brown argus nor the northern brown argus.

Taxonomy

Subspecies include:

• Erebia aethiops ssp. aethifps (Esper, 1777) (Europe)
• Erebia aethiops ssp. caledonia (Verity, 1911 (Scotland)
• Erebia aethiops ssp. deprezzivaga Verity, 1935
• Erebia aethiops ssp. fogarasica Warren, 1931
• Erebia aethiops ssp. isouata Goltz, 1939 (Shansi)

Distribution and habitat

Erebia aethiops is widespread in the Palearctic realm, including most of Europe, the Alps and Apennines, Balkans, Asia Minor, the Urals, and the Caucasus. English colonies are found at Arnside Knott and Smardale Gill both in Cumbria. This species is found in most of northern Scotland, and in western areas in south Scotland. In Europe and Russia, these butterflies prefer the edge of pine forests and tall damp grassland in hills and mountains up to 2400 m.

Description

Erebia aethiops has a wingspan of 42–46 mm. Antennae are clavate (club shaped). The background colour of the wings varies from dark brown to black brown, with reddish-yellow bands, black eyespots with white pupils and greyish wing fringes, weakly chequered in the females. On the forewing there are three or four eyespots, usually two apical plus a third detached, while on the hindwings there are four or five.

The underside of the hindwings shows a whitish or cream banding but there is a slightly marked sexual dimorphism. In Scotland, the subspecies caledonia has a narrower reddish-yellow band and never contains more than three small eyespots. Three black spots are present on each hindwing. The egg is oval shaped, with 20 to 28 longitudinal ribs. The mature caterpillar is yellow brown, with a brown head. The pupae are yellow brown and relatively short and compact, reaching a length of 12.5–124.5 millimetres (0.49–4.90 in).

This species is rather similar to Erebia ligea, but the fringes on the edges of the upper side of the wings are more light grey, while in E. ligea they are white.Wheeler (1903) gives a short description of both species.

• Erebia aethiops ♂
• Erebia aethiops ♂ △
• Erebia aethiops ♀
• Erebia aethiops ♀ △

Biology

This species is univoltine. Adults fly from June to August. The eggs are laid singly on the leaves of grass. The larvae hatch after two weeks. The hibernating larva is reported to feed on Agrostis, Dactylis, Poa annua, Bromus erectus and other grasses. In Scotland the main host plants are purple moor-grass (Molinia caerulea) and blue moor-grass (Sesleria caerulea).

Gallery

• Egg
• Caterpillar
• Pupa
• E. a. caledonia
  Argyll, Scotland

References

1. Lepiforum.de
2. ^ "Erebia Dalman, 1816" at Markku Savela's Lepidoptera and Some Other Life Forms
3. ^ Steven Cheshire Argus British Butterflies
4. Fauna europaea
5. Matt Rowling Euro Butterflies
6. Wheeler, George.The butterflies of Switzerland and the alps of central Europe. London: Elliot Stock, 1903.

External links

• Moths and Butterflies of Europe and North Africa
• www.schmetterling-raupe.de

• Erebia
• Butterflies of Europe
• Butterflies of Asia
• Butterflies described in 1777
• Taxa named by Eugenius Johann Christoph Esper