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Late twenties Croydon flatmates Mark and Jeremy are introduced. Mark is a history-loving office worker who has never been in a relationship and is attracted to his colleague Sophie. Jez is a lazy, unemployed slacker and aspiring musician who lives in Mark's high-rise flat and leeches off him after breaking up with Big Suze and leaving her "love shack", where the couple cohabited for 18 months.

The two men are competing to try to become their unhappily married next-door neighbour Toni's fuck buddy. She hosts a party, at which Mark fails in his attempt to seduce her while talking to her about the Battle of Stalingrad. Meanwhile, Jez meets her sister, whom he erroneously believes has leukaemia because she has another sister who has the disease. After this misunderstanding causes an argument, Mark and Jez are told by Toni's neighbour, Barry, to leave.

During the next day, Mark picks up a metal pole that had been discarded in the street. He uses it to chase away a group of children who had been harassing and kicking him in the street during the previous night. Sophie is horrified to see this through the window of the bus that she is on.

Mark tells Sophie that he likes her and ineptly tries to woo her. She is in a relationship with their colleague Jeff. Jez and Super Hans' plans to form a band are halted by disagreements between them, including what the band's name should be. Toni tries to involve Jez in pyramid selling, and he tries to involve Mark. Mark tries to get Jez a job at his workplace, JLB Credit, but Jez is determined not to become employed, so he deliberately gives a bad impression of himself at his interview and tries to involve the interviewer in pyramid selling.

Mark is shopping at the supermarket at 8pm on a Friday, when Jez walks in and invites him to a party that is happening that evening. At the party, Mark strikes up a rapport with sixth-form college student goth, Valerie. When all three go to a bowling alley, Jez brings Toni. At the alley, they encounter Sophie as well as Jeff, who is on a date with her. Mark has sex with Valerie at his flat, but stops when she starts to strangle him, which she does because her ex enjoyed it. Jez has sex with Toni at her flat. She makes a lot of noise so that Tony, who is in another room in the flat, hears them. Toni and Tony are separating, and argue during her sex with Jez about which of their possessions he may take with him.

Mark questions his sexual orientation when he becomes infatuated with his boss, charismatic loan manager Alan Johnson, who asks Mark to join him on a business venture. Mark watches interracial gay porn in order to try to work out if he is sexually attracted to men. Jez and Johnson quickly take a disliking to each other. Mark drives Johnson's BMW without his permission or a licence, after Jez encourages him to, and drives into a skip. At the flat, Jez tells Johnson that Mark is gay and is sexually attracted to Johnson. Mark admits to Johnson that he has feelings for him, but that he is 85% sure that he is straight.

Jez tries to remember "the bad thing" that he did with Hans whilst they were on a drug binge together, which he eventually remembers was fellatio.

Tony has moved back in with Toni; Jez is jealous and tells her that he loves her. Mark gets into Sophie's email account by guessing her password. He reads her emails, which include her thoughts about him and Jeff. Mark wants Sophie to find him appealing so that he can poach her from Jeff, so Mark and Jez to go to Sophie's dance class ("Rainbow Rhythms"). Sophie dances with a man, and Mark quickly becomes unpopular because of his lack of enthusiasm and for rejecting another man who tried to dance with him. Jez dances with Nancy, a beautiful young American Christian hippie, who then dances with fellow class member Gwyn. Jez and Nancy begin a taboo-breaking relationship, involving anal sex and Jez blacking up. Mark, Jez, Nancy, Sophie, Toni, Tony and Gwyn go to the pub, where Nancy surprises Jez when she describes their relationship as open. At Nancy's suggestion, she, Mark, Sophie and Jez visit Gwyn's countryside house in Tollesbury, Essex, rather than their previous plan of visiting a stone circle. Whilst at Gwyn's, they skinny dip in a nearby lake, then play spin the bottle at his house, which leads to Nancy and Gwyn having sex together. At the office, Sophie is angry when she sees Mark reading her emails; she then walks off.

At work, Mark asks two security guards to show him on the CCTV what his colleagues are doing. He sees Sophie and Jeff kissing. Mark is filmed drinking beer in a park in a television report about nuisance drinkers while he and Jez are spying on Sophie and Jeff. After seeing Mark drinking on TV, Johnson makes Mark go to an alcoholics support group, where Mark has to tell the group that he is an alcoholic, then give details of his drinking. After Johnson sees Mark drinking beer in a pub, he ends Mark's chances of attending the business trip in Aberdeen with Sophie. Back at the office, Mark goads Jeff into hitting him, thinking it will be recorded on CCTV and cause action to be taken against Jeff. Mark is disappointed to find out from security that it was not recorded.

Mark accompanies Jez, who is filling in with Super Hans in a band, to Dartmouth, Devon, where the band is playing. Mark is hoping to find shoe shop assistant April (played by Catherine Shepherd) whom he had recently first met in the shop. She is a student at Dartmouth University (where he and Jez met when they were students together there) and has the "magical combo of beauty and low self-esteem". Mark goes to the university, and, pretending to be a mature student, is welcomed into April's Ancient History tutorial that is led by Professor MacLeish (Peter Capaldi). Jez misses out on the frontman spot when he is delayed by being caught by a shop's manager, Mr Rashid, whilst trying to shoplift a chocolate bar. Rashid holds Jez and Mark in his store room, until they climb out of the window. Mark and April go to a get-together at the professor's house. Jez arrives, and the professor immediately takes a disliking to him. April invites Mark back to her room on campus; they kiss, but she rejects his suggestion of sex.

Sophie tries to set Mark up with her friend Karen, but Mark still pines after Sophie, even more so when Sophie tells him that she has broken up with Jeff permanently. Nancy's visa has expired. She asks Jez to marry her so that she can stay in the UK, which he immediately and enthusiastically agrees to. Nancy leaves the wedding reception to attend a job interview. Mark gives a speech. After the reception, Tony leaves Toni, who invites Jez to hers and succeeds in persuading him to have sex with her. When Jez tells Nancy about his cheating, Nancy angrily leaves him.

Jez tries to break up with his girlfriend Michelle (played by Shelley Blond). She persuades him to remain in a relationship with her when she arranges a threesome with an acquaintance of hers, Vicky, on the condition that Michelle pegs him afterwards. Big Suze returns with a new boyfriend, muscular former monk Stu. Jez resents Stu because he wants Suze himself, and he is sexually attracted to Stu. Suze becomes Michelle's lodger.

Mark experiences a reduction in self-confidence after he is mugged in a subway by two youths. Mark and Sophie are starting a relationship. Mark buys a folding knife, which Sophie is horrified to find he is carrying whilst on their second date.

Mark and Jez's Canadian friend from university, Merry, (Meredith MacNeill) arrives at their flat and stays the night. Her mother died recently and she is suffering a manic episode. She sexually propositions Mark, but he declines her advances. She gives away to Jez and Hans the disused pub which she owns. However, Jez disagrees with Hans' proposed new name for the pub – Free the Paedos. Mark phones the NHS to have her sectioned shortly afterwards.

Mark is due to attend a business trip in Frankfurt, so Jez invites Suze to a magic mushroom party at the flat. Sick with gastric flu, Mark comes home to find Hans having sex with a woman in his bed. Hans breaks the bathroom door off its hinges when he has difficulty opening it. Mark hires a carpenter, Andy, to repair the door. Andy leaves before finishing the job, and breaking the door off its hinges again when Jez tells him to finish it before he leaves. Mark is drugged and locked in his room by Jez, who hopes to seduce Big Suze. A comedy of errors ensues, with Jez trying to keep Big Suze from noticing Mark, whilst Mark is desperate to break out of his room to go to the toilet. Mark phones Johnson, who comes round and tells Jez to release Mark. Johnson sees that Mark is ill, so he removes Mark from a project at work. Everyone except Mark and Jez leave the flat.

Mark's sister Sarah is visiting after splitting from her husband, Simon. She starts a fling with Jez on her first night at the flat, despite Mark trying to discourage her from doing so by pointing out that Jez is married, has chlamydia and is in love with Suze. Suze is staying at the flat while asbestos is removed from her flat. Mark becomes attracted to her, but tries to hide that from Jez. Jez juggles pretending to be in love with Sarah whilst simultaneously attempting to worm his way back into the affections of Suze, who has told him that she has split up with Stu. Sarah wants to stay at the flat for a few months, but then changes her mind and goes back to Simon. Jez is delighted by that, because he just wanted a fling. In addition, he tired of her because she talks too much, is emotionally demanding and he finds her similarities with Mark off-putting during sex. Jez tells Suze, in front of Mark, that Mark loves her. Mark denies it. Sarah briefly returns, suspecting that Simon is having an affair. Jez is horrified by Sarah's return, so he walks out of the flat and asks Mark to tell her that their fling is over. Mark does so, and Sarah goes home.

Jez is called up for jury service. At first, he thinks it will be a black kid up on a petty theft case, and that he will vote not guilty. He finds out it is a young white woman, Carla, who is being tried for credit card fraud and initially decides to vote guilty. While the court case is ongoing Jez and Carla meet in a café and he is attracted to her, which makes him decide to vote not guilty. Jez arranges to meet Carla again, despite Mark's warning that it will be contempt of court. Sophie visits the flat with a friend. She and her friend want to go out, and Mark unsuccessfully tries to persuade Sophie to stay in. Mark, Jez, Sophie, her friends and Carla go to a gay nightclub, where most of them take drugs. The group then go to the flat, but Mark finds Sophie's friends very disagreeable and tells them to leave. Jez finds out that Carla is a habitual criminal, that he does not want her in his life, so he changes his vote to guilty.

Jez is angry with Mark for wearing an identical hoodie to him.

Mark's doubts over his engagement to Sophie continue whilst he and Jez visit Sophie's unhappily married parents in the countryside for her birthday. Jez is angry with Mark when he realises that he was only brought along as company for Sophie's annoying, needy, younger brother Jamie, who has no friends. Sophie's father, Ian, takes Mark and Jez hunting with shotguns. Jamie tags along, but Ian does not allow him to have a gun. Mark shoots a bird down, and is told by Ian to wring its neck to kill it. Mark pulls its head off, spurting blood over himself. Ian and Mark have an uncomfortable conversation in a local pub. Mark tells Jez that he has decided to call the wedding off. Sophie's mother, Penny, seduces Jez, which Mark is horrified about when he finds out about it. Ian overhears Mark tell Jez of his lack of love for Sophie. Ian makes Mark promise to cancel the wedding, and then takes his revenge on Dan, with whom Ian believes Penny has been unfaithful, by burning down his barn by throwing a petrol bomb at it. Mark decides that he will marry Sophie when she is given her late grandmother's cottage as a birthday present. Ian tries to make Mark tell the truth about not loving Sophie, but he refuses to, and no-one believes Ian when he says it. Mark and Jez then tell the family that it was Ian who burned the barn down. Sophie stays at the house, as Ian leaves in disgrace to visit his sister. Penny invites Jez to visit again, but he decides against it. Jamie unsuccessfully tries to get Jez and Mark to take him to London, as Jez drives away with Mark in the passenger seat.

Seeing her in the flat, Johnson offers Jez £530 to sleep with Big Suze. When Jez proposes this deal to her, she rejects that idea and walks away whilst upset. When Jez goes to Suze's house, he finds that she is now sexually involved with Johnson.

Johnson puts Mark in charge of formulating Project Zeus. It is to be departmental merger of Marketing and Sales, in which the other staff are Jeff, Sophie, Gerrard, Lisa and Katie. He is angry with them for not making progress. Mark, Sophie, their colleague Gerrard and Johnson go to Kettering for a JLB conference. Suze accompanies Johnson and Jez turns up to pursue Suze. Mark, Gerrard, Johnson and Big Suze go to a lap dancing club, where Suze pays for a lap dance for Mark. At the hotel, Sophie tells Jez that she has only had sex with four men in her life. This surprises Jez, who has had sex with more men than that, despite the large majority of his sexual partners having been women. After insulting his team for failing to come up with a plan, Mark realises that he has nothing to present to the JLB board. Mark decides to take Jez's advice and run away, but then bumps into Sophie. Upon Sophie's advice, he returns to face the truth. He tells the attendees that integrating Sales and Marketing does not work and recommends JLB Credit move into humanitarian work. He falsely claims that he is suffering from brain cancer and will be dead within a month. Johnson confirms that.

In order to avoid spending time with Sophie, Mark joins a gym. Whilst there, he is assigned a tall, handsome, athletic young man, Matt, as his personal trainer. Mark tells him that he is marrying, but is not sure he wants to. Mark discovers that Jez's estranged wife Nancy is working at the same gym. She tells Mark that she has been using crystal meth and assistant managed the IMAX Bradford for 18 months. He tells Jez about Nancy, and Jez then becomes desperate to get back together with her. Jez takes a job as a cleaner at the gym in order to try to get back with her. He is disappointed when she tells him that she does not want to get back with him and that she wants to have sex with Matt. Nancy tells Jez they can be friends and encourages him to date their Polish colleague Eva. He agrees to Nancy's suggestion of a cinema double date: her and Matt; him and Eva. He tries to sabotage things between Nancy and Matt by telling him that Nancy hates him and telling Nancy that Matt only wants casual sex. Jez traps Eva in a flotation tank in order to try to get her out of the way, but Nancy hears her and releases her. Sophie thinks she and Mark spend too little time together since he joined the gym. She decides to join the gym as well, in order to spend more time with her. Mark is horrified at that, because Matt will likely tell Sophie how Mark feels about her. He and Jez decide to try to get him fired so they can both be happy. They succeed after falsely claiming to the gym's manager that Matt defecated in the swimming pool and touched Mark's penis whilst massaging Mark. He is sacked, but Mark and Jez later see him during a judo class, which he is participating in. Mark and Jez try to protect themselves by using Sophie and Nancy as human shields.

Sophie has arranged for her and Mark to attend couples counselling, costing him a pound per minute. He is horrified when she tells the counsellor that he is not satisfying her sexually and that he ejaculates far too soon.

It is Mark's wedding day, but he is torn about whether he should go through with it or not. Jez tries to discourage him, but Mark decides to continue. Nancy, who has stayed at the flat overnight, says that she will not attend, because she is meeting her father, who is visiting from the US. Super Hans vomits in Mark's hat and shoes, having taken too many drugs the night before. Mark tries to avoid the wedding by proposing to a barista, then trying to get run over then insulting the driver in a failed attempt to provoke him into beating him up. Nancy phones from the Heathrow Hilton to tell Jez to say that she has argued with her father and that she wants to come to the wedding afterall. Soon after, Jez receives a text telling him that Super Hans is giving Nancy a lift from the Heathrow Hilton on a friend's moped instead. Jez tells Mark that he snogged Sophie at the JLB conference, which Mark tries to use as a reason to cancel the wedding. He tells Sophie's mother Penny that he has to see Sophie about her infidelity. Penny refuses to let Mark in, telling him that it is bad luck for the groom to see the bride on the wedding day prior to the service and that a drunken snog is insignificant. At the church, Mark hides upstairs whilst he tries to make up his mind about whether or not to marry Sophie. Jez wants to go to the toilet, but Mark refuses to let him do so. He then wets himself; his urine goes through the floorboards and leaks through to the ground floor where Sophie and the wedding guests are gathered. Mark and Sophie force themselves to go through with the wedding however both end up crying during the ceremony. Whilst being driven to the reception, Sophie becomes even more distraught and flees the car telling Mark he's "horrible" and that she wants an annulment. Jez sees Hans and Nancy embracing each other on a bench outside the church, then gets into the car with Mark.

Sophie's father, Ian, and her cousin Barney visit briefly, but Jez keeps the chain on the door and does not let them in. Barney tells Jez that she is going back to work tomorrow — three weeks earlier than she said she would — and gives Jez some music CDs he has worked on.

Mark tries to deflect the bad image that has arisen around him since the marriage failed, but the only ones who take Mark's side are Johnson (who suspends Sophie when she goes too far) and a female IT worker called Dobby (played by Isy Suttie), whom he meets in the staff canteen and who moves between different branches of JLB. She gently dry humps Mark in the stationery cupboard; he ejaculates and leaves. Mark tries to resist her because he is scared that he will be treated worse by his colleagues if he starts a new relationship at work.

After listening to the CDs, Jez and Hans decide to form a band with the talented but naïve Barney, so that they can get him to play the music while they make all the money. The band get a gig at a venue called The Fuck Bunker, but they lose their concentration when Barney locks himself in the toilet after fellating Hans while in a k-hole. Mark invites Dobby to the Fuck Bunker, where she leads him to the women's toilets to have sex. Before they do anything, they are interrupted by Sophie vomiting. When Sophie reveals that she is married to Mark, Dobby angrily leaves. Barney leaves and gets into Ian's car, losing the band their slot. Mark helps Sophie into the same car. Ian is angry when Jez says that Barney sucked Hans off, then he drives off with Sophie and Barney.

Mark is angry with Jez for eating his food. Jez tells Mark that he has spent all of the money his mother gave him, resulting in him being forced to find money from somewhere else. Jez donates sperm amongst other things but he cannot earn enough cash to support himself.

Mark uses the promotion he was given at the end of series 5 to buy a new sofa and get Jez a job at JLB Credit's call centre. A fire alarm on Jez's first day causes all the staff to flee the building. It turns out that the alarm was a hoax by Johnson to evacuate the building and tell them that everyone at JLB Credit's London branch is being made redundant. Mark decides to start a campaign group to get everyone's job back, so that he can have some money to pay for the sofa and keep seeing Dobby. He impersonates JLB's Frankfurt boss Steffan Strauss, while having a toothbrush moustache drawn on with marker. It goes well, but Steffan arrives and offers Mark a cheque for £15,000 to end the campaign. He agrees and takes the money, but soon after is encouraged to break into the office with Jez, Jeff, Dobby and Gerrard, which he does to get Dobby to like him more. They vandalise the office, but are interrupted by Johnson and Steffan, who arrive at the office as Jez and Dobby push the photocopier down the stairs. Steffan takes the cheque back.

Mark discovers that Jez's girlfriend Elena is in a long-term relationship, when he hears her talking on her mobile phone. Her partner has been away and is moving back in with Elena. Elena does not want to tell Jez; she pressures Mark to tell him. Jez is initially disappointed, but is relieved when Elena tells her that her partner is a woman. Mark's former boss, Johnson, offers him the chance to go into business with him. However, Mark's excitement is dampened when Alan begins to show signs of becoming mentally ill due to his fall from power at JLB. Mark gets a job as a waiter at Gail's Mexican restaurant, Banditos, only to discover that Johnson's claims were legitimate, before being amply dismissed for working at the restaurant. Jez and Elena struggle to keep their affair secret. Big Suze is disappointed with Johnson and comes to Jez and Mark's flat. Johnson comes to the flat, cons Mark out of more money and Jez arrives soon later. Johnson wrongly assumes that Big Suze is having sex with Jez and so punches him.

As a last desperate hurrah before the baby arrives, Mark and Jez decide to throw a party. Mark sees it as his final opportunity to reignite his relationship with Dobby, while Jez think it will be a chance to improve his relationship with Elena. Hans brings a snake to the party. Sophie and Ian turn up and Mark and Sarah negotiate with them about visitation for Mark's unborn baby, before Mark vomits in a bin. Dobby leaves with Gerrard. Gail proposes to Elena, who accepts. Sarah is initially hostile to Jez, but later invites him into bed with her.

Elena and Gail's wedding is imminent, but Jez is busy plotting how to get rid of Gail and win Elena back. Elena tells Jez that she and Gail are moving to Quebec after the wedding. Mark, although conflicted about becoming a father, is worried that Jeff is becoming a potential rival dad. Sophie convinces a reluctant Mark to learn to drive so that he can take her to the hospital. However, he gives up trying to learn during his first lesson, because the instructor is terrible. Elena does not want Jez at the wedding, but Gail invites him. Jez and Gail get into a rowing boat and Gail accidentally falls into the water. Mark pretends he has passed his test and tries to drive Sophie to the hospital. Mark gets out of the car and Jez drives, almost running over Gail. Sophie realises that Mark cannot drive and Jez tells Gail about his affair with Elena. Sophie drives herself to the hospital with Mark and Jez in the back seat.

Mark, Jez and Sophie are at the hospital, where she is giving birth to Mark's baby. Elsewhere in the hospital, Jez meets Zahra, who is visiting her boyfriend, Ben, who is in a coma. Ben moves slightly, disappointing Jez, who is hoping for him to die in order to leave Zahra available.

With Dobby going out with Simon, Mark and Gerrard unite and try to break them up. However, when Dobby is once again single, they compete over her again. Dobby chooses Mark. Ben, having recovered, gives Jez a job in his music merchandising business. Jez tries to use his position to get Super Hans a record deal in return for a place in the band, but Hans rejects Jez for his poor saxophone performance. Ben thinks the band is awful and tells Jez not to sign them.

Mark picks Jeremy up from Zahra's flat on the way to Mark's son's christening. After getting themselves locked between the flat and the front door of the building, Mark and Jez argue over their strategy of escape. They find a key to the flat and let themselves in. Zahra and Ben return and, after initially hiding, they reveal themselves. They falsely claim to Ben that they use the flat to have sex together, which he does not believe. Mark and Jez miss baby Ian's christening.

Zahra has dumped Ben for cheating on her, and invites Jez to move in with her. At a party, Dobby overhears Mark insulting her while talking to Jeff and Johnson, and leaves angrily. Mark and Jez follow her trail to Hans' party, which has become so degenerate that Hans and Jez both emerge shaken after only a few seconds. Mark, Jez, Hans and Hans' girlfriend Yoko go to Suze's party, where they find that Dobby has gone to Gerrard's party. He refuses to allow the group to enter, but Mark sees and talks to Dobby through the fence. Mark tunnels under the fence and apologises to her. He asks her to move in with him, and she accepts. After it emerges that Jez made sexual hand gestures at Yoko during Suze's party, Zahra dumps Jez, and Hans beats him with a broom. Jez's living arrangements are now uncertain.

Jez and Hans give up their band. Jez begins therapy, paid for by Mark, but changes his mind whilst in the waiting room and instead goes to an Indian restaurant. Mark discovers this and treats Jez to an Indian takeaway while he is still full so he breaks down and admits that he had spent Mark's money on a meal for himself instead of therapy as originally promised. Mark is increasingly worried that Dobby will move in with his rival, Gerrard, who in illness has become dependent on her company. Mark's concern increases when Gerrard hints to Dobby and Mark that he would gladly allow her to live with him rent-free. Mark vandalises Dobby's microwave oven in an attempt to rush her into moving in with him. Gerrard dies of the flu and Dobby helps his sister to arrange the funeral. Mark attends interviews for a job at a bathroom supplies company, after Hans, who works there, suggests it to him. Dobby tells Mark that Gerrard left her a sum of money in his will.

Dobby suggests to Mark that they go interrailing together, an idea that Mark tries to discourage her from. Hans becomes Mark's boss at the bathroom supplies company where they work. Jez agrees to leave the flat and move in with Hans. Mark employs mentally unstable decorator Andy (Liam Noble) to repaint the kitchen. Jez tries to help Andy by giving him free 'life coaching'. Mark enrols on an 18-month MBA evening course, where he meets Stephanie (Josephine Butler), an attractive older woman whose husband left her. Jez encourages Mark to have a fling with her. She invites Mark to her place, where they drink wine and chat. Despite the conversation going well, he leaves without telling her upon seeing a note that Dobby had left him with his lunch. Jez realises that his attempt to help Andy failed and, at Andy's request, hits him several times. Mark tells Dobby of the course that he is on and that he loves her, disappointing Jez, who wants Dobby himself. Mark and Dobby tell each other that they will interrail when they both have enough free time.

Mark, in an attempt to deal with a damp patch on his bedroom wall to hasten the asthmatic Dobby to move in, launches a campaign to be elected chairman of the freehold committee of Apollo House. He is elected and authorises repair work to be done on his flat. Mark is sacked from his job as Jez helps him fraudulently sell taps to a bathroom supplier, claiming that they are used by Mark Ronson, before being rumbled when they are seen together by the buyer that they had just conned. Jez, desperate to avoid living with Hans, sparks up an old relationship with Mark's sister Sarah, and moves in with her and her five-year-old son, Joshy. He cannot cope with the demands she makes on him, so he wants to move back to Mark's. He refuses, so Jez stays with Andy, sleeping on his floor in a rubbish sack. He cannot cope with that, so comes back to Mark's just after Dobby has arrived to move in. Jez asks if he can stay the night and after that sleeps on Hans' floor in a sleeping bag. Dobby suggests to Mark that Jez stay at Mark's for longer and that she is fine with delaying moving in. Jez and Dobby play a video game together.

Jez moves into a sleeping bag at Hans' place, but the room has snakes in it. Dobby is offered a job in New York by ex-boyfriend Simon. She is uncertain about the offer, so Mark tries to persuade her to stay. He tries to get information about the position from him whilst playing five-a-side football in Hackney, but makes a fool of himself due to his lack of knowledge of and ability at the sport. He decides to take her away for the weekend in the Quantocks, where he will propose to her. Jez comes along and tells Mark that he is in love with her. Mark tells Dobby that Jez is insane. Mark and Jez physically fight whilst Dobby walks away. She texts Mark to tell him that she is taking the NY job and is flying there that night.

Mark and Jez meet for the first time in six months to attend the stag do for a supposedly sober Hans. Jez is reluctant to apologise to Mark. Jez is sleeping in Hans' bath, until he evicts him from the flat by having Jez pretend to be in possession of Hans' cocaine. Mark has a new job at a bank, which Johnson helped him to obtain. Mark's colleague Jerry has moved in with Mark. Mark grants Jez a loan at his bank. Jez persuades Mark to evict Jerry and allow him to move back in. Jez calls Hans to assist with the eviction, where Jerry is bagged up, waterboarded with beer and left in a lift.

Hans asks Mark to be the best man at his wedding to Molly because he wants a conventional guy to perform that role. It is revealed during the wedding that his real name is Simon. Dobby, having taken on a new hipster personality in New York, returns to attend the wedding and brings Gregory, her American boyfriend. Jez coaches Megan at her place, where he meets her boyfriend, Joe. Mark puts webcams in the flat. He views the webcams on a laptop, along with Dobby, Gregory and Hans who all see Jez have sex with Joe in Mark's bedroom. Mark, Jez, Dobby and Hans travel to rural Norfolk for the wedding. Gregory finds out that Mark was following Dobby's movements on her phone and punches him, only after Mark finds himself no longer enamoured with Dobby. Jez gives an awful speech at the reception.

Mark tracks down April (whom he had unsuccessfully tried to seduce by returning to his alma mater, Dartmouth University, and pretending to be a History student years earlier, in series 2, episode 4), at a book signing. She tells him that she writes with her husband Angus and that she lectures at a college. He invites her to a dinner party at the flat that he claims he is holding tomorrow. April says that her husband will be unable to attend because he will be going to a conference, so Mark tries to rekindle his relationship with her. She later tells him that Angus will be attending after all. Due to a lack of ingredients in the flat, combined with the short notice of the dinner party, Mark has to resort to using the only ingredients available in his kitchen, concocting an egg, lettuce and bean casserole as well as a lettuce cocktail, both of which he passes of as Moroccan. Jez starts having sex with Megan, whilst still having sex with Joe. Megan turns up at the flat whilst Joe is there. Mark tells Jez, Joe and Megan to stay for dinner, to convince Angus that the dinner party is genuine. Joe and Megan find out that they're both having sex with Jez, so they decide to try a three-way relationship, but they disagree on who will have sex with whom on which days.

Jez and Hans publish their first track on a football video on YouTube, but they disagree of the authorship of the lyrics. Jez tags along with Mark when he takes his son Ian for playtime at a soft play centre in Thornton Heath called the Kid Kave. Sophie visits the flat and tells Mark that she thinks her boyfriend, a tutor, is having an affair with the mother of one of his pupils. Sophie suggests that she and Mark reconcile. April arrives at the door whilst Sophie is still in the flat; Mark suggests that April and her niece join him and Ian at Kid Kave, but April declines his suggestion and tells Mark that she does not want a relationship with him. Jez tells Mark that he brought one of Hans' snakes in his bag. April later turns up at Kid Kave with her niece. She and Mark have sex in a toilet. Sophie turns up at Kid Kave whilst drunk and falls asleep in a ball pit. Mark tries to decide between miserable stability with Sophie, or whether to continue to pursue April. The snake escapes, so Mark and Jez leave.

Jez turns 40 and Mark is throwing a party for him. Jez has told Joe (who is in his late twenties) that he is turning 39. Jez starts drinking his own urine, believing it will help him stay youthful. Mark is dismissed from his job for not following proper procedure when granting Jez his loan. Mark tracks down April with the intention of poaching her from Angus. Mark and April go to a café, where they drink coffee. April tells Mark that she and Angus are going on a cruise together, but she receives a blunt and dismissive text from Angus, telling her that he is going to Ibiza instead of the cruise. Mark and April have sex in the café's toilet. Mark returns to the flat and tells Jez that he is going on the cruise with April. Mark discovers that Jez and Hans have kidnapped Angus and are holding him at the flat. Guests, including April, arrive for Jez's party. Angus comes into the room where the party is, telling everyone what happened. Joe finds out that Jez is 40 and Jez tells Joe that he cannot keep up with Joe's partying lifestyle; Joe decides to go back to Megan. April leaves with Angus. Hans splits from Molly and says he is emigrating to Macedonia.

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