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{{Short description|Overview of and topical guide to change in the heritable characteristics of organisms}}
{{See also|Index of evolutionary biology articles|Evolution (disambiguation)}}
<!--... Attention: THIS IS AN OUTLINE <!--... Attention: THIS IS AN OUTLINE


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See ] See ] and
for more details. ] for more details.
Further improvements Further improvements
to this outline are on the way to this outline are on the way
...--> ...-->
The following ] is provided as an overview of and topical guide to ]:
{{See also|Index of evolutionary biology articles|Evolution (disambiguation)}}
]
The following ] is provided as an overview of and topical guide to evolution:
{{Evolutionary biology}}
]
''']''' change in ]s of ] over generations due to ], ], ], and ]. Also known as '''descent with modification'''. Over time these evolutionary processes lead to formation of new species (]), changes within lineages (]), and loss of species (]). "Evolution" is also another name for ], the subfield of ] concerned with studying ] that produced the ] on Earth. In ], '''evolution''' is change in the ] ] of ] over generations due to ], ], ], and ]. Also known as '''descent with modification'''. Over time these evolutionary processes lead to formation of new species (]), changes within lineages (]), and loss of species (]). "Evolution" is also another name for ], the subfield of ] concerned with studying ] that produced the ] on Earth.


==Fundamentals about evolution== ==Fundamentals about evolution==
Line 23: Line 25:


===Introduction=== ===Introduction===
* ] – non-technical introduction * {{Annotated link|Introduction to evolution}}
* {{Annotated link|Evolution}}
* ] – more technical introduction
* {{Annotated link|Evolution as fact and theory}}
* ] – basis for scientific acceptance as ] (well-substantiated explanation) based on ] (empirical data and objective verifiable observation)


===Basic principles=== ===Basic principles===
* {{annotated link|Macroevolution}}
* ] – change above level of species, including:
** {{annotated link|Speciation}}
** ] – evolutionary process by which new biological species arise
*** Natural speciation *** Natural speciation
**** {{annotated link|Allopatric speciation}}
**** ] – speciation that occurs after biological populations become isolated from each other (also known as geographic speciation)
**** ] – speciation that occurs in biological populations with adjacent ranges **** {{annotated link|Peripatric speciation}}
**** {{annotated link|Parapatric speciation}}
**** ] – speciation that occurs in biological populations with adjacent, but not significantly overlapping ranges
**** {{annotated link|Sympatric speciation}}
**** ] – speciation that occurs in biological populations that inhabit the same geographic region
*** Artificial speciation *** Artificial speciation
**** {{annotated link|Animal husbandry}}
**** ] – management and care of farm animals, typically involving ]
**** {{annotated link|Plant breeding}}
**** ] – ] of plants to produce desired characteristics
**** ] – direct manipulation of an organism's genome using ] **** {{annotated link|Genetic engineering}}
*** {{annotated link|Hybrid speciation}}
*** ] – speciation wherein hybridization between two different species leads to a new species
** {{annotated link|Despeciation}}
** ] – loss of a unique species due to combining with another previously distinct species
** {{annotated link|Anagenesis}}
** ] – changes in a lineage that result in a new morphospecies distinct in form from an ancestral species ("phyletic transformation")
** {{annotated link|Extinction}}
** ] – end of a lineage such that there are no longer living populations of a species (or other taxon)
* {{annotated link|Microevolution}}
* ] – change within species or populations, due to ], selection (] and ]), ], and ]
** {{annotated link|Artificial selection}}
** ] – the process by which humans increase particular traits in a lineage or population by choosing which individuals have offspring together (also called ])
** {{annotated link|Natural selection}}
** ] – differential survival and reproduction of individuals due to differences in heritable traits (]), a key mechanism of ]
*** {{annotated link|Sexual selection}}
*** ] – a mode of natural selection wherein members of one gender choose mates of the other gender to mate with, resulting in distinct gender-based differences
** {{annotated link|Mutation}}
** ] – a permanent change of the genome of an organism (nucleotide sequence), a key mechanism of ]
** {{annotated link|Gene flow}}
** ] – movement of genes from one population to another (through migration, dispersal, transport of pollen, etc.)
** {{annotated link|Genetic drift}}
** ] – change in frequency of a gene variant (allele) in a population due to random sampling


===Subfields=== ===Subfields===
* {{annotated link|Biology}}
* ] – the study of distribution of species and ecosystems in geographic space and through geological time
** {{annotated link|Evolutionary biology}}
* ] – the study of genetics in natural populations
*** {{annotated link|Evolutionary developmental biology}}
* ] – study of the evolutionary processes that produced the diversity of life on Earth. More specifically, it studies the descent of species, and the origin of new species.
** {{annotated link|Genetics}}
** ] – the study of developmental processes to determine the ancestral relationships and discover how developmental processes evolved (informally called evo-devo)
* {{annotated link|Biogeography}}
* ] – the study of ecology with explicit consideration of evolutionary histories of species, or conversely the study of evolution that incorporates an understanding of ecological interactions between the species
* {{annotated link|Ecological genetics}}
* ] – the study of changes in functional characteristics over generations as the result of selection
* {{annotated link|Evolutionary biology}}
* ] – branch of biological classification that classifies organisms based on phylogenetic relationship (shared descent), progenitor-descendant relationship (serial descent), and degree of evolutionary change
** {{annotated link|Evolutionary developmental biology}}
* ] – study of evolution using controlled experiments to test hypotheses and theories
* {{annotated link|Evolutionary ecology}}
* ] – study of change in sequence composition of cellular molecules such as DNA, RNA, and proteins across generations
* {{annotated link|Evolutionary physiology}}
* ] – study of evolutionary history, development, and relationships among groups of organisms
* {{annotated link|Evolutionary taxonomy}}
* ] – study of distribution and change in frequency of alleles within populations
* {{annotated link|Experimental evolution}}
* ] – study of evolution of life based on the fossil record
* {{annotated link|Molecular evolution}}
** ] – study of ancient viruses based on fossil viruses
* {{annotated link|Phylogenetics}}
** ] – summary of key dates and events in the history of ]
* {{annotated link|Population genetics}}
* ] – study of diversification of living forms, both past and present, and relationships among living organisms through time
* {{annotated link|Paleontology}}
** {{annotated link|Paleovirology}}
** {{annotated link|Timeline of paleontology}}
* {{annotated link|Systematics}}


===History=== ===History===
* {{annotated link|Charles Darwin}}
* ] – English naturalist and geologist, best known for his contributions to evolutionary theory
** {{annotated link|On the Origin of Species|''On the Origin of Species''}}
:* '']'' – work of scientific literature by Charles Darwin, considered to be foundation of evolutionary biology (published 1859)
:* ] – artistic depictions concerning Charles Darwin and evolutionary theory ** {{annotated link|Caricatures of Charles Darwin and his evolutionary theory in 19th-century England}}
** {{annotated link|Darwinism}}
* ] – historical account of evolutionary thought from antiquity through present
* {{annotated link|History of evolutionary thought}}
:* By period or event
** By period or event
::* ]
*** {{annotated link|Evolutionary ideas of the Renaissance and Enlightenment}}
::* ]
*** {{annotated link|Transmutation of species}}
::* ]
*** {{annotated link|1860 Oxford evolution debate}}
::* ]
*** {{annotated link|Neo-Darwinism}}
::* ]
*** {{annotated link|The eclipse of Darwinism}}
::* ]
*** {{annotated link|Evolutionary progress}}
::* ], early 20th century synthesis of ideas from various fields of biology (particularly genetics, systematics, ecology, and paleontology)
*** {{annotated link|Scopes Trial}}
::* ]
*** {{annotated link|Modern synthesis (20th century)|Modern synthesis}}
:* By field
**** {{annotated link|Extended evolutionary synthesis}}
::* ]
*** {{annotated link|Evolutionary biology#Current research topics|Current research}}
::* ]
** By field
::* ]
*** {{annotated link|Evolutionary developmental biology}}
::* ]
* ] *** {{annotated link|History of evolutionary psychology}}
*** {{annotated link|History of molecular evolution}}
*** {{annotated link|History of paleontology}}
* {{annotated link|Social effect of evolutionary theory}}


==Evolutionary theory and modelling== ==Evolutionary theory and modelling==
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===Population genetics=== ===Population genetics===


* ] * {{annotated link|Population genetics}}
* Process * Process
** ] ** {{annotated link|Mutation}}
** ] ** {{annotated link|Selection (biology)|Selection}}
*** ] *** {{annotated link|Natural selection}}
**** ] **** {{annotated link|Sexual selection}}
*** ] *** {{annotated link|Artificial selection}}
*** ] *** {{annotated link|Ecological selection}}
** ] ** {{annotated link|Gene flow}}
** ] ** {{annotated link|Genetic drift}}
*** ] *** {{annotated link|Small population size}}
*** ] *** {{annotated link|Population bottleneck}}
*** ] *** {{annotated link|Founder effect}}
*** ] *** {{annotated link|Coalescent theory}}
* Variation * Variation
** ] ** {{annotated link|Genetic variation}}
*** ] *** {{annotated link|Genetic diversity}}
*** ] *** {{annotated link|Gene frequency}}
*** ] *** {{annotated link|Polymorphism (biology)}}
* Key concepts * Key concepts
** ] ** {{annotated link|Hardy–Weinberg principle|Hardy-Weinberg law}}
** ] ** {{annotated link|Genetic linkage}}
** ] ** {{annotated link|Identity by descent}}
** ] <!--Haldane's general selection model--> ** {{annotated link|Linkage disequilibrium}} <!--Haldane's general selection model-->
** ] ** {{annotated link|Fisher's fundamental theorem of natural selection|Fisher's fundamental theorem}}
** ] ** {{annotated link|Neutral theory of molecular evolution|Neutral theory}}
** ] ** {{annotated link|Shifting balance theory}}
** ] ** {{annotated link|Price equation}}
** ] ** {{annotated link|Coefficient of relationship}}
** ] ** {{annotated link|Fitness (biology)|Fitness}}
** ] ** {{annotated link|Heritability}}
* Effects of selection * Effects of selection
** ] ** {{annotated link|Genetic hitchhiking}}
** ] ** {{annotated link|Negative selection (natural selection)}}
* Related topics * Related topics
** ] ** {{annotated link|Microevolution}}
** ] ** {{annotated link|Evolutionary game theory}}
** ] ** {{annotated link|Fitness landscape}}
** ] ** {{annotated link|Genetic genealogy}}
** {{annotated link|Quantitative genetics}}
** ], branch of population genetics that deals with phenotypes which vary continuously


===Evolutionary phenomena=== ===Evolutionary phenomena===


* ] * {{annotated link|Adaptation}}
* ] * {{annotated link|Adaptive radiation}}
* ] * {{annotated link|Coevolution}}
* ] * {{annotated link|Concerted evolution}}
* ] * {{annotated link|Convergent evolution}}
** ] ** {{annotated link|List of examples of convergent evolution}}
* ] * {{annotated link|Divergent evolution}}
** ] ** {{annotated link|Divergent evolution in animals}}
* ] * {{annotated link|Evolution of ageing}}
* ] * {{annotated link|Evolution of biological complexity}}
* ] * {{annotated link|Evolution of multicellularity}}
* ] * {{annotated link|Evolution of photosynthesis}}
* ] * {{annotated link|Evolution of sexual reproduction}}
* ] * {{annotated link|Evolutionary arms race}}
* ] * {{annotated link|Evolutionary capacitance}}
* ] * {{annotated link|Evolutionary fauna}}
* ] * {{annotated link|Evolutionary pressure}}
* ] * {{annotated link|Evolutionary radiation}}
* ] * {{annotated link|Evolutionary trap}}
* {{annotated link|Evolvability}}
* ]
* {{annotated link|Exaptation}}
* ] –
* {{annotated link|Extinction}}
* ] – capacity of a system for adaptive evolution. Beneficial mutations are always rare, but if they are too rare, then adaptation cannot occur. Biological genomes are structured in ways that make beneficial changes less unlikely than they would otherwise be. Evolution has created not just fitter organisms, but populations of organisms that are better able to evolve.
* {{annotated link|Fitness (biology)}}
* ]
** {{annotated link|Inclusive fitness}}
* ]
*** {{annotated link|Kin selection}}
** ]
** {{annotated link|Reproductive success}}
*** ]
* {{annotated link|Genetic recombination}}
** ]
* ] * {{annotated link|Horizontal gene transfer in evolution}}
* {{annotated link|Human evolution (origins of society and culture)}}
* ]
* {{annotated link|Inversion (evolutionary biology)}}
* ]
* {{annotated link|Mosaic evolution}}
* ]
* ] * {{annotated link|Parallel evolution}}
* ] * {{annotated link|Quantum evolution}}
* ] * {{annotated link|Recurrent evolution}}
* {{annotated link|Robustness (evolution)}}
* ]
* {{annotated link|Speciation}}
* ]
* ]
* ]


===Modelling=== ===Modelling===
* ] * {{annotated link|Emergent evolution}}
* ] * {{annotated link|Epic of evolution}}
* ] * {{annotated link|Evolution window}}
* ] * {{annotated link|Evolutionary dynamics}}
* ] * {{annotated link|Evolutionary game theory}}
* ] * {{annotated link|Evolutionary graph theory}}
* ] * {{annotated link|Evolutionary invasion analysis}}
* ] * {{annotated link|Largest-scale trends in evolution}}


==Taxonomy, systematics, and phylogeny== ==Taxonomy, systematics, and phylogeny==


===Fundamentals=== ===Fundamentals===
* ] * {{annotated link|Taxonomy (biology)|Taxonomy}}
** ] ** {{annotated link|Alpha taxonomy}}
** ] ** {{annotated link|Biological classification}}
** ] ** {{annotated link|Binomial nomenclature}}
** ] ** {{annotated link|Evolutionary taxonomy}}
** ] ** {{annotated link|Catalogue of life}}
** ] ** {{annotated link|Homonym (biology)}}
** ] ** {{annotated link|Integrated Taxonomic Information System}}
** ] ** {{annotated link|International Code of Zoological Nomenclature}}
** ] ** {{annotated link|International Code of Nomenclature for algae, fungi, and plants}}
** ] ** {{annotated link|Linnaean taxonomy}}
** ] ** {{annotated link|Phenetics}}
** ] ** {{annotated link|Species 2000}}
** ] ** {{annotated link|Taxon}}
** ] ** {{annotated link|Taxonomic rank}}
** ] ** {{annotated link|Type (biology)}}
** ] ** {{annotated link|Species description}}
* {{annotated link|Systematics}}
* ] – study of diversification of living forms, both past and present
** ] ** {{annotated link|Cladogram}}
** ] ** {{annotated link|Phylogenetic tree}}
* ] * {{annotated link|Phylogenetics}}
** ] ** {{annotated link|Cladistics}}
** ] ** {{annotated link|Computational phylogenetics}}
** {{annotated link|Common descent}}
** ] – how different lineages of organisms share a most recent common ancestor
** ] – ** {{annotated link|Evidence of common descent}}
** ] ** {{annotated link|Evolutionary grade}}
** ] ** {{annotated link|Lineage (evolution)}}
** ] ** {{annotated link|Molecular phylogenetics}}
** {{annotated link|Most recent common ancestor}}
** ] – most recent individual from which all organisms in a group are directly descended. It is impossible to identify the specific MRCA of a large set of individuals, but an estimate of the time at which the MRCA lived can often be given.


===Basic concepts of phylogenetics=== ===Basic concepts of phylogenetics===
* ] * {{annotated link|Phylogenetic tree}}
* ] * {{annotated link|Phylogenetic network}}
* ] * {{annotated link|Long branch attraction}}
* {{annotated link|Clade}}
* ] vs ]
* {{annotated link|Evolutionary grade|Grade}}
* ]
* {{annotated link|Ghost lineage}}


===Inference methods=== ===Inference methods===
* ] * {{annotated link|Maximum parsimony (phylogenetics)}}
* ] * {{annotated link|Minimum evolution}}
* ''Probabilistic methods'' * ''Probabilistic methods''
** ] ** {{annotated link|Maximum likelihood estimation}}
** ] ** {{annotated link|Bayesian inference in phylogeny|Bayesian inference}}
* ] * {{annotated link|Distance matrices in phylogeny}}
** ] ** {{annotated link|Neighbor joining}}
** {{annotated link|UPGMA|Unweighted Pair Group Method with Arithmetic Mean|abbreviation=UPGMA}}
** ]
** ] ** {{annotated link|Least squares inference in phylogeny}}
* ] * {{annotated link|Three-taxon analysis}}


===Current topics=== ===Current topics===
* {{annotated link|PhyloCode|aka=International Code of Phylogenetic Nomenclature}}
* ]
* ] * {{annotated link|DNA barcoding}}
* ] * {{annotated link|Molecular phylogenetics}}
* ] * {{annotated link|Phylogenetic comparative methods}}
* ] * {{annotated link|Phylogenetic network}}
* ] * {{annotated link|Phylogenetic niche conservatism}}
* ] * {{annotated link|List of phylogenetics software}}
* ] * {{annotated link|Phylogenomics}}
* ] * {{annotated link|Phylogeography}}
* ] * {{annotated link|DNA phylogeny}}


===Group Traits=== ===Group Traits===
* ] * {{annotated link|Symplesiomorphy}}
* ] * {{annotated link|Synapomorphy|Apomorphy}}
* ] * {{annotated link|Synapomorphy}}
* ] * {{annotated link|Autapomorphy}}


===Group Types=== ===Group Types===
* ] * {{annotated link|Monophyly}}
* ] * {{annotated link|Paraphyly}}
* ] * {{annotated link|Polyphyly}}


==Evolution of biodiversity== ==Evolution of biodiversity==
* {{annotated link|Biodiversity}}
* ] – variety of different types of life found on the Earth and the variations within species.<ref name=UN>{{cite web |url=http://www.unep-wcmc.org/what-is-biodiversity_50.html |title=What is biodiversity? |publisher=United Nations Environment Programme, World Conservation Monitoring Centre}}</ref> It is a measure of the variety of organisms present in different ecosystems. This can refer to genetic variation, ecosystem variation, or species variation (number of species)<ref name=UN/> within an area, biome, or planet. Terrestrial biodiversity tends to be greater near the equator,<ref>{{cite journal |last=Gaston |first=Kevin J. |title=Global patterns in biodiversity|journal=Nature|date=11 May 2000 |volume=405 |issue=6783 |pages=220–227 |doi=10.1038/35012228 |pmid=10821282}}</ref> which seems to be the result of the warm climate and high primary productivity.<ref>{{cite journal | last=Field | first=Richard | author2=Hawkins, Bradford A.; Cornell, Howard V.; Currie, David J.; Diniz-Filho, J. Alexandre F.; Guégan, Jean-François; Kaufman, Dawn M.; Kerr, Jeremy T.; Mittelbach, Gary G.; Oberdorff, Thierry; O’Brien, Eileen M.; Turner, John R. G.|title=Spatial species-richness gradients across scales: a meta-analysis|journal=Journal of Biogeography|date=1 January 2009|volume=36|issue=1|pages=132–147|doi=10.1111/j.1365-2699.2008.01963.x}}</ref> Biodiversity is not distributed evenly on Earth. It is richest in the tropics. Marine biodiversity tends to be highest along coasts in the Western Pacific, where sea surface temperature is highest and in the mid-latitudinal band in all oceans.


===Origin and evolutionary history of life=== ===Origin and evolutionary history of life===
* ] * {{annotated link|Abiogenesis}}
* {{annotated link|History of life}}
* ] – outline of major events during evolution of life on Earth
* ] – more concise outline * {{annotated link|Timeline of the evolutionary history of life}}


===Evolution of organisms=== ===Evolution of organisms===


====Evolution of tetrapods==== ====Evolution of tetrapods====
* ] * {{annotated link|Evolution of tetrapods}}
** ] ** {{annotated link|Evolution of dinosaurs}}
***] *** {{annotated link|Evolution of birds}}
** ] ** {{annotated link|Evolution of mammals}}
*** ] *** {{annotated link|Evolution of cetaceans}}
*** ] *** {{annotated link|Evolution of the horse|Evolution of horses}}
*** ] *** {{annotated link|Evolution of primates}}
**** ] **** {{annotated link|Evolution of humans}}
***** ] ***** {{annotated link|Evolution of human intelligence}}
***** ] ***** {{annotated link|Human evolutionary genetics}}
***** ] ***** {{annotated link|Sexual selection in human evolution}}
***** ] ***** {{annotated link|Timeline of human evolution}}
**** ] **** {{annotated link|Evolution of lemurs}}
*** ] *** {{annotated link|Evolution of sirenians}}
** ] ** {{annotated link|Evolution of reptiles}}


====Evolution of other animals==== ====Evolution of other animals====
* ] * {{annotated link|Evolution of brachiopods}}
* ] * {{annotated link|Evolution of cephalopods}}
* ] * {{annotated link|Evolution of fish}}
* ] * {{annotated link|Evolution of insects}}
** ] ** {{annotated link|Evolution of butterflies}}
** ] ** {{annotated link|Peppered moth evolution}}
* ] * {{annotated link|Evolution of molluscs}}
* ] * {{annotated link|Evolution of spiders}}


====Evolution of plants==== ====Evolution of plants====
* ] * {{annotated link|Evolution of plants}}
** ] ** {{annotated link|Evolutionary anachronism}}
** ] ** {{annotated link|Plant evolution}}
** ] ** {{annotated link|Plant evolutionary developmental biology}}
** ] ** {{annotated link|Timeline of plant evolution}}


====Evolution of other taxa==== ====Evolution of other taxa====
* ] * {{annotated link|Evolution of fungi}}
* ] * {{annotated link|Evolution of viruses}}
** ] ** {{annotated link|Evolution of influenza}}
* ] * {{annotated link|E. coli long-term evolution experiment}}


===Evolution of cells, organs, and systems=== ===Evolution of cells, organs, and systems===
* ] * {{annotated link|Evolution of cells}}
* ] * {{annotated link|Evolution of flagella}}
* ] * {{annotated link|Evolution of mammalian auditory ossicles}}
* ] * {{annotated link|Evolution of nervous systems}}
* ] * {{annotated link|Evolution of snake venom}}
* ] * {{annotated link|Evolution of the brain}}
* ] * {{annotated link|Evolution of the eye}}
** ] ** {{annotated link|Evolution of color vision}}
** ] ** {{annotated link|Evolution of color vision in primates}}
* ] * {{annotated link|Immune system#Other mechanisms and evolution|Immune system}}
* ] * {{annotated link|Metabolism#Evolution|Evolution of metabolism}}


===Evolution of molecules and genes=== ===Evolution of molecules and genes===
* ] * {{annotated link|Directed evolution}}
* ] * {{annotated link|Error threshold (evolution)}}
* {{annotated link|Gene-centered view of evolution}}
* ]
* {{annotated link|Genome evolution}}
* ]
* ] * {{annotated link|Hologenome theory of evolution}}
* ] * {{annotated link|Models of DNA evolution}}
* ] * {{annotated link|Molecular evolution}}
* ] ** {{annotated link|History of molecular evolution}}
** ] ** {{annotated link|Neutral theory of molecular evolution}}
** ] ** {{annotated link|Nearly neutral theory of molecular evolution}}
** ] * {{annotated link|Neutral network (evolution)}}
* ] * {{annotated link|RNA-based evolution}}
* ]


===Evolution of behaviour=== ===Evolution of behaviour===
* ] * {{annotated link|Co-operation (evolution)}}
* ] * {{annotated link|Evolution of biparental care in tropical frogs}}
* ] * {{annotated link|Evolution of emotion}}
* ] * {{annotated link|Empathy#Evolution|Evolution of empathy}}
* ] * {{annotated link|Evolution of eusociality}}
* ] * {{annotated link|Monogamy in animals}}
* ] * {{annotated link|Reciprocal altruism}}
* ] * {{annotated link|Reciprocity (evolution)}}


===Evolution of other processes=== ===Evolution of other processes===
* ] * {{annotated link|Evolution of ageing}}
** ] ** {{annotated link|Death#Evolution of aging and mortality|Evolution of aging and mortality}}
** ] ** {{annotated link|Programmed cell death#Evolutionary origin|Origin of programmed cell death}}
* ] * {{annotated link|Origin of avian flight}}
* ] * {{annotated link|Evolution of biological complexity}}
* ] * {{annotated link|Mosaic evolution}}
* ] * {{annotated link|Evolution of multicellularity}}
* ] * {{annotated link|Evolution of sexual reproduction}}
** ]s ** {{annotated link|Mating type}}s
** ] ** {{annotated link|Anisogamy#Evolution of anisogamy|Gamete differentiation/sexes}}
** ] ** {{annotated link|Sex-determination system#Evolution of sex-determination systems|Sex-determination}}
** ] ** {{annotated link|Biological life cycle|Life cycles/nuclear phases}}


==Applications in other disciplines== ==Applications in other disciplines==


* {{annotated link|Applications of evolution}}
* ] – practical applications in fields such as ecology, artificial intelligence, medicine, and computer science
* ] – * {{annotated link|Biological anthropology}}
* ] – * {{annotated link|Evolutionary aesthetics}}
* ] – * {{annotated link|Evolutionary anthropology}}
* ] – * {{annotated link|Evolutionary computation}}
** {{annotated link|Evolutionary algorithm}}
* ] –
* {{annotated link|Evolutionary economics}}
** ] –
** {{annotated link|Kenneth Boulding's evolutionary perspective}}
* ] –
* ] – * {{annotated link|Evolutionary epistemology}}
* ] – * {{annotated link|Evolutionary ethics}}
* ] – * {{annotated link|Evolutionary linguistics}}
* ] – * {{annotated link|Evolutionary medicine}}
* ] – * {{annotated link|Evolutionary neuroscience}}
* {{annotated link|Evolutionary psychology}}
** ] –
** {{annotated link|Biosocial criminology}}
** ] –
** {{annotated link|Criticism of evolutionary psychology}}
** ] –
** ] – ** {{annotated link|Evolution of morality}}
** {{annotated link|Evolution of schizophrenia}}
** ] –
** ] – ** {{annotated link|Evolutionary aesthetics}}
** {{annotated link|Evolutionary approaches to depression}}
** ] –
*** ] – ** {{annotated link|Evolutionary developmental psychology}}
** ] – *** {{annotated link|Evolutionary developmental psychopathology}}
** {{annotated link|Evolutionary educational psychology}}
** ] –
** ] – ** {{annotated link|Evolutionary ethics}}
** {{annotated link|Evolutionary leadership theory}}
** ] –
** ] – ** {{annotated link|Evolutionary musicology}}
** ] – ** {{annotated link|Evolutionary origin of religions}}
** ] – ** {{annotated link|Evolutionary psychology of language}}
** ] – ** {{annotated link|Evolutionary psychology of parenting}}
** ] ** {{annotated link|Evolutionary psychology of religion}}
** {{annotated link|Theoretical foundations of evolutionary psychology}}
* ] –
* {{annotated link|Evolutionary robotics}}
* {{annotated link|Evolutionary systems}}
* {{annotated link|Sociobiology}}
** {{annotated link|Sociocultural evolution}}
** {{annotated link|Cultural evolution}}
* {{annotated link|Universal Darwinism}}


==Evolutionary issues== ==Evolutionary issues==


===Controversy about evolution=== ===Controversy about evolution===
* ] (]) * {{annotated link|Creation–evolution controversy}}
** {{annotated link|Outline of the creation–evolution controversy}}
* ] * {{annotated link|Criticism of evolutionary psychology}}
* ] * {{annotated link|Evolutionary argument against naturalism}}
* ] * {{annotated link|Level of support for evolution}}
* ] * {{annotated link|Objections to evolution}}
* ] * {{annotated link|Social effects of evolutionary theory}}
* ] * {{annotated link|Theology of creationism and evolution}}


===Religious and philosophical views of evolution=== ===Religious and philosophical views of evolution===
* ] * {{annotated link|Acceptance of evolution by religious groups}}
* Atheistic evolution * {{annotated link|Conscious evolution}}
* ] * {{annotated link|Buddhism and evolution}}
* ] * {{annotated link|Catholic Church and evolution}}
* ] * {{annotated link|Hindu views on evolution}}
* ] * {{annotated link|Islamic views on evolution}}
* ] ** {{annotated link|Ahmadiyya views on evolution}}
:* ] * {{annotated link|Jewish views on evolution}}
* ] * {{annotated link|Mormon views on evolution}}
* ] * {{annotated link|Theistic evolution}}
* ]


=== Influence of evolutionary theory === === Influence of evolutionary theory ===
* ] * {{annotated link|Social effects of evolutionary theory}}
* See also '']'' * See also '']''


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* '']'' – book by zoologist and evolutionary biologist ], canonical publication of ], first published in 1942 by ] * '']'' – book by zoologist and evolutionary biologist ], canonical publication of ], first published in 1942 by ]
* '']'' – technical book on macroevolutionary theory by the Harvard paleontologist Stephen Jay Gould * '']'' – technical book on macroevolutionary theory by the Harvard paleontologist Stephen Jay Gould
* ] * {{annotated link|Evolutionary Biology (textbook)|''Evolutionary Biology''}}


===Journals=== ===Journals===


* {{annotated link|Evolution (journal)|''Evolution''}}
* ] – monthly scientific journal published by the Society for Study of Evolution (also called ''International Journal of Organic Evolution'')
* ] * {{annotated link|Evolutionary Anthropology (journal)|''Evolutionary Anthropology''}}
* {{annotated link|Evolutionary Bioinformatics|''Evolutionary Bioinformatics''}}
* ] –
* '']'' * {{annotated link|Evolutionary Psychology (journal)|''Evolutionary Psychology''}}
* '']'' * {{annotated link|Journal of Evolutionary Biology|''Journal of Evolutionary Biology''}}
* {{annotated link|Journal of Zoological Systematics and Evolutionary Research|''Journal of Zoological Systematics and Evolutionary Research''}}
* ] – Elsevier journal of review articles about ecology and evolution
* {{annotated link|Trends (journals)|''Trends in Ecology & Evolution''|abbreviation=TREE}}


===Organizations=== ===Organizations===
<!--* ] – --> <!--* {{annotated link|Center for Evolutionary Psychology}} -->
<!--* ] – --> <!--* {{annotated link|Department of Ecology and Evolutionary Biology, University of Colorado at Boulder}} -->
* ] – * {{annotated link|European Society for Evolutionary Biology}}
* ] – * {{annotated link|Society for the Study of Evolution}}
* ] – * {{annotated link|Evolutionary psychology research groups and centers}}
* ] – * {{annotated link|I. M. Sechenov Institute of Evolutionary Physiology and Biochemistry}}
* ] – * {{annotated link|Max Planck Institute for Evolutionary Anthropology}}
* ] – * {{annotated link|Max Planck Institute for Evolutionary Biology}}
* ] – * {{annotated link|National Evolutionary Synthesis Center}}
* ] – * {{annotated link|Systematic and Evolutionary Biogeography Association}}
* ] – * {{annotated link|Evolutionary Informatics Lab}}


==Evolution scholars and researchers== ==Evolution scholars and researchers==
* ] – * {{annotated link|List of evolutionary psychologists}}
* ] – * {{annotated link|List of members of the National Academy of Sciences (Evolutionary biology)}}


===Prominent evolutionary biologists=== ===Prominent evolutionary biologists===
* ] * {{annotated link|Charles Darwin}}
* ] * {{annotated link|Theodosius Dobzhansky}}
* ] * {{annotated link|Richard Dawkins}}
* ] * {{annotated link|Stephen Jay Gould}}
* ] * {{annotated link|J. B. S. Haldane}}
* ] * {{annotated link|Julian Huxley}}
* ] * {{annotated link|Thomas Henry Huxley}}
* ] * {{annotated link|Ronald Fisher}}
* ] * {{annotated link|Ernst Mayr}}
* {{annotated link|Alfred Russel Wallace}}
* ]
* {{annotated link|Sewall Wright}}


==See also== ==See also==
{{portal|Evolutionary biology}} {{portal|Evolutionary biology}}
* {{annotated link|Outline of biology}}

* ] ** {{annotated link|Outline of genetics}}
** ]

<!-- place the following, remove those that don't belong in this outline --> <!-- place the following, remove those that don't belong in this outline -->
* {{annotated link|Biogeography}}

* {{annotated link|Conscious evolution}}
* ] –
* {{annotated link|Ecology and Evolutionary Biology}}
* ] – claim that humanity has acquired the ability to choose what the species Homo sapiens becomes in the future, based on recent advancements in science, medicine, technology, psychology, sociology, and spirituality. Conscious evolution assumes that human beings may be positioned at the crest of the ongoing evolution of the universe.
* {{annotated link|Effective evolutionary time}}
* ] – degree program, in some North American universities, offering integrated studies in the disciplines of ecology and evolutionary biology.
* {{annotated link|Evolutionary acquisition of neural topologies}}
* ] – hypothesis that attempts to explain gradients, in particular latitudinal gradients, in species diversity. It was originally named "time hypothesis".<ref name="Latitudinal gradients(1978a)">K. Rohde: Latitudinal gradients in species diversity and their causes. I. A review of the hypotheses explaining the gradients. Biologisches Zentralblatt 97, 393-403, 1978a.</ref><ref name="Latitudinal gradients(1978b)">K. Rohde: Latitudinal gradients in species diversity and their causes. II. Marine parasitological evidence for a time hypothesis. Biologisches Zentralblatt 97, 405-418, 1978b.</ref>
* ] – * {{annotated link|Evolutionary anachronism}}
* ] * {{annotated link|Evolutionary approaches to depression}}
* {{annotated link|Evolutionary argument against naturalism}}
* ] –
* ] – * {{annotated link|Evolutionary art}}
** {{annotated link|Evolutionary music}}
* ] –
* {{annotated link|Evolutionary baggage}}
* ] – part of the genome of a population that was advantageous in past individuals but is disadvantageous under the pressures exerted by natural selection today.<ref>Appenzeller, T. 1999. "Test tube evolution catches time in a bottle." ''Science.'' 284: 2108-2110</ref>
* {{annotated link|Evolutionary Humanism}}
* ] – peer-reviewed open access scientific journal focusing on computational biology in the study of evolution.
* {{annotated link|Evolutionary informatics}}
* ] – philosophy that mixes teachings about Enlightenment from Eastern traditions with a Western scientific understanding of evolution.
* ] – * {{annotated link|Evolutionary landscape}}
* {{annotated link|Evolutionary Principle}}
* ] – subfield of informatics addressing the practice of information processing in, and the engineering of information systems for, the study of biological evolution, as well as the study of information in evolutionary systems, natural and artificial.
* {{annotated link|Extinction}}
* ] – metaphor;<ref name=Wright1932>Wright, Sewall (1932) The Roles of Mutation, Inbreeding, Crossbreeding, and Selection in Evolution. ''Proceedings of the Sixth International Congress of Genetics'' 1: 356-366</ref> (construct) used to think about and visualize the processes of ] (e.g. ] and ]) acting on a biological entity <ref name=Wright1988>Wright, Sewall (1988) Surfaces of Selective Value Revisited. ''The American Naturalist'' 131(1):115-123</ref> ( e.g., a ], ], ], ]).<ref name=Lee2008>Lee, Carol E. & Gelebiuk, Gregory W. (2008) Evolutionary origins of invasive populations. "Evolutionary Applications" 1: 427-448.</ref> This entity can be viewed as searching or moving through a search space. For example, the search space of a gene would be all possible nucleotide sequences. The search space is only part of an evolutionary landscape. The final component is the "y-axis," which is usually ].
** {{annotated link|Extinction event}}
* ] –
** {{annotated link|Human extinction}}
* ] –
** {{annotated link|Local extinction}}
* ] – death of an entire species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Extinction is the end of the evolution of a species, but related branches of the taxonomy may live on.
* {{annotated link|MEGA, Molecular Evolutionary Genetics Analysis}}
** ] – widespread and rapid decrease in the amount of life on Earth. Such an event is identified by a sharp reduction in the diversity and abundance of macroscopic life. Also known as a mass extinction or biotic crisis.
* {{annotated link|Sloshing bucket model of evolution}}
** ] – hypothesized end of the human species. Various scenarios have been discussed in science, popular culture and religion ''(see ])''
* {{annotated link|Spandrel (biology)}}
** ] (extirpation) – condition of a species (or other taxon) that ceases to exist in the chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinction can be reversed by reintroduction of the species to the area from other locations; wolf reintroduction is an example of this.* '']'' –
* {{annotated link|Speculative evolution}}
* ] – freely available software for conducting statistical analysis of molecular evolution and for constructing phylogenetic trees.
* ] – * {{annotated link|Transitional fossil}}

==References==
{{Reflist}}


==External links== ==External links==
{{sisterlinks|Evolution}} {{sister project links|Evolution}}


;General information ;General information
* {{In Our Time|Evolution|p00545gl|Evolution}} * {{In Our Time|Evolution|p00545gl|Evolution}}
* {{cite web |url = https://www.newscientist.com/topic/evolution |title = Evolution |publisher = New Scientist |accessdate = May 30, 2011 }} * {{cite web |url = https://www.newscientist.com/topic/evolution |title = Evolution |publisher = New Scientist |access-date = May 30, 2011 }}
* {{cite web |url = http://nationalacademies.org/evolution/ |title = Evolution Resources from the National Academies |publisher = ] |accessdate = May 30, 2011 }} * {{cite web |url = http://nationalacademies.org/evolution/ |title = Evolution Resources from the National Academies |publisher = ] |access-date = May 30, 2011 }}
* {{cite web |url = http://evolution.berkeley.edu/ |title = Understanding Evolution: your one-stop resource for information on Evolution |publisher = University of California, Berkeley |accessdate = May 30, 2011 }} * {{cite web |url = http://evolution.berkeley.edu/ |title = Understanding Evolution: your one-stop resource for information on Evolution |publisher = University of California, Berkeley |access-date = May 30, 2011 }}
* {{cite web |url = https://www.nsf.gov/news/special_reports/darwin/textonly/index.jsp |title = Evolution of Evolution – 150 Years of Darwin's "On the Origin of Species" |publisher = ] |accessdate = May 30, 2011 }} * {{cite web |url = https://www.nsf.gov/news/special_reports/darwin/textonly/index.jsp |title = Evolution of Evolution – 150 Years of Darwin's "On the Origin of Species" |publisher = ] |access-date = May 30, 2011 }}
* – ], ] (August 2016). * – ], ] (August 2016).


;Experiments concerning the process of biological evolution ;Experiments concerning the process of biological evolution
* {{cite web |url = http://myxo.css.msu.edu/index.html |title = Experimental Evolution – Michigan State University|author = Lenski RE|authorlink=Richard Lenski|accessdate = July 31, 2013}} * {{cite web |url = http://myxo.css.msu.edu/index.html |title = Experimental Evolution – Michigan State University|author = Lenski RE|author-link=Richard Lenski|access-date = July 31, 2013}}
* , ''Proceedings of the National Academy of Sciences of the USA'' * , ''Proceedings of the National Academy of Sciences of the USA''


;Online lectures ;Online lectures
* {{cite web |url = http://www.molbio.wisc.edu/carroll/Fittest.html |title = The Making of the Fittest |author = Carroll SB |authorlink = Sean B. Carroll |accessdate = May 30, 2011 |deadurl = yes |archiveurl = https://web.archive.org/web/20110718090526/http://www.molbio.wisc.edu/carroll/Fittest.html |archivedate = July 18, 2011 |df = }} * {{cite web |url = http://www.molbio.wisc.edu/carroll/Fittest.html |title = The Making of the Fittest |author = Carroll SB |author-link = Sean B. Carroll |access-date = May 30, 2011 |url-status = dead |archive-url = https://web.archive.org/web/20110718090526/http://www.molbio.wisc.edu/carroll/Fittest.html |archive-date = July 18, 2011 }}
* {{cite web |url = http://oyc.yale.edu/ecology-and-evolutionary-biology/principles-of-evolution-ecology-and-behavior/ |title = Principles of Evolution, Ecology and Behavior |author = Stearns SC |authorlink = Stephen C. Stearns |accessdate = August 30, 2011 |deadurl = yes |archiveurl = https://web.archive.org/web/20110323050059/http://oyc.yale.edu/ecology-and-evolutionary-biology/principles-of-evolution-ecology-and-behavior |archivedate = March 23, 2011 |df = }} * {{cite web |url = http://oyc.yale.edu/ecology-and-evolutionary-biology/principles-of-evolution-ecology-and-behavior/ |title = Principles of Evolution, Ecology and Behavior |author = Stearns SC |author-link = Stephen C. Stearns |access-date = August 30, 2011 |url-status = dead |archive-url = https://web.archive.org/web/20110323050059/http://oyc.yale.edu/ecology-and-evolutionary-biology/principles-of-evolution-ecology-and-behavior |archive-date = March 23, 2011 }}


{{Outline footer}} {{Outline footer}}


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Latest revision as of 21:37, 9 October 2024

Overview of and topical guide to change in the heritable characteristics of organisms See also: Index of evolutionary biology articles and Evolution (disambiguation)

The following outline is provided as an overview of and topical guide to evolution:

A diagram showing the relationships among various groups of organisms
Part of a series on
Evolutionary biology
Darwin's finches by John Gould
Processes and outcomes
Natural history
History of evolutionary theory
Fields and applications
Social implications

In biology, evolution is change in the heritable characteristics of biological organisms over generations due to natural selection, mutation, gene flow, and genetic drift. Also known as descent with modification. Over time these evolutionary processes lead to formation of new species (speciation), changes within lineages (anagenesis), and loss of species (extinction). "Evolution" is also another name for evolutionary biology, the subfield of biology concerned with studying evolutionary processes that produced the diversity of life on Earth.

Fundamentals about evolution

Life timeline
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−4500 —–—–−4000 —–—–−3500 —–—–−3000 —–—–−2500 —–—–−2000 —–—–−1500 —–—–−1000 —–—–−500 —–—–0 — Water Single-celled life Photosynthesis Eukaryotes Multicellular life P
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 Arthropods MolluscsFlowersDinosaurs MammalsBirdsPrimatesH
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Earth formed
Earliest water
LUCA
Earliest fossils
LHB meteorites
Earliest oxygen
Pongola glaciation*
Atmospheric oxygen
Huronian glaciation*
Sexual reproduction
Earliest multicellular life
Earliest fungi
Earliest plants
Earliest animals
Cryogenian ice age*
Ediacaran biota
Cambrian explosion
Hirnantian glaciation*
Earliest tetrapods
Karoo ice age*
Earliest apes / humans
Quaternary ice age*
(million years ago)*Ice Ages


Introduction

Basic principles

  • Macroevolution – Evolution on a scale at or above the level of species
    • Speciation – Evolutionary process by which populations evolve to become distinct species
      • Natural speciation
        • Allopatric speciation – Speciation that occurs between geographically isolated populations
        • Peripatric speciation – speciation in which a new species is formed from an isolated smaller peripheral populationPages displaying wikidata descriptions as a fallback
        • Parapatric speciation – Speciation within a population where subpopulations are reproductively isolated
        • Sympatric speciation – Evolution of a new species from an ancestor in the same location
      • Artificial speciation
      • Hybrid speciation – Form of speciation involving hybridization between two different species
    • Despeciation – Loss of a species of animal due to its combining with another species
    • Anagenesis – Gradual evolutionary change in a species without splitting
    • Extinction – Termination of a taxon by the death of its last member
  • Microevolution – Change in allele frequencies that occurs over time within a population
    • Artificial selection – Breeding used to develop desired characteristics
    • Natural selection – Mechanism of evolution by differential survival and reproduction of individuals
      • Sexual selection – Mode of natural selection involving the choosing of and competition for mates
    • Mutation – Alteration in the nucleotide sequence of a genome
    • Gene flow – Transfer of genetic variation from one population to another
    • Genetic drift – Concept in genetics

Subfields

History

Evolutionary theory and modelling

See also Basic principles (above)

Population genetics

Evolutionary phenomena

Modelling

Taxonomy, systematics, and phylogeny

Fundamentals

Basic concepts of phylogenetics

  • Phylogenetic tree – Branching diagram of evolutionary relationships between organisms
  • Phylogenetic network – Graph used to visualize evolutionary relationships, including reticulation events
  • Long branch attraction – Systematic error in phylogenetics
  • Clade – Group of a common ancestor and all descendants
  • Grade – Non-monophyletic grouping of organisms united by morphological or physiological characteristics
  • Ghost lineage – Phylogenetic lineage that is inferred to exist but has no fossil record

Inference methods

Current topics

Group Traits

  • Symplesiomorphy – An ancestral character or trait state shared by two or more taxa
  • Apomorphy – Two concepts on heritable traitsPages displaying short descriptions of redirect targets
  • Synapomorphy – Two concepts on heritable traitsPages displaying short descriptions of redirect targets
  • Autapomorphy – Distinctive feature, known as a derived trait, that is unique to a given taxon

Group Types

  • Monophyly – Property of a group of including all taxa descendant from a common ancestral species
  • Paraphyly – Type of taxonomic group
  • Polyphyly – Property of a group not united by common ancestry

Evolution of biodiversity

Origin and evolutionary history of life

Evolution of organisms

Evolution of tetrapods

Evolution of other animals

Evolution of plants

Evolution of other taxa

Evolution of cells, organs, and systems

Evolution of molecules and genes

Evolution of behaviour

Evolution of other processes

Applications in other disciplines

Evolutionary issues

Controversy about evolution

Religious and philosophical views of evolution

Influence of evolutionary theory

Publications and organizations concerning evolution

Books

Journals

Organizations

Evolution scholars and researchers

Prominent evolutionary biologists

See also

External links

General information
Experiments concerning the process of biological evolution
Online lectures
Misplaced Pages outlines
Categories: