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Bolosauridae Temporal range: Latest Carboniferous or earliest Permian to Middle Permian, Asselian–268 PreꞒ Ꞓ O S D C P T J K Pg N
Life restoration of Belebey vegrandis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	†Parareptilia
Order:	†Procolophonomorpha
Node:	†Procolophoniformes
Family:	†Bolosauridae Cope, 1878

Bolosauridae is an extinct family of parareptiles known from the latest Carboniferous (Gzhelian) or earliest Permian (Asselian) to the early Guadalupian epoch (latest Roadian stage) of North America, China, Germany, Russia and France. The bolosaurids were unusual for their time period by being bipedal, the oldest known tetrapods to have been so. Their teeth suggest that they were herbivores. The bolosaurids were a rare group and died out without any known descendants. The following cladogram shows the phylogenetic position of the Bolosauridae, from Johannes Müller, Jin-Ling Li and Robert R. Reisz, 2008.

References

1. Marcello Ruta; Juan C. Cisneros; Torsten Liebrect; Linda A. Tsuji; Johannes Muller (2011). "Amniotes through major biological crises: faunal turnover among Parareptiles and the end-Permian mass extinction". Palaeontology. 54 (5): 1117–1137. doi:10.1111/j.1475-4983.2011.01051.x.
2. Jocelyn Falconnet (2012). "First evidence of a bolosaurid parareptile in France (latest Carboniferous-earliest Permian of the Autun basin) and the spatiotemporal distribution of the Bolosauridae". Bulletin de la Société Géologique de France. 183 (6): 495–508. doi:10.2113/gssgfbull.183.6.495.
3. Johannes Müller; Jin-Ling Li & Robert R. Reisz (2008). "A new bolosaurid parareptile, Belebey chengi sp. nov., from the Middle Permian of China and its paleogeographic significance". Naturwissenschaften. 95 (12): 1169–1174. doi:10.1007/s00114-008-0438-0. PMID 18726080.

v t e Sauropsida (Reptilia sensu lato)
Kingdom: Animalia Phylum: Chordata Superclass: Tetrapoda Clade: Amniota Tetrapodomorpha see Tetrapodomorpha Reptiliomorpha see Reptiliomorpha Sauropsida see below↓
Sauropsida Sauropsida †Australothyris †Erpetonyx †Microleter †Acleistorhinidae Acleistorhinus Carbonodraco Colobomycter Delorhynchus Feeserpeton Karutia Klastomycter †Bolosauridae Belebey Bolosaurus Eudibamus †Mesosauria Brazilosaurus Mesosaurus Stereosternum †Millerettidae Broomia Heleophilus Milleretta Millerosaurus Milleropsis Nanomilleretta †Procolophonia see Procolophonia Diapsida see below↓
Diapsida Diapsida †Adelosaurus? †Claudiosaurus †Cryptovaranoides †Dolerosaurus †Elachistosuchus †Eunotosaurus? †Feralisaurus †Heleosuchus †Kenyasaurus †Kudnu †Lanthanolania †Maiothisavros †Microzemiotes †Orovenator †Palacrodon †Palaeagama †Saurosternon †Sphodrosaurus †Younginidae Galesphyrus Thadeosaurus Youngina †Tangasauridae Acerosodontosaurus? Hovasaurus Tangasaurus †Weigeltisauridae Coelurosauravus Glaurung Rautiania Weigeltisaurus †Drepanosauromorpha Ancistronychus Avicranium Dolabrosaurus Drepanosaurus Hypuronector Kyrgyzsaurus Longisquama? Megalancosaurus Protoavis? Skybalonyx Unguinychus Vallesaurus †Kuehneosauridae Icarosaurus Idiosaura? Kuehneosaurus Kuehneosuchus Pamelina Rhabdopelix? †Choristodera see Choristodera †Ichthyosauromorpha see Ichthyosauromorpha †Sauropterygia see Sauropterygia †Thalattosauria see Thalattosauria Sauria (Crown-Reptilia) Lepidosauromorpha see Lepidosauromorpha Includes Lepidosauria includes rhynchocephalians and squamates Archelosauria Archosauromorpha see Archosauromorpha Includes Archosauria includes birds and crocodilians Pantestudines see Pantestudines Includes Testudines turtles
See also Possible sauropsids Araeoscelidia Captorhinidae Protorothyrididae Recumbirostra Varanopidae Other taxa Anapsida Captorhinida Eureptilia Parareptilia Lanthanosuchoidea Millerosauria Procolophonomorpha Ankyramorpha Neodiapsida Eosuchia Younginiformes Avicephala Eolacertilia Euryapsida
Category

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• Procolophonomorphs
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• Paleozoic reptiles of Asia
• Paleozoic reptiles of Europe
• Paleozoic reptiles of North America
• Prehistoric reptile families
• Taxa named by Edward Drinker Cope
• Permian reptile stubs