Crocidura - Misplaced Pages

Genus of mammals

Crocidura Temporal range: Late Miocene–recent PreꞒ Ꞓ O S D C P T J K Pg N

Greater white-toothed shrew, C. russula
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Eulipotyphla
Family:	Soricidae
Subfamily:	Crocidurinae
Genus:	Crocidura Wagler, 1832
Type species
Sorex leucodon (Hermann, 1780)
Species
See text.

The genus Crocidura is one of nine genera of the shrew subfamily Crocidurinae. Members of the genus are commonly called white-toothed shrews or musk shrews, although both also apply to all of the species in the subfamily. With over 180 species, Crocidura contains the most species of any mammal genus. The name Crocidura means "woolly tail", because the tail of Crocidura species are covered in short hairs interspersed with longer ones.

They are found throughout all tropical and temperate regions of the Old World, from South Africa north to Europe, and east throughout Asia, as far east as the Malay Archipelago. One species, the possibly extinct Christmas Island shrew (C. trichura), also inhabited Christmas Island. They likely originated in Africa or Asia Minor during the Miocene, spread to Europe by the early Pliocene, and spread to eastern Asia and the Mediterranean by the Pleistocene.

List of species

Further information: List of crocidurines

Extant species

Javan hidden shrew (C. abscondita)
Sanetti shrew (C. afeworkbekelei)
Cyrenaica shrew (C. aleksandrisi)
East African highland shrew (C. allex)
Andaman shrew (C. andamanensis)
Anhui white-toothed shrew (C. anhuiensis)
Annamite shrew (C. annamitensis)
Ansell's shrew (C. ansellorum)
Arabian shrew (C. arabica)
Jackass shrew (C. arispa)
Armenian shrew (C. armenica)
Asian gray shrew (C. attenuata)
Hun shrew (C. attila)
Southwest Peninsula white-toothed shrew (C. australis)
Bailey's shrew (C. baileyi)
Balete's white-toothed shrew (C. baletei)
Kinabalu shrew (C. baluensis)
Batak shrew (C. batakorum)
Mindanao shrew (C. beatus)
Beccari's shrew (C. beccarii)
Bottego's shrew (C. bottegi)
Bale shrew (C. bottegoides)
Short-tailed white-toothed shrew (C. brevicauda)
Thick-tailed shrew (C. brunnea)
Buettikofer's shrew (C. buettikoferi)
African dusky shrew (C. caliginea)
Canarian shrew (C. canariensis)
Caspian shrew (C. caspica)
Thick-tailed Sulawesi white-toothed shrew (C. caudicrassa)
Sulawesi hairy-tailed shrew (C. caudipilosa)
Cinderella shrew (C. cinderella)
Congo white-toothed shrew (C. congobelgica)
Cranbrook's shrew (C. cranbooki)
Long-footed shrew (C. crenata)
Crosse's shrew (C. crossei)
Reddish-gray musk shrew (C. cyanea)
Dent's shrew (C. denti)
Desperate shrew (C. desperata)
Dhofar shrew (C. dhofarensis)
Long-tailed musk shrew (C. dolichura)
Dongjiangyuan white-toothed shrew (C. dongyangjiangensis)
Doucet's musk shrew (C. douceti)
Dracula shrew (C. dracula)
Dsinezumi shrew (C. dsinezumi)
Ivory Coast white-toothed shrew (C. eburnea)
Eisentraut's shrew (C. eisentrauti)
Elgon shrew (C. elgonius)
Elongated shrew (C. elongata)
Heather shrew (C. erica)
Fingui white-toothed shrew (C. fingui)
Fischer's shrew (C. fischeri)
Greater red musk shrew (C. flavescens)
Flower's shrew (C. floweri)
Bornean shrew (C. foetida)
Fox's shrew (C. foxi)
Southeast Asian shrew (C. fuliginosa)
Savanna shrew (C. fulvastra)
Smoky white-toothed shrew (C. fumosa)
Bicolored musk shrew (C. fuscomurina)
Gathorne's shrew (C. gathornei)
Glass's shrew (C. glassi)
Gmelin's white-toothed shrew (C. gmelini)
Goliath shrew (C. goliath)
Peters's musk shrew (C. gracilipes)
Large-headed forest shrew (C. grandiceps)
Greater Mindanao shrew (C. grandis)
Grasse's shrew (C. grassei)
Luzon shrew (C. grayi)
Greenwood's shrew (C. greenwoodi)
Güldenstädt's shrew (C. gueldenstaedtii)
Guy's shrew (C. guy)
Harenna shrew (C. harenna)
Sinharaja shrew (Crocidura hikmiya)
Hildegarde's shrew (C. hildegardeae)
Hill's shrew (C. hilliana)
Lesser red musk shrew (C. hirta)
Andaman spiny shrew (C. hispida)
Horsfield's shrew (C. horsfieldii)
Hutan shrew (C. hutanis)
Indochinese shrew (C. indochinensis)
Jackson's shrew (C. jacksoni)
Jenkins' shrew (C. jenkinsi)
Jouvenet's shrew (C. jouvenetae)
Katinka's shrew (C. katinka)
Kego shrew (Crocidura kegoensis)
Kivu shrew (C. kivuana)
Lamotte's shrew (C. lamottei)
Kivu long-haired shrew (C. lanosa)
Ussuri white-toothed shrew (C. lasiura)
Latona's shrew (C. latona)
Sulawesi shrew (C. lea)
Sumatran giant shrew (C. lepidura)
Bicolored shrew (C. leucodon)
Sulawesi tiny shrew (C. levicula)
Naked-tail shrew (C. littoralis)
Savanna swamp shrew (C. longipes)
Lucina's shrew (C. lucina)
Ludia's shrew (C. ludia)
Moonshine shrew(C. luna)
Mauritanian shrew (C. lusitania)
Lwiro shrew (C. lwiroensis)
MacArthur's shrew (C. macarthuri)
MacMillan's shrew (C. macmillani)
Nyiro shrew (C. macowi)
Malayan shrew (C. malayana)
Manenguba shrew (C. manengubae)
Makwassie musk shrew (C. maquassiensis)
Swamp musk shrew (C. mariquensis)
Gracile naked-tailed shrew (C. maurisca)
Javanese shrew (C. maxi)
Mduma's shrew (C. mdumai)
Central Sulawesi white-toothed shrew (C. mediocris)
Small elongated white-toothed shrew (C. microelongata)
Mindoro shrew (C. mindorus)
Sri Lankan long-tailed shrew (C. miya)
Kilimanjaro shrew (C. monax)
Sunda shrew (C. monticola)
Montane white-toothed shrew (C. montis)
Munissi’s shrew (C. munissii)
West African long-tailed shrew (C. muricauda)
Mossy forest shrew (C. musseri)
Ugandan musk shrew (C. mutesae)
Somali dwarf shrew (C. nana)
Savanna dwarf shrew (C. nanilla)
Narcondam shrew (C. narcondamica)
Sumatran white-toothed shrew (C. neglecta)
Peninsular shrew (C. negligens)
Negros shrew (C. negrina)
Newmark's shrew (C. newmarki)
Nicobar shrew (C. nicobarica)
Nigerian shrew (C. nigeriae)
Blackish white-toothed shrew (C. nigricans)
Black-footed shrew (C. nigripes)
African black shrew (C. nigrofusca)
Nimba shrew (C. nimbae)
Mount Nimba giant forest shrew (C. nimbasilvanus)
Sibuyan shrew (C. ninoyi)
Niobe's shrew (C. niobe)
Typical Sulawesi white-toothed shrew (C. normalis)
West African pygmy shrew (C. obscurior)
African giant shrew (C. olivieri)
West Sulawesi white-toothed shrew (C. ordinaria)
Oriental shrew (C. orientalis)
Ryukyu shrew (C. orii)
North African white-toothed shrew (C. pachyura)
Palawan shrew (C. palawanensis)
Pallid-footed white-toothed shrew (C. pallida')
Panay shrew (C. panayensis)
Sumatran long-tailed shrew (C. paradoxura)
Tiny Sulawesi white-toothed shrew (C. parva)
Small-footed shrew (C. parvipes)
Sahelian tiny shrew (C. pasha)
Pale gray shrew (C. pergrisea)
Guramba shrew (C. phaeura)
Dr. Phan Luong shrew (C. phanluongi))
Phu Hoc shrew (C. phuquocensis)
Cameroonian shrew (C. picea)
Pitman's shrew (C. pitmani)
Flat-headed shrew (C. planiceps)
Fraser's musk shrew (C. poensis)
Polia's shrew (C. polia)
North Peninsula white-toothed shrew (C. pseudorhoditis)
Kashmir white-toothed shrew (C. pullata)
Southern elongated white-toothed shrew (C. quasielongata)
Rainey's shrew (C. raineyi)
Negev shrew (C. ramona)
Chinese white-toothed shrew (C. rapax)
Egyptian pygmy shrew (C. religiosa)
Sulawesi white-handed shrew (C. rhoditis)
Roosevelt's shrew (C. roosevelti)
Greater white-toothed shrew (C. russula)
Sa Pa shrew (C. sapaensis)
Ugandan lowland shrew (C. selina)
Lesser rock shrew (C. serezkyensis)
Asian lesser white-toothed shrew (C. shantungensis)
Siberian shrew (C. sibirica)
Sicilian shrew (C. sicula)
Lesser gray-brown musk shrew (C. silacea)
Mebado white-toothed shrew (C. similturba)
Desert musk shrew (C. smithii)
Sokolov's shrew (C. sokolovi)
Somali shrew (C. somalica)
West-Central Sulawesi white-toothed shrew (C. solita)
Narrow-headed shrew (C. stenocephala)
Lesser white-toothed shrew (C. suaveolens)
Iranian shrew (C. susiana)
Taiwanese gray shrew (C. tanakae)
Tanzanian shrew (C. tansaniana)
Tarella shrew (C. tarella)
Saharan shrew (C. tarfayensis)
Telford's shrew (C. telfordi)
Dark white-toothed shrew (C. tenebrosa)
Timor shrew (C. tenuis)
Thalia's shrew (C. thalia)
Therese's shrew (C. theresae)
São Tomé shrew (C. thomensis)
Christmas Island shrew (C. trichura) (possibly extinct)
Turbo shrew (C. turba)
Ultimate shrew (C. ultima)
Javan ghost shrew (C. umbra)
Usambara shrew (C. usambarae)
Savanna path shrew (C. viaria)
Mamfe shrew (C. virgata)
Voi shrew (C. voi)
Voracious shrew (C. vorax)
Banka shrew (C. vosmaeri)
Lesser Ryukyu shrew (C. watasei)
Whitaker's shrew (C. whitakeri)
Wimmer's shrew (C. wimmeri)
Hainan Island shrew (C. wuchihensis)
Xanthippe's shrew (C. xantippe)
Beletta shrew (C. yaldeni)
Yankari shrew (C. yankariensis)
Mikhail Zaitsev’s shrew (C. zaitsevi)
Zaphir's shrew (C. zaphiri)
Zarudny's rock shrew (C. zarudnyi)
Upemba shrew (C. zimmeri)
Cretan shrew (C. zimmermanni)

Extinct species

†Crocidura balsamifera (known only from embalmed Crocidura mummies from Ancient Egypt)

Fossil species

†Crocidura abdallahi (Pleistocene of Morocco)
†Crocidura darelbeidae (Pleistocene of Morocco)
†Crocidura jaegeri (Pleistocene of Morocco)
†Crocidura kornfeldi (Pliocene to Pleistocene of Europe)
†Crocidura kapsominensis (Miocene of Kenya)
†Crocidura maghrebiana (Pleistocene of Morocco)
†Crocidura marocana (Pleistocene of Morocco)
†Crocidura tadjerensis (Pleistocene of Morocco)
†Crocidura thomasi (Pleistocene of Morocco)

A significant diversity of extinct Crocidura species is known from the early-mid Pleistocene of Morocco, but by the majority of these species went extinct between the Middle to Late Pleistocene boundary, and were replaced by modern species. Indeterminate Crocidura remains are known from the Miocene-aged rocks in the Potwar Plateau of Pakistan, concurrent with fossils from the Shivalik Fossil Beds.

Cultural significance

Crocidura shrews were embalmed in Ancient Egypt, being associated with the dark aspect of the god Horus. Many of these mummies have been uncovered during excavations at the Falcon Necropolis, providing important information about the former diversity of shrews in this area.

References

Hutterer, R. (2005). "Genus Crocidura". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 224–255. ISBN 978-0-8018-8221-0. OCLC 62265494.
Wilson, D. E.; Reeder, D. M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ Jenkins, Paulina D.; Darrin P. Lunde & Clive B. Moncrieff (2009). "Descriptions of New Species of Crocidura (Soricomorpha: Soricidae) from Mainland Southeast Asia, with Synopses of Previously Described Species and Remarks on Biogeography" (PDF). Bulletin of the American Museum of Natural History. 331: 356–405. doi:10.1206/582-10.1. S2CID 84803490.
Palmer, T.S. (1904). "Index generum mammalium : a list of the genera and families of mammals". USGS Report: 204. Bibcode:1904usgs.rept....4P.
"white-toothed shrew | mammal genus | Britannica". www.britannica.com. Retrieved 2022-02-20.
^ Rofes, Juan; Cuenca-Bescós, Gloria (2011-01-01). "Evolutionary history and biogeography of the genus Crocidura (Mammalia, Soricidae) in Europe, with emphasis on Crocidura kornfeldi". Mammalian Biology. 76 (1): 64–78. Bibcode:2011MamBi..76...64R. doi:10.1016/j.mambio.2009.12.001. ISSN 1616-5047.
^ Jacob A., Esselstyn; Anang S., Achmadi; Heru, Handika; Mark T., Swanson; Thomas C., Giarla; Kevin C., Rowe (2021-12-15). "Fourteen New, Endemic Species of Shrew (Genus Crocidura) from Sulawesi Reveal a Spectacular Island Radiation". Bulletin of the American Museum of Natural History. 454 (1): 1–108. doi:10.1206/0003-0090.454.1.1.
Meegaskumbura; et al. (2007-12-19). "Crocidura hikmiya, a new shrew (Mammalia: Soricomorpha: Soricidae) from Sri Lanka" (PDF). Zootaxa. 1665: 19–30. doi:10.11646/zootaxa.1665.1.2. Archived from the original (PDF) on 2016-03-03. Retrieved 2007-12-19.
Lunde, D.P.; Musser, G.G. & Ziegler, T. (2004). "Description of a new species of Crocidura (Soricomorpha: Soricidae, Crocidurinae) from Ke Go Nature Reserve, Vietnam". Mammal Study. 29: 27–36. doi:10.3106/mammalstudy.29.27.
Kamalakannan, Manokaran; Sivaperuman, Chandrakasan; Kundu, Shantanu; Gokulakrishnan, Govindarasu; Venkatraman, Chinnadurai; Chandra, Kailash (2021-05-03). "Discovery of a new mammal species (Soricidae: Eulipotyphla) from Narcondam volcanic island, India". Scientific Reports. 11 (1): 9416. Bibcode:2021NatSR..11.9416K. doi:10.1038/s41598-021-88859-4. ISSN 2045-2322. PMC 8093265. PMID 33941819.
Esselstyn, J.A.; Goodman, S.M. (2010). "New species of shrew (Soricidae: Crocidura) from Sibuyan Island, Philippines". Journal of Mammalogy. 91 (6): 1467–1472. doi:10.1644/10-MAMM-A-002.1.
Jenkins, P.; Abramov, A.; Bannikova, А.; Rozhnov, V. (2013). "Bones and genes: Resolution problems in three Vietnamese species of Crocidura (Mammalia, Soricomorpha, Soricidae) and the description of an additional new species". ZooKeys (313): 61–79. Bibcode:2013ZooK..313...61J. doi:10.3897/zookeys.313.4823. PMC 3701231. PMID 23840165.
Jenkins, P. D.; Abramov, A. V.; Rozhnov, V. V.; Makarova, O. V. (2007-09-19). "Description of two new species of white-toothed shrews belonging to the genus Crocidura (Soricomorpha: Soricidae) from Ngoc Linh Mountain, Vietnam" (PDF). Zootaxa. 1589: 57–68. doi:10.11646/zootaxa.1589.1.5.
Demos, T.C.; Achmadi, A.S.; Handika, H.; Maharadatunkamsi; Rowe, K.C.; Esselstyn, J.A. (2016). "A new species of shrew (Soricomorpha: Crocidura) from Java, Indonesia: possible character displacement despite interspecific gene flow". Journal of Mammalogy: gyw183. doi:10.1093/jmammal/gyw183.
Jenkins, P. D.; Abramov, A. V.; Rozhnov, V. V.; Makarova, O. V. (2007-09-19). "Description of two new species of white-toothed shrews belonging to the genus Crocidura (Soricomorpha: Soricidae) from Ngoc Linh Mountain, Vietnam" (PDF). Zootaxa. 1589: 57–68. doi:10.11646/zootaxa.1589.1.5.
^ Woodman, Neal; Ikram, Salima; Rowland, Joanne (2021-04-07). "A new addition to the embalmed fauna of ancient Egypt: Güldenstaedt's White-toothed Shrew, Crocidura gueldenstaedtii (Pallas, 1811) (Mammalia: Eulipotyphla: Soricidae)". PLOS ONE. 16 (4): e0249377. Bibcode:2021PLoSO..1649377W. doi:10.1371/journal.pone.0249377. ISSN 1932-6203. PMC 8026016. PMID 33826664.
^ Cornette, Raphaël; Stoetzel, Emmanuelle; Baylac, Michel; Moulin, Sibyle; Hutterer, Rainer; Nespoulet, Roland; El Hajraoui, Mohammed Abdeljalil; Denys, Christiane; Herrel, Anthony (2015-10-15). "Shrews of the genus Crocidura from El Harhoura 2 (Témara, Morocco): The contribution of broken specimens to the understanding of Late Pleistocene–Holocene palaeoenvironments in North Africa". Palaeogeography, Palaeoclimatology, Palaeoecology. 436: 1–8. Bibcode:2015PPP...436....1C. doi:10.1016/j.palaeo.2015.06.020. ISSN 0031-0182.
Mein, Pierre; Pickford, Martin (2006). "Late Miocene micromammals from the Lukeino Formation (6.1 to 5.8 Ma), Kenya". Publications de la Société Linnéenne de Lyon. 75 (4): 183–223. doi:10.3406/linly.2006.13628.
Flynn, Jacobs, Kimura, Taylor & Tomida (March 2020). "SIWALIK FOSSIL SORICIDAE: A CALIBRATION POINT FOR THE MOLECULAR PHYLOGENY OF SUNCUS". Paludicola. 12 (4): 247–258.{{cite journal}}: CS1 maint: multiple names: authors list (link)

Extant species of Eulipotyphla

Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Infraclass: Eutheria
Superorder: Laurasiatheria

Family: Erinaceidae
subfamily: Erinaceinae

Atelerix	Four-toed hedgehog (A. albiventris) North African hedgehog (A. algirus) Southern African hedgehog (A. frontalis) Somali hedgehog (A. sclateri)
Erinaceus	Amur hedgehog (E. amurensis) Southern white-breasted hedgehog (E. concolor) West European hedgehog (E. europaeus) Northern white-breasted hedgehog (E. roumanicus)
Hemiechinus	Long-eared hedgehog (H. auritus) Indian long-eared hedgehog (H. collaris)
Mesechinus	Daurian hedgehog (M. dauuricus) Hugh's hedgehog (M. hughi) Gaoligong forest hedgehog (M. wangi) Small-toothed forest hedgehog (M. miodon) Eastern forest hedgehog (M. orientalis)
Paraechinus	Desert hedgehog (P. aethiopicus) Brandt's hedgehog (P. hypomelas) Indian hedgehog (P. micropus) Bare-bellied hedgehog (P. nudiventris)

Family: Erinaceidae
subfamily: Galericinae

Echinosorex	Moonrat (E. gymnura)
Hylomys	Dwarf gymnure (H. parvus) Javan short-tailed gymnure (H. suillus) Bornean short-tailed gymnure (H. dorsalis) Max's short-tailed gymnure (H. maxi) Dalat gymnure (H. macarong) Northern short-tailed gymnure (H. peguensis) Leuser gymnure (H. vorax)
Neohylomys	Hainan gymnure (N. hainanensis)
Neotetracus	Shrew gymnure (N. sinensis)
Otohylomys	Long-eared gymnure (O. megalotis)
Podogymnura	Dinagat gymnure (P. aureospinula) Mindanao gymnure (P. truei)

Family: Soricidae
subfamily: Crocidurinae

Crocidura
(White-toothed
shrews)

Family: Soricidae
subfamily: Crocidurinae (continued)

Diplomesodon	Piebald shrew (D. pulchellus)
Feroculus	Kelaart's long-clawed shrew (F. feroculus)
Palawanosorex	Palawan moss shrew (P. muscorum)
Paracrocidura (Large-headed shrews)	Grauer's large-headed shrew (P. graueri) Greater large-headed shrew (P. maxima) Lesser large-headed shrew (P. schoutedeni)
Ruwenzorisorex	Ruwenzori shrew (R. suncoides)
Scutisorex	Hero shrew (S. somereni) Thor's hero shrew (S. thori)
Solisorex	Pearson's long-clawed shrew (S. pearsoni)
Suncus	Taita shrew (S. aequatorius) Black shrew (S. ater) Day's shrew (S. dayi) Etruscan shrew (S. etruscus) Sri Lankan shrew (S. fellowesgordoni) Bornean pygmy shrew (S. hosei) Least dwarf shrew (S. infinitesimus) Greater dwarf shrew (S. lixus) Madagascan pygmy shrew (S. madagascariensis) Malayan pygmy shrew (S. malayanus) Climbing shrew (S. megalura) Flores shrew (S. mertensi) Asian highland shrew (S. montanus) Asian house shrew (S. murinus) Remy's pygmy shrew (S. remyi) Anderson's shrew (S. stoliczkanus) Lesser dwarf shrew (S. varilla) Jungle shrew (S. zeylanicus)
Sylvisorex (Forest shrews)	Dudu Akaibe's pygmy shrew (S. akaibei) Cameroonian forest shrew (S. cameruniensis) Corbet's forest shrew (S. corbeti) Grant's forest shrew (S. granti) Howell's forest shrew (S. howelli) Bioko forest shrew (S. isabellae) Johnston's forest shrew (S. johnstoni) Kongana shrew (S. konganensis) Moon forest shrew (S. lunaris) Mount Cameroon forest shrew (S. morio) Greater forest shrew (S. ollula) Lesser forest shrew (S. oriundus) Rain forest shrew (S. pluvialis) Volcano shrew (S. vulcanorum)

Family: Soricidae
subfamily: Soricinae

Anourosoricini

Anourosorex (Asian mole shrews)	Assam mole shrew (A. assamensis) Giant mole shrew (A. schmidi) Chinese mole shrew (A. squamipes) Taiwanese mole shrew (A. yamashinai)

Blarinellini

Blarinella (some Asiatic short-tailed shrews)	Asiatic short-tailed shrew (B. quadraticauda) Burmese short-tailed shrew (B. wardi)
Parablarinella (some Asiatic short-tailed shrews)	Indochinese short-tailed shrew (P. griselda) Anhui short-tailed shrew (P. latimaxillata)

Blarinini

Blarina
(American short-tailed
shrews)

Cryptotis
(Small-eared
shrews)

C. mexicana group: Mexican small-eared shrew (C. mexicana); Nelson's small-eared shrew (C. nelsoni); Grizzled Mexican small-eared shrew (C. obscura); Phillips' small-eared shrew (C. phillipsii)

C. goldmani set: Central Mexican broad-clawed shrew (C. alticola); Goldman's broad-clawed shrew (C. goldmani); Goodwin's broad-clawed shrew (C. goodwini); Guatemalan broad-clawed shrew (C. griseoventris); C. lacertosus; C. mam; Oaxacan broad-clawed shrew (C. peregrina)

C. nigrescens group: Eastern Cordillera small-footed shrew (C. brachyonyx); Colombian small-eared shrew (C. colombiana); Honduran small-eared shrew (C. hondurensis); Yucatan small-eared shrew (C. mayensis); Darién small-eared shrew (C. mera); Merriam's small-eared shrew (C. merriami); Blackish small-eared shrew (C. nigrescens)

C. thomasi group: Southern Colombian small-eared shrew (C. andinus); Ecuadorian small-eared shrew (C. equatoris); Rainer's small-eared shrew (C. huttereri); Medellín small-eared shrew (C. medellinia); Merida small-eared shrew (C. meridensis); Wandering small-eared shrew (C. montivaga); Peruvian small-eared shrew (C. peruviensis); Scaly-footed small-eared shrew (C. squaipes); Tamá small-eared shrew (C. tamensis); Thomas's small-eared shrew (C. thomasi)

C. parva group: Central American least shrew (C. orophila); North American least shrew (C. parva); Tropical small-eared shrew (C. tropicalis)

Ungrouped / relict: Enders's small-eared shrew (C. endersi); Talamancan small-eared shrew (C. gracilis); Big Mexican small-eared shrew (C. magna)

Nectogalini

Chimarrogale (Asiatic water shrews)	Malayan water shrew (C. hantu) Himalayan water shrew (C. himalayica) Bornean water shrew (C. phaeura) Japanese water shrew (C. platycephalus) Chinese water shrew (C. styani) Sumatran water shrew (C. sumatrana)
Chodsigoa	Van Sung's shrew (C. caovansunga) De Winton's shrew (C. hypsibia) Lamulate shrew (C. lamula) Lowe's shrew (C. parca) Pygmy brown-toothed shrew (C. parva) Salenski's shrew (C. salenskii) Smith's shrew (C. smithii) Lesser Taiwanese shrew (C. sodalis)
Episoriculus	Hodgsons's brown-toothed shrew (E. caudatus) Taiwanese brown-toothed shrew (E. fumidus) Long-tailed brown-toothed shrew (E. leucops) Long-tailed mountain shrew (E. macrurus)
Nectogale	Elegant water shrew (N. elegans)
Neomys	Mediterranean water shrew (N. anomalus) Eurasian water shrew (N. fodiens) Transcaucasian water shrew (N. teres)
Soriculus	Himalayan shrew (S. nigrescens)

Notiosoricini

Megasorex	Mexican shrew (M. gigas)
Notiosorex	Cockrum's gray shrew (N. cockrumi) Crawford's gray shrew (N. crawfordi) Large-eared gray shrew (N. evotis) Villa's gray shrew (N. villai)

Family: Soricidae
subfamily: Soricinae (tribe: Soricini)

Sorex
(Long-tailed
shrews)

Subgenus
Otisorex

S. vagrans complex: Glacier Bay water shrew (S. alaskanus); Baird's shrew (S. bairdii); Marsh shrew (S. bendirii); Montane shrew (S. monticolus); New Mexico shrew (S. neomexicanus); Pacific shrew (S. pacificus); American water shrew (S. palustris); Fog shrew (S. sonomae); Vagrant shrew (S. vagrans)

S. cinereus group: Kamchatka shrew (S. camtschatica); Cinereus shrew (S. cinereus); Prairie shrew (S. haydeni); Saint Lawrence Island shrew (S. jacksoni); Paramushir shrew (S. leucogaster); Southeastern shrew (S. longirostris); Mount Lyell shrew (S. lyelli); Portenko's shrew (S. portenkoi); Preble's shrew (S. preblei); Pribilof Island shrew (S. pribilofensis); Olympic shrew (S. rohweri); Barren ground shrew (S. ugyunak)

Subgenus
Sorex

S. alpinus group: Alpine shrew (S. alpinus); Ussuri shrew (S. mirabilis)

S. araneus group: Valais shrew (S. antinorii); Common shrew (S. araneus); Udine shrew (S. arunchi); Crowned shrew (S. coronatus); Siberian large-toothed shrew (S. daphaenodon); Iberian shrew (S. granarius); Caucasian shrew (S. satunini)

S. arcticus group: Arctic shrew (S. arcticus); Maritime shrew (S. maritimensis)

S. tundrensis group: Tien Shan shrew (S. asper); Gansu shrew (S. cansulus); Tundra shrew (S. tundrensis)

S. minutus group: Buchara shrew (S. buchariensis); Kozlov's shrew (S. kozlovi); Caucasian pygmy shrew (S. volnuchini)

S. caecutiens group: Laxmann's shrew (S. caecutiens); Taiga shrew (S. isodon); Eurasian least shrew (S. minutissimus); Eurasian pygmy shrew (S. minutus); Flat-skulled shrew (S. roboratus); Shinto shrew (S. shinto); Long-clawed shrew (S. unguiculatus)

S. gracillimus group: Slender shrew (S. gracillimus)

S. raddei group: Radde's shrew (S. raddei)

S. samniticus group: Apennine shrew (S. samniticus)

incertae sedis

Family: Soricidae
subfamily: Myosoricinae

Congosorex (Congo shrews)	Phillips's shrew (C. phillipsorum) Greater Congo shrew (C. polli) Lesser Congo shrew (C. verheyeni)
Myosorex (Mouse shrews)	Babault's mouse shrew (M. babaulti) Montane mouse shrew (M. blarina) Bururi forest shrew (M. bururiensis) Dark-footed mouse shrew (M. cafer) Eisentraut's mouse shrew (M. eisentrauti) Geata mouse shrew (M. geata) Nyika mouse shrew (M. gnoskei) Kihaule's mouse shrew (M. kihaulei) Long-tailed forest shrew (M. longicaudatus) Oku mouse shrew (M. okuensis) Rumpi mouse shrew (M. rumpii) Schaller's mouse shrew (M. schalleri) Sclater's mouse shrew (M. sclateri) Thin mouse shrew (M. tenuis) Forest shrew (M. varius) Kilimanjaro mouse shrew (M. zinki)
Surdisorex (African mole shrews)	Aberdare mole shrew (S. norae) Mount Kenya mole shrew (S. polulus)

Family: Talpidae

Scalopinae
(New World moles
and relatives)

Condylura	Star-nosed mole (C. cristata)
Parascalops	Hairy-tailed mole (P. breweri)
Scalopus	Eastern mole (S. aquaticus)
Scapanulus	Gansu mole (S. oweni)
Scapanus (Western North American moles)	Broad-footed mole (S. latimanus) Coast mole (S. orarius) Townsend's mole (S. townsendii)

Talpinae
(Old World moles
and relatives)

Desmana	Russian desman (D. moschata)
Dymecodon	True's shrew mole (D. pilirostris)
Euroscaptor	Greater Chinese mole (E. grandis) Kloss's mole (E. klossi) Long-nosed mole (E. longirostris) Himalayan mole (E. micrura) Small-toothed mole (E. parvidens) Vietnamese mole (E. subanura)
Galemys	Pyrenean desman (G. pyrenaicus)
Mogera	Echigo mole (M. etigo) Insular mole (M. insularis) Kano mole (M. kanoana) Kobe mole (M. kobeae) Small Japanese mole (M. imaizumii) Large mole (M. robusta) Sado mole (M. tokudae) Japanese mole (M. wogura) Senkaku mole (M. uchidai)
Neurotrichus	American shrew mole (N. gibbsii)
Oreoscaptor	Japanese mountain mole (O. mizura)
Parascaptor	White-tailed mole (P. leucura)
Scaptochirus	Short-faced mole (S. moschatus)
Scaptonyx	Long-tailed mole (S. fusicaudus)
Talpa	Altai mole (T. altaica) Aquitanian mole (T. aquitania) Blind mole (T. caeca) Caucasian mole (T. caucasica) European mole (T. europaea) Père David's mole (T. davidiana) Levant mole (T. levantis) Martino's mole (T. martinorum) Spanish mole (T. occidentalis) Ognev's mole (T. ognevi) Roman mole (T. romana) Balkan mole (T. stankovici) Talysch mole (T. talyschensis)
Urotrichus	Japanese shrew mole (U. talpoides)

Uropsilinae
(Chinese shrew-like
moles)

Uropsilus	Equivalent-teeth shrew mole (U. aequodonenia) Anderson's shrew mole (U. andersoni) Black-backed shrew mole (U. atronates) Dabie Mountains shrew mole (U. dabieshanensis) Gracile shrew mole (U. gracilis) Inquisitive shrew mole (U. investigator) Snow Mountain shrew mole (U. nivatus) Chinese shrew mole (U. soricipes)

Family: Solenodontidae

Atopogale	Cuban solenodon (A. cubana)
Solenodon	Hispaniolan solenodon (S. paradoxus)

Taxon identifiers
Crocidura	Wikidata: Q935495 Wikispecies: Crocidura ADW: Crocidura AFD: Crocidura BOLD: 3064 CoL: 3W2V EoL: 13833 EPPO: 1KROKG Fauna Europaea: 305465 Fauna Europaea (new): f6773a8a-8a05-4fc6-ab85-55ebfd94f649 GBIF: 2435528 iNaturalist: 46557 IRMNG: 1036727 ITIS: 633452 MSW: 13700023 NBN: NBNSYS0000132032 NCBI: 36801 Open Tree of Life: 809501 Paleobiology Database: 40463

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