Paleobiota of the Green River Formation

Geological formation in the Intermountain West of the United States

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The Green River Formation is a geological formation located in the Intermountain West of the United States, in the states of Colorado, Utah, and Wyoming. It comprises sediments deposited during the Early Eocene in a series of large freshwater lakes: Lake Gosiute, Lake Uinta, and Fossil Lake (the last containing Fossil Butte National Monument). It preserves a high diversity of freshwater fish, birds, reptiles, and mammals, with some sections of the formation (including Fossil Lake and the Parachute Creek member of Lake Uinta) qualifying as Konservat-Lagerstätten due to their extremely well-preserved fossils.

Cartilaginous fish

Family	Genus	Species	Member	Abundance	Notes	Image
†Asterotrygonidae	†Asterotrygon	†A. maloneyi Carvalho, Maisey & Grande, 2004	Fossil Lake		An Asterotrygonidae stingray of uncertain higher affinities.
†Heliobatidae	†Heliobatis	†H. radians Marsh, 1877	Fossil Lake		A Heliobatid stingray of uncertain higher affinities. H. radians synonymy Dasyatis species Haseman, 1912 Palaeodasybatis discus Fowler, 1947 Xiphotrygon acutidens Cope, 1879 Xyphotrygus species Romer, 1971

Bony fish

Primarily based on Grande (2001), with changes where necessary:

Acipenseriformes

Family	Genus	Species	Member	Abundance	Notes	Image
Polyodontidae	†Crossopholis	†C. magnicaudatus Cope, 1883	Fossil Lake		A paddlefish.

Lepisosteiformes

Family	Genus	Species	Member	Notes
Lepisosteidae	Atractosteus	†A. atrox (Leidy, 1873)	Fossil Lake	An alligator gar relative. A. atrox is the largest Green River gar.
	Atractosteus	†A. simplex (Leidy 1873)	Fossil Lake Lake Gosiute?	An alligator gar relative.
	†Cuneatus	†C. cuneatus (Cope, 1884)	Fossil Lake? Lake Gosiute Lake Uinta	A Cuneatini tribe gar.
	†Cuneatus	†C. wileyi Grande, 2010	Lake Uinta	A Cuneatini tribe gar.
	Lepisosteus	†L. bemisi Grande, 2010	Fossil Lake	A Lepisosteini tribe gar.
	†Masillosteus	†M. janeae Grande, 2010	Fossil Lake	A Cuneatini tribe gar.

Amiiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Amiidae	Amia	†A. pattersoni Grande & Bemis, 1998	Fossil Lake Lake Gosiute		An amiid bowfin.
Amiidae	†Cyclurus	†C. gurleyi (Romer & Fryxell, 1928)	Fossil Lake	Only 8 specimens known	An amiid related to modern bowfins.

Hiodontiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Hiodontidae	Hiodon	†H. falcatus (Grande, 1979)	Fossil Lake		A hiodontid. First described as Eohiodon falcatus (1979).

Osteoglossiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Osteoglossidae	†Phareodus	†P. encaustus (Cope, 1871)	Fossil Lake Lake Gosiute		A phareodontine bonytongue.
Osteoglossidae	†Phareodus	†P. testis (Leidy, 1873)	Fossil Lake		A phareodontine bonytongue.

Ellimmichthyiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Armigatidae	†Diplomystus	†D. dentatus Cope, 1877	Fossil Lake Lake Gosiute Lake Uinta		An armigatid ellimmichthyiform.

Clupeiformes

Family	Genus	Species	Member	Abundance	Notes
Clupeidae	†Gosiutichthys	†G. parvus Grande, 1982	Lake Gosiute		A clupeine clupeid herring relative.
	†Knightia	†K. alta (Leidy, 1873)	Fossil Lake Lake Gosiute Lake Uinta		A pellonuline clupeid herring relative.
	†Knightia	†K. eocaena Jordan, 1907	Fossil Lake Lake Gosiute Lake Uinta	The most common articulated vertebrate fossil worldwide.	A pellonuline clupeid herring relative.

Gonorynchiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Gonorynchidae	†Notogoneus	†N. osculus Cope, 1885	Fossil Lake		A beaked sandfish. Last gonorynchid known from North America. Bottom feeding trace fossils are attributed to N. osculus.

Cypriniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Catostomidae	†Amyzon	†A. gosiutensis Grande, Eastman, & Cavender, 1982	Lake Gosiute		An ictiobine sucker.	A. aggregatum, a related species

Siluriformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Hypsidoridae	†Hypsidoris	†H. farsonensis Lundberg & Case, 1970	Lake Gosiute		A hypsidorid catfish.
Ictaluridae	†Astephus	†A. antiquus (Leidy, 1873)	Fossil Lake Lake Gosiute Lake Uinta		An ictalurid catfish.

Salmoniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Esocidae	Esox	†E. kronneri Grande, 1999	Fossil Lake	1 specimen	A pike or muskellunge species. Likely native to surrounding streams or rivers and not the lakes.

Percopsiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Percopsidae	†Amphiplaga	†A. brachyptera Cope, 1877	Fossil Lake		A trout-perch relative. Includes the jr synonym Erismatopterus endlichi (1877)
Percopsidae	†Erismatopterus	†E. levatus (Cope, 1870)	Lake Gosiute Lake Uinta		A trout-perch relative. First described as Cyprinodon levatus (1870) includes the jr synonym Erismatopterus rickseckeri (1871)

Carangiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Latidae	†Mioplosus	†M. labracoides Cope, 1877	Fossil Lake Lake Gosiute Lake Uinta		A lates perches relative.

Acanthuriformes

Family	Genus	Species	Member	Abundance	Notes
Moronidae	†Cockerellites	†C. liops (Cope, 1877)	Fossil Lake	common	A planktivorous/insectivorous temperate bass. First described as Priscacara liops (1877) First moved to Cockerellites liops in 1923.
	†Hypsiprisca	†H. hypsacantha (Cope, 1886)	Fossil Lake	rare	A temperate bass. First named Priscacara hypsacantha 1886)
	†Hypsiprisca	†H. sp.	Fossil Lake	rare	An undescribed species of Hypsiprisca. A slenderer species with a more rounded tail fin outline.
	†Priscacara	†P. serrata Cope, 1877	Fossil Lake Lake Uinta		A temperate bass.
		†P. sp. a	Fossil Lake	1 specimen	An undescribed Priscacara Specimens have a rounder body and weak dorsal spines.
		†P. sp. b.	Fossil Lake	1 specimen	An undescribed Priscacara Possibly a new species or a variant specimen with 4 anal spines.

Incertae sedis

Family	Genus	Species	Member	Abundance	Notes	Image
†Asineopidae	†Asineops	†A. squamifrons Cope, 1870	Fossil Lake Lake Gosiute Lake Uinta		An acanthomorph of uncertain affinities Possibly a percopsiform, polymixiiform, or perciform.
†Asineopidae	†Asineops	A. sp.	Fossil Lake		An undescribed Asineops species Restricted to Fossil Lake.

Amphibians

Frogs

Family	Genus	Species	Member	Abundance	Notes	Image
Pelodytidae	†Aerugoamnis	†A. paulus Henrici et al., 2013	Fossil Lake	2 specimens	A stem-relative of parsley frogs.

Salamanders

Family	Genus	Species	Member	Abundance	Notes	Image
Amphiumidae	†Paleoamphiuma	†P. tetradactylum Rieppel & Grande, 1998	Fossil Lake		A sirenid salamander.

Reptiles

Squamates

Family	Genus	Species	Member	Abundance	Notes
Boidae	†Boavus	†B. idelmani Gilmore, 1938	Fossil Lake		A boa.
Dactyloidae	†Afairiguana	†A. avius Conrad et al., 2007	Fossil Lake		A stem-anole.
Shinisauridae	†Bahndwivici	†B. ammoskius Conrad, 2006	Fossil Lake		A relative of the Chinese crocodile lizard.
Tropidophiidae?	†Dunnophis	D. sp.	Fossil Lake		An undescribed possible dwarf boa.
Varanidae	†Saniwa	†S. ensidens Leidy, 1870	Fossil Lake		A stem-monitor lizard.
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A possible gecko or iguanid Preserved with soft tissue and skin, but bones dissolved.
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A relative of Shinisaurids Preserved with soft tissue and skin, but bones dissolved.

Crocodilians

Family	Genus	Species	Member	Abundance	Notes
Alligatoridae	†Tsoabichi	†T. greenriverensis Brochu, 2010	Fossil Lake		A caiman.
Incertae sedis	†Borealosuchus	†B. wilsoni (Mook, 1959)	Fossil Lake		A eusuchian crocodyliform of uncertain affinities. Moved from Leidyosuchus wilsoni (1959)
	†Borealosuchus	Cf. †B. sp.	Fossil Lake		Indeterminate eusuchian bones close to Borealosuchus.
	"Crocodylus"	†"Crocodylus" acer Cope, 1882	Lake Uinta (Manti beds)	1 specimen	A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.
	"Crocodylus"	Cf. †"Crocodylus" affinis Cope, 1871	Lake Gosiute		Bones close in morphology to †"Crocodylus" affinis A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.

Turtles

Family	Genus	Species	Member	Abundance	Notes
Baenidae	†Chisternon	†C. undatum (Leidy, 1871)	Fossil Lake		A baenid.
Dermatemydidae	†Baptemys	†B. wyomingensis Leidy, 1873	Fossil Lake		A dermatemydid.
Dermatemydidae	†Baptemys	†B. Cf. wyomingensis Leidy, 1873	Lake Unita		A dermatemydid.
Geoemydidae	†Echmatemys	†E. haydeni (Cope, 1873)		1 specimen	A geoemydid tortoise.
		†E. naomi (Leidy, 1871)	Fossil Lake	1 specimen	A geoemydid tortoise.
		†E. wyomingensis Leidy, 1872	Fossil Lake		A geoemydid tortoise.
Testudinidae	†Hadrianus	†Cf. †H. corsoni (Leidy, 1871)	Fossil Lake		A tortoise species. A single complete hatchling
Trionychidae	†Axestemys	A. sp.	Fossil Lake		A softshell turtle.
	†Oliveremys	†O. uintaensis Cope, 1873	Fossil Lake		A softshell turtle.
	†Plastomenus	†P. thomasii (Cope, 1872	Fossil Lake		A softshell turtle.

Birds

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Lithornithiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Lithornithidae	†Calciavis	†C. grandei	Fossil Lake		A lithornithid.
†Lithornithidae	†Pseudocrypturus	†P. cercanaxius	Fossil Lake		A lithornithid.

Anseriformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Presbyornithidae	†Presbyornis	†P. pervetus	Fossil Lake		A presbyornithid anseriform.
Incertae sedis	†Paakniwatavis	†P. grandei	Fossil Lake		An anseriform of uncertain affinities.

Galliformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Gallinuloididae	†Gallinuloides	†G. wyomingensis	Fossil Lake		A gallinuloid galliform.

Coliiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Sandcoleidae	†Anneavis	†A. annae	Fossil Lake		A sandcoleid mousebird.

Leptosomiformes

Family	Genus	Species	Member	Abundance	Notes	Image
incertae sedis	†Plesiocathartes	P. major	Fossil Lake		A stem-cuckoo roller.
incertae sedis	†Plesiocathartes	†P. wyomingensis	Fossil Lake		A stem-cuckoo roller

Coraciiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Primobucconidae	†Primobucco	†P. mcgrewi	Fossil Lake		A primobucconid coraciiform.
Incertae sedis	†Paracoracias	†P. occidentalis			A coraciiform.

Piciformes

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Neanis	†N. schucherti	Fossil Lake		A possible piciform of uncertain affinities.
Incertae sedis	†Pristineanis	†P. kistneri			A possible piciform of uncertain affinities.

Strisores

Family	Genus	Species	Member	Notes
Eocypselidae	†Eocypselus	†E. rowei	Fossil Lake	A eocypselid apodiform.
Incertae sedis	†Fluvioviridavis	†F. platyrhamphus	Fossil Lake	A basal strisore, possibly a stem-frogmouth.
Incertae sedis	†Prefica	†P. nivea	Fossil Lake	A stem-oilbird.

Musophagiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Musophagidae?	†Foro	†F. panarium	Fossil Lake		A possible stem-turaco.

Mirandornithes

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Juncitarsus	†J. merkeli	Fossil Lake		A mirandornithalean, possibly an early phoenicopteriform.

Suliformes

Family	Genus	Species	Member	Abundance	Notes	Image
Fregatidae	†Limnofregata	L. azygosternon	Fossil Lake		An early frigatebird.
Fregatidae	†Limnofregata	L. hasegawai	Fossil Lake		An early frigatebird

Pelecaniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Threskiornithidae	†Rhynchaeites	†R. sp.		1 specimen	An undescribed ibis, known from a largely complete specimen.	R. messelensis, a related species
Threskiornithidae	†Vadaravis	†V. brownae	Fossil Lake		Potentially a stem-ibis, but may be an indeterminate aequornithine.

Charadriiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Indeterminate	Indeterminate	Indeterminate	Lake Uinta	1 trace specimen	A shorebird Potentially the earliest record of recurvirostrids Known from a 3 footprint trackway.

Gruiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Messelornithidae	†Messelornis	†M. nearctica	Fossil Lake		A messelornithid.	Specimen of a related species

Eufalconimorphae

Family	Genus	Species	Member	Notes
†Halcyornithidae	†Cyrilavis	†C. colburnorum	Fossil Lake	A halcyornithid.
†Halcyornithidae	†Cyrilavis	†C. olsoni	Fossil Lake	A halcyornithid.
†Messelasturidae	†Tynskya	†T. eocaena	Fossil Lake	A messelasturid.
†Psittacopedidae	†Eofringillirostrum	†E. boudreauxi	Fossil Lake	A psittacopedid.
†Quercypsittidae	†Avolatavis	†A. tenens	Fossil Lake	A quercypsittid.
†Zygodactylidae	†Eozygodactylus	†E. americanus	Fossil Lake	A zygodactylid.
†Zygodactylidae	†Zygodactylus	†Z. grandei	Fossil Lake	A zygodactylid.

Neoaves incertae sedis

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Nahmavis	†N. grandei	Fossil Lake	1 specimen	Either a stem-gruiform or a stem-charadriiform.

Two other genera, "Eoeurypyga" (a stem-sunbittern) and "Wyomingcypselus" (an early apodiform) are mentioned only in a 2002 dissertation, and are presently nomina nuda.

Avian ichnofossils

Family	Genus	Species	Member	Notes
†Avipedidae	†Avipeda	Cf. †A. phoenix Vyalov, 1960	Lake Unita	Avian trackways with three forward facing toes without inter-digit webbing.
†Ignotornidae	†Quadridigitus	†Q. semimembranus Lockley et al., 2021	Lake Unita	Avian trackways with three forward facing toes, a hallux, and inter-digit webbing.
†Jindongornipodidae	†Jindongornipes	†J. falkbuckleyi Lockley et al., 2021	Lake Unita	Large avian trackways with four toes, and wide spread between digits II and IV.
Incertae sedis	†Presbyornithiformipes	†P. feduccii Yang et al., 1995	Lake Uinta	Duck-like trackways with dabble marks attributed to Presbyornis.

Mammals

Partially based on Grande (1984). Aside from the few well-preserved mammals found in Fossil Lake, a majority of Green River mammals are based on isolated bones and teeth:

Metatheria

Family	Genus	Species	Member	Abundance	Notes	Image
†Herpetotheriidae	†Herpetotherium	†H. innominatum	Lake Gosiute Lake Uinta		An opossum-like basal metatherian Originally described as Peratherium innominatum.	H. huntii, a related species
†Herpetotheriidae	†Herpetotherium	†H. knighti	Lake Gosiute		An opossum-like basal metatherian.	H. huntii, a related species

Cimolesta

Family	Genus	Species	Member	Abundance	Notes	Image
†Pantolestidae	†Palaeosinopa	†P. didelphoides	Fossil Lake		A semi-aquatic pantolestid.

Chiroptera

Family	Genus	Species	Member	Abundance	Notes
†Icaronycteridae	†Icaronycteris	†I. gunnelli	Fossil Lake	2 specimens	An icaronycterid bat.
†Icaronycteridae	†Icaronycteris	†I. index	Fossil Lake	2 specimens	An icaronycterid bat.
†Onychonycteridae	†Onychonycteris	†O. finneyi	Fossil Lake	2 specimens	An onychonycterid bat.

Eulipotyphla

family	Genus	Species	Member	Abundance	Notes	Image
Nyctitheriidae	Nyctitherium	N. nitidum	Lake Gosiute		A nyctitheriid. Originally described as Talpavus nitidus.
Nyctitheriidae	Nyctitherium	N. sp.	Lake Uinta		A nyctitheriid. Originally described as Talpavus nitidus.

Pan-Carnivora

Family	Genus	Species	Member	Notes	Image
Miacidae	Vulpavus	V. profectus	Lake Gosiute	A miacid.
Miacidae	Indeterminate	indeterminate	Lake Gosiute Lake Uinta	Indeterminate miacid bones.
Sinopidae	Sinopa	S. minor	Lake Uinta	A sinopid hyaenodont.	S. rapax, a related species
Viverravidae	Viverravus	V. minutus	Lake Uinta	A viverravid.
Incertae sedis	"Miacis"	"M." gracilis	Lake Uinta	A carnivoraform of uncertain affinities.

Paraxonia

family	Genus	Species	Member	Abundance	Notes	Image
Mesonychidae	†Mesonyx?	Indeterminate	Lake Gosiute		Indeterminate mesonychid bones. First described as Mesonyx obtusidens Cope, 1872

Pan-Perissodactyla

family	Genus	Species	Member	Abundance	Notes
†Brontotheriidae	†Lambdotherium	†L. popoagicum	Fossil Lake		A small-sized basal brontothere.
†Brontotheriidae	†L. sp.	Lake Unita		A small-sized basal brontothere. Reported from Powder Wash, Utah by Cope (1880)
Equidae	†Protorohippus	†Aff. P. venticolus	Fossil Lake	2 specimens	An equid closely affiliated with P. venticolus.
Equidae	Indeterminate	Indeterminate	Lake Gosiute Lake Unita<	Indeterminate equiid bones. First identified as Hyracotherium sp.
†Hyopsodontidae	†Hyopsodus	†H. minusculus	Lake Gosiute Lake Uinta		A hyopsodontid.
		†H. paulus (=H. vicarius)	Lake Uinta		A hyopsodontid. includes H. vicarius specimens.
		†H. wortmani	Fossil Lake		A hyopsodontid.
↑Hyrachyidae	↑Hyrachyus	Indeterminate	Lake Unita		Undeterminate Hyrachyus bones.
Indeterminate	Indeterminate	Indeterminate	Fossil Lake	1 specimen	A juvenile tapiromorph, the largest mammal known from the formation. Initially described as Heptodon calciculus.

Apatotheria

Family	Genus	Species	Member	Abundance	Notes	Image
Apatemyidae	Apatemys	A. chardini Jepsen, 1930	Fossil Lake	1 specimen	An apatemyid.

Rodentia

Family	Genus	Species	Member	Notes	Image
Ischyromyidae	Paramys	P. sp.	Lake Uinta	A ischyromyid rodent.	P. delicatus
Ischyromyidae	Thisbemys	T. sp.	Lake Gosiute	An undescribed ischyromyid rodent.	T. plicatus
Sciuravidae	Sciuravus	S. eucristadens (Burke, 1937)	Lake Uinta (Douglas Creek Member)	A sciuravid rodent. Originally described as Viverravus eucristadens (1937).
Sciuravidae	Sciuravus	S. sp.	Lake Uinta	An undescribed sciuravid.

Primatomorpha

Family	Genus	Species	Member	Abundance	Notes	Image
Microsyopidae	Microsyops	M. elegans	Lake Gosiute		A microsyopid stem-primate.	M. cardiorestes, a related species
Microsyopidae	Uintasorex	U. parvulus	Lake Gosiute		A microsyopid stem-primate.
Notharctidae	Notharctus	N. sp.	Lake Gosiute		A notharctid adapiform.	N. tenebrosus
Notharctidae	Smilodectes	S. gracilis	Lake Uinta		A notharctid adapiform. Formerly included in Notharctus matthewi.
Omomyidae	Anemorhysis	A. sp.	Lake Unita	1 specimen (lost?)	An omomyid primate. First named as Uintalacus nettingi (1958) Deemed Anemorhysis sp indeterminate (1976)
	Omomys	O. carteri	Lake Uinta		An omomyid primate. Includes O. pucillus
	Tetonius	T. sp.	Lake Uinta		An omomyid primate.	T. homunculus
	Washakius	W. insignis	Lake Gosiute		An omomyid primate.

Arthropoda

Arachnida

Acariformes

Family	Genus	Species	Member	Abundance	Notes	Image
Cheyletidae	Indeterminate	Indeterminate	Lake Unita	1 specimen	A Chelytidae mite. Found in thin sectioning of rock samples.

Araneae

Family	Genus	Species	Member	Abundance	Notes
Araneidae	Araneus	A. sp	Lake Gosiute	6 specimens	Possible orbweb-spiders, known from partial fossils First placed as Epeira sp.
Hersiliidae	Hersiliola (sensu lato)	H. sp.	Lake Uinta (Parachute Creek Member)	1 male	A tree trunk spider.
Selenopidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	1 specimen	A female or juvenile selenopid spider.
Thomisidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	2 males, 1 juv female	Crab spiders.
Uloboridae	Miagrammopes	M. sp.	Lake Uinta (Parachute Creek Member)	1 female, 2 males	A uloborid spider.
Uloboridae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)		A uloborid spider.
Unidentified	†Araneaovoius (Scudder, 1878)	†A. columbiae		1 specimen	An orb-web spider egg sack. First described as "Aranea" columbiae (1878)

Parasitiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Ixodidae	Ixodes	†"I." tertiarius Scudder, 1885	Lake Uinta (Laney Member)	1 specimen	A punitive ixodid hard tick. Type specimen declared lost species considered nomen dubium by Dunlop (2011)

Scorpiones

Family	Genus	Species	Member	Abundance	Notes	Image
Buthidae	†Uintascorpio	†U. halandrasi Perry, 1995	Lake Uinta (Parachute Creek Member)	1 specimen	A buthid scorpion.

Crustacea

Branchiopoda

Family	Genus	Species	Member	Abundance	Notes	Image
Cyclestheriidae	†Cyclestherioides	†C. wyomingensis	Lake Gosiute (Laney)		A cyclestheriid clam shrimp.
Lynceidae	†Prolynceus	†P. laneyensis	Lake Gosiute (Laney)		A clam shrimp.

Decapods

Family	Genus	Species	Member	Abundance	Notes	Image
Palaemonidae	†Bechleja	†B. rostrata	Fossil Lake		A palaemonid shrimp.
Cambaridae	Procambarus	†P. primaevus	Fossil Lake		A cambarid crayfish.

Ostracoda

Family	Genus	Species	Member	Notes
Candonidae	Candona	C. artesensis Swain, 1964	Fossil Lake Lake Unita	A candonid ostracod.
		C. rangliensis Swain, 1964	Fossil Lake Lake Unita	A candonid ostracod.
		C. whitei Scott & Smith, 1951	Fossil Lake Lake Gosiute Lake Unita	A candonid ostracod.
	Tuberocypris	T. centronotus Swain, 1964	Lake Unita	A candonid ostracod.
†Cyprideidae	†Cypridea	†C. bisulcata Swain, 1949	Lake Unita	A cyprideid ostracod.
Cyprididae	†Advenocypris	†A. umbonata Swain, 1964	Lake unita	A cypridid ostracod.
	Hemicyprinotus	†H. watsonensis Swain, 1964	Lake unita	A cypridid ostracod. One described subspecies H. watsonensis emaciata
	Heterocypris	†H. riflensis Swain, 1964	Lake unita	A cypridid ostracod.
	Heterocypris	†H. whiteriverensis Swain, 1964	Lake unita	A cypridid ostracod.
	Potamocypris	†P. williamsi Swain, 1964	Fossil Lake Lake Unita	A cypridid ostracod.
	†Procyprois	†P. ravenridgensis Swain, 1964	Lake Unita	A cypridid ostracod. Male and female specimens show shell sexual dimorphism
	Pseudoeucypris	†P. pagei (Swain) Swain, 1956	Lake Unita	A cypridid ostracod. First described as Candona pagei (1949) Moved from Cypris pagei (1956)
	Scottia	†S. subquadrata Swain, 1964	Lake Unita	A cypridid ostracod.
	Stenocypris	Cf. S. sinuata G.W. Müller, 1898	Lake Unita	A cypridid ostracod.
Limnocytheridae	Limnocythere	L. sp.	Lake Unita	Indistinct limnocytherid fossils.
Limnocytheridae	Metacypris	†M. paracordata Swain, 1964	Lake Unita	A limnocytherid ostracod.
Notodromadidae	Cyprois	†C. ephraimensis Swain, 1964	Fossil Lake Lake Gosiute Lake Unita	A notodromadid ostracod.

Myriapoda

Family	Genus	Species	Member	Abundance	Notes	Image
Parajulidae	"Julus"	"J." telluster Scudder, 1878	Lake Gosiue	1 specimen	A parajulidae millapede in need of redescription. First described as Iulus telluster (1878).

Insecta

Coleoptera

Adephaga

Family	Genus	Species	Member	Abundance	Notes
Carabidae	Bembidium	†B. exoletum Scudder, 1876	Lake Unita	1 specimen	A Bembidiini tribe trechine ground beetle.
	†Carabites	†C. eocenicus Cockerell, 1921	Lake Unita		A ground beetle of uncertain subfamily affiliation.
	†Carabites	†C. exanimus Scudder, 1892	Lake Uinta	1 specimen	A ground beetle of uncertain subfamily affiliation.
	Harpalus	†H. veterum Cockerell, 1921	Lake Uinta		A Harpalini tribe harpaline ground beetle.
	Lebia	†L. protospiloptera Cockerell, 1921	Lake Unita		A Lebiini tribe harpaline ground beetle.
	†Neothanes	†N. testeus (Scudder, 1878)	Lake Gosiute	2 specimens	A ground beetle of uncertain subfamily affiliation. First named as Cychrus testeus (1878)
	Platynus	†P. caesus Scudder, 1890	Lake Gosiute	2 specimens	A Platynini tribe platynine ground beetle.
	Platynus	†P. senex Scudder, 1878	Lake Gosiute	1 specimen	A Platynini tribe platynine ground beetle.
Dytiscidae	Laccophilus	Indeterminate	Lake Uinta	1 fragment	An indeterminate Laccophilini tribe predaceous diving beetle.

Elateriformia

Family	Genus	Species	Member	Abundance	Notes
Byrrhidae	"Nosodendron"	†"N." tritavum Scudder, 1890	Lake Gosiute	1 specimen	A pill beetle in need of redescription. First named as a Nosodendrid beetle.
Elateridae	Cardiophorus	†C. exhumatus Cockerell, 1925	Lake Unita	1 specimen	A cardiophorine click beetle.
	Ctenicera	†C. velata (Scudder, 1876)	Lake Gosiute	1 specimen	A Prosternini tribe dendrometrine click beetle. First named as Corymbites velatus (1876).
	Oxygonus	†O. mortuus Scudder, 1876	Lake Uinta	1 specimen	A Prosternini tribe dendrometrine click beetle.
Eucnemidae	Epiphanis	†E. deletus Scudder, 1876	Lake Unita	1 specimen	An Epiphanini tribe melasine false click beetle.

Polyphaga - Bostrichiformia

Family	Genus	Species	Member	Abundance	Notes
Ptinidae	Anobium	†A. deceptum Scudder, 1878	Lake Gosiute	1 specimen	An Anobiini tribe anobiine death-watch beetle.
		†A. lignitum Scudder, 1878	Lake Gosiute	1 specimen	An Anobiini tribe anobiine death-watch beetle.
		†A. ovale Scudder, 1878	Lake Gosiute	2 specimens	An Anobiini tribe anobiine death-watch beetle.
	Stegobium	†S. defunctus (Scudder, 1876)	Lake Gosiute	1 specimen	A Stegobiini tribe anobiine death-watch beetle. First described as Sitodrepa defuncta (1876).

Polyphaga - Cucujiformia

Family	Genus	Species	Member	Abundance	Notes
Anthribidae	Choragus	†C. fictilis Scudder, 1890	Lake Gosiute	1 specimen	A Choragini tribe choragine fungus weevil.
	Euparius	†E. elusus (Scudder, 1878)	Lake Gosiute	2 specimens	A Cratoparini tribe anthribine fungus weevil. First named as Cratoparis? elusus (1878)
	Euparius	†E. repertus (Scudder, 1878)	Lake Gosiute	1 specimen	A Cratoparini tribe anthribine fungus weevil. First named as Cratoparis repertus (1878)
	Ormiscus	†O. partitus (Scudder, 1890)	Lake Gosiute	1 specimen	A Zygaenodini tribe anthribine fungus weevil. First named as Hormiscus partitus (1890)
	†Pseudochirotenon	†P. eocaenicus Legalov, 2018	Lake Unita	1 specimen	An Ecelonerini tribe anthribine fungus weevil.
	Trigonorhinus	†T. pristinus (Scuddr, 1876)	Lake Gosiute	1 specimen	A Trigonorhinini tribe anthribine fungus weevil. First named as Brachytarsus pristinus (1876)
	Tropideres	†T. remotus Scudder, 1893	Lake Gosiute	1 specimen	A Tropiderini tribe anthribine fungus weevil.
Attelabidae	†Teretrum	†T. quiescitum Scudder, 1893	Lake Unita	1 specimen	An Auletini tribe rhynchitine leaf-rolling weevil.
Belidae	†Paltorhynchus	†P. bisulcatus Scudder, 1893	Lake Unita	2 specimens	A Metrioxenini tribe oxycorynite belid weevil.
Brentidae	Apionion	†A. evestigatum (Scudder, 1893)	Lake Unita	1 specimen	An Apionini tribe apionine primitive weevil.
	†Palaeonanophyes	†P. zherikhini Legalov, 2015	Lake Unita		A Nanophyini tribe nanophyine primitive weevil.
	Perapion	†P. rasnitsyni Legalov, 2018	Lake Uinta	1 specimen	An Aplemonini tribe apionine primitive weevil.
Cerambycidae	Clytus	†C. pervetustus (Cockerell, 1921)	Lake Uinta		A Clytini tribe cerambycine long-horn beetle. First named as Lema pervetusta (1921)
Cerambycidae	Saperda	†S. caroli Vitali, 2015	Lake Unita		A Saperdini tribe lamiine long-horn beetle.
Chrysomelidae	Bruchus	†B. anilis Scudder, 1876	Lake Unita	1 specimen	A Bruchini tribe bruchine leaf beetle.
	Cryptocephalus	†C. vetustus Scudder, 1878	Lake Gosiute	2 specimens	A Cryptocephalini tribe cryptocephaline leaf beetle.
	†Denaeaspis	†D. chelonopsis Chaboo & Engel, 2009	Lake Unita	1 specimen	An Imatidiini tribe Cassidine tortoise beetle.
	†Eosacantha	†E. delocranioides Chaboo & Engel, 2009	Lake Unita	1 specimen	An Imatidiini tribe Cassidine tortoise beetle.
	†Palaeatalasis	†P. monrosi Legalov, 2021	Lake Unita	1 specimen	A Megamerini tribe sagrine leaf beetle.
	†Pulchritudo	†P. attenboroughi Krell & Vitali, 2021	Lake Uinta	1 specimen	A sagrine leaf beetle
Cryptophagidae	Antherophagus	†A. priscus Scudder, 1876	Lake Gosiute	4 specimens	A Cryptophagini tribe cryptophagine silken fungus beetle.
Curculionidae	Anthonomus	†A. revictus Scudder, 1890	Lake Gosiute	2 specimens	An Anthonomini tribe curculionine weevil.
	Anthonomus	†A. soporus Scudder, 1890	Lake Gosiute Lake Unita	7 specimens	An Anthonomini tribe curculionine weevil.
	†Archaeoheilus	†A. deleticius (Scudder, 1893)	Lake Uinta	1 specimen	A Molytini tribe molytine weevil. First named as Pachylobius deleticius (1893) Moved from Hylobius deleticius (2015)
		†A. ovalis Legalov, 2018	Lake Uinta	1 specimen	A Molytini tribe molytine weevil.
		†A. packardii (Scudder, 1893)	Lake Gosiute	1 specimen	A Molytini tribe molytine weevil. First named as Hylobius packardii (1893).
		†A. provectus (Scudder, 1876)	Lake Gosiute	2 specimens	A Molytini tribe molytine weevil. First named as Hylobius provectus (1876).
		†A. scudderi Legalov, 2018	Lake Uinta	1 specimen	A Molytini tribe molytine weevil.
	†Calandrites	†C. cineratius Scudder, 1893	Lake Gosiute Lake Unita	13 specimens	A weevil of uncertain subfamilial affiliation.
	†Calandrites	†C. defessus Scudder, 1893	Lake Gosiute Lake Unita	4 specimens	A weevil of uncertain subfamilial affiliation.
	Ceutorhynchus	†C. degravatus Scudder, 1893	Lake Unita	1 specimen	A Ceutorhynchini tribe ceutorhynchine weevil.
	"Coniatus"	†"C." refractus Scudder, 1893	Lake Uinta		A Tropiphorini tribe entimine weevil of uncertain genus affiliations.
	Cossonus	†C. rutus Scudder, 1893	Lake Unita	1 specimen	A Cossonini tribe cossonine weevil.
	Cryptorhynchus	†C. annosus Scudder, 1893)	Lake Gosiute Lake Unita	3 specimen	A Cryptorhynchini tribe cryptorhynchine weevil.
	Cryptorhynchus	†"C". durus Scudder, 1893)	Lake Unita	1 specimen	A Tropiphorini tribe entimine weevil of uncertain generic affiliations.
	"Dryocoetes"	†"D". carbonarius Scudder, 1876	Lake Gosiute	1 specimen	A scolytine bark beetle. Considered as genus and tribe incertae sedis by Legalov (2015).
	Entimus	†E. primordialis Scudder, 1876	Lake Uinta	1 specimen	An Entimini tribe entimine weevil.
	Epicaerus	†E. dilapsus (Scudder, 1893)	Lake Gosiute	1 specimen	A Geonemini tribe entimine weevil. First named as Phyxelis dilapsus (1893).
		†E. effossus (Scudder, 1876)	Lake Gosiute lake Uinta	numerous	A Geonemini tribe entimine weevil. First named as Eudiagogus effossus (1876).
		†E. eradicatus (Scudder, 1893)	Lake Unita	3 specimens	A Geonemini tribe entimine weevil. First named as Phyxelis eradicatus (1893).
		†E. evigoratus (Scudder, 1893)	Lake Unita	3 specimens	A Geonemini tribe entimine weevil. First named as Phyxelis evigoratus (1893).
		†E. exanimis (Scudder, 1876)	Lake Gosiute Lake Unita	numerous	A Geonemini tribe entimine weevil. First named as Eudiagogus exanimis (1876).
		†E. excissus (Scudder, 1893)	Lake Unita	1 specimen	A Geonemini tribe entimine weevil. First named as Phyxelis excissus (1893).
		†E. fodinarum (Scudder, 1876)	Lake Gosiute	3 specimens	A Geonemini tribe entimine weevil. First named as Sitona fodinarum (1893).
		†E. saxatilis (Scudder, 1876)	Lake Gosiute Lake Unita	numerous	A Geonemini tribe entimine weevil. First named as Eudiagogus saxatilis (1876).
		†E. subterraneus (Scudder, 1893)	Lake Gosiute Lake Unita	Numerous	A Geonemini tribe entimine weevil. First named as Scythropus subterraneus (1893).
		†E. terrosus (Scudder, 1878)	Lake Gosiute Lake Uinta	4 specimens	A Geonemini tribe entimine weevil. First named as Eudiagogus terrosus (1878).
	Eudiagogus	†E. vossi Legalov, 2018	Lake Uinta	1 specimen	A Eudiagogini tribe entimine weevil.
	†Extinctocentrinus	†E. brevirostris Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.
	Geraeus	†G. anvilis Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.
		†G. diruptus (Scudder, 1893)	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil. First named as Centrinus diruptus (1893)
		†G. fossilis Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.
	"Gymnetron"	†G. lecontei Scudder, 1878	Lake Gosiute	1 specimen	A curculionine weevil of uncertain genus and tribal affilliations.
	"Lepyrus"	†L. evictus (Scudder, 1893)	Lake Gosiute	1 specimen	A Tropiphorini tribe entimine weevil of uncertain genus affiliation.
	†Limalophus	†L. compositus (Scudder, 1893)	Lake Gosiute Lake Unita	3 specimens	A Tropiphorini tribe entimine weevil.
		†L. contractus (Scudder, 1893)	Lake Gosiute	6 specimens	A Tropiphorini tribe entimine weevil.
		†L. receptus (Scudder, 1893)	Lake Unita	1 specimen	A Tropiphorini tribe entimine weevil. First named as Artipus receptus (1893).
	Listronotus	†L. muratus Scudder, 1890	Lake Gosiute	1 specimen	A Listroderini tribe cyclomine weevil.
	†Lithogeraeus	†L. anicilla (Scudder, 1893)	Lake Unita	5 specimens	An Apostasimerini tribe conoderine weevil. First described as Aulobaris ancilla (1893)
		†L. circumscripta (Scudder, 1893)	Lake Unita	5 specimens	An Apostasimerini tribe conoderine weevil. First described as Aulobaris circumscripta (1893)
		†L. comminuta (Scudder, 1893)	Lake Unita	4 specimens	An Apostasimerini tribe conoderine weevil. First described as Aulobaris comminuta (1893)
		†L. greenriverensis Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	Mitostylus	†M. abacus (Scudder, 1893)	Lake Unita	1 specimen	A Sciaphilini tribe entimine weevil. First named as Scythropus abacus (1893).
		†M. obdurefactus (Scudder, 1893)	Lake Unita	5 specimens	A Sciaphilini tribe entimine weevil. First named as Exomias obdurefactus (1893).
		†M. seculorum (Scudder, 1890)	Lake Gosiute	1 specimen	A Sciaphilini tribe entimine weevil. First named as Tanymecus seculorum (1890).
	Neoptochus	Indeteminate	Lake Unita	2 specimens	Indeterminate Cyphicerini tribe Entiminae weevil fossils.
	Nicentrus	†N. curvirostris Legalov, 2018	Lake Uinta	1 specimen	An Apostasimerini tribe conoderine weevil.
	Ophryastes	†"O". compactus Scudder, 1878	Lake Gosiute	1 specimen	A Tropiphorini tribe entimine weevil. of uncertain genus affinities (2011, 2015).
		†O. grandis Scudder, 1893	Lake Unita	1 specimen	A Tropiphorini tribe entimine weevil. Considered nomen dubium (2015).
		†O. petrarum Scudder, 1893	Lake Unita	2 specimens	A Tropiphorini tribe entimine weevil. Considered nomen dubium (2011, 2015).
	†Ophryastites	†O. digressus Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain tribal affiliation.
	†Ophryastites	†O. dispertitus Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain tribal affiliation.
	†Otiorhynchites	†O. commutatus Scudder. 1893	Lake Unita	1 specimen	A weevil of uncertain subfamilial affiliations.
		†O. fossilis Scudder. 1893	Fossil Lake	1 specimen	A weevil of uncertain subfamilial affiliations.
		†O. tysoni Scudder. 1893	Lake Gosiute Lake Unita	2 specimens	A weevil of uncertain subfamilial affiliations.
	Otiorhynchus	†"O". flaccus Scudder, 1893	Lake Gosiute	1 specimen	An Otiorhynchini tribe entimine weevil. Considered genus and subfamily incertae sedis by Legalov (2015).
		†O. subteractus Scudder, 1893	Lake Unita	1 specimen	An Otiorhynchini tribe entimine weevil. Considered genus incertae sedis by Legalov (2015).
		†"O". tumbae (Scudder, 1890)	Lake Gosiute	1 specimen	An entimine weevil. Considered genus and tribus incertae sedis by Legalov (2015). First named Otiorhynchus dubius (1878)
	"Pachylobius"	†"P". compressus Scudder, 1893	Lake Gosiute Lake Unita	9 specimens	A weevil elytron of uncertain subfamily and genus affiliations.
	"Pachylobius"	†"P". depraedatus Scudder, 1893	Lake Unita	4 specimens	A weevil elytron of uncertain subfamily and genus affiliations.
	†Palaeomallerus	†P. longirostris Legalov, 2018	Lake Uinta	1 specimen	A Palaeomallerini tribe conoderine weevil.
	"Phyllobius"	†"P". antecessor Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain subfamily and genus affiliations.
		†"P". avus Scudder, 1893	Lake Gosiute Lake Unita	3 specimens	A weevil of uncertain subfamily and genus affiliations.
		†"P". carcerarius Scudder, 1893	Lake Unita	2 specimens	A weevil of uncertain subfamily and genus affiliations.
	"Polygraphus"	†"P". wortheni (Scudder, 1876)	Lake Gosiute	1 specimen	A scolytine bark beetle. Considered genus and tribus incertae sedis by Legalov (2015)
Procas	†P. vinculatus Scudder, 1893	Lake Unita	2 specimens	An Erirhinini tribe erirhinine weevil.
†Primocentron	†P. wickhami Legalov, 2018	Lake Uinta	1 specimen	An Eustylini tribe entimine weevil.
†Pseudophaops	†P. perditus (Scudder, 1876)	Lake Gosiute	2 specimens	A Eustylini tribe entimine weevil. First named Otiorhynchus perditus (1876)
†Pseudophaops	†P. wickhami Legalov, 2018	Lake Uinta		A Tropiphorini tribe entimine weevil.
"Rhyssomatus"	†"R". tabescens Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain generic affiliations.
†Sciabregma	†S. rugosa Scudder, 1893	Lake Unita	1 specimen	A Sciabregmini tribe molytine weevil.
†Sciabregma	†S. tenuicornis Cockerell, 1921	Lake Unita		A Sciabregmini tribe molytine weevil.
†Scythropus	†S. somniculosus Scudder, 1893	Lake Unita	1 specimen	A weevil of uncertain subfamilial affiliations.
"Sitona"	†"S". paginarum Scudder, 1893	Lake Gosiute Lake Unita	4 specimens	A Tropiphorini tribe entimine weevil of uncertain genus affiliations.
†Steganus	†S. barrandei Scudder, 1893	Lake Unita	1 specimen	A conoderine weevil.
†Syntomostylus	†S. rudis Scudder, 1893	Lake Unita	3 specimens	A weevil of uncertain subfamilial affiliations.
"Trypodendron"	†"T". impressus (Scudder, 1876)	Lake Gosiute	4 specimens	A scolytine bark beetle. Considered genus and tribus incertae sedis by Legalov (2015) Jr synonym includes Dryocoetes impressus (1878 in paleontology)
Erotylidae	Tritoma	†T. binotata Scudder, 1878	Lake Gosiute	1 specimen	A Tritomini tribe erotyline pleasing fungus beetle. First named as Mycotretus binotata (1878)
Laemophloeidae	Parandrita	†P. vestita Scudder, 1890	Lake Gosiute	4 specimens	A lined flat bark beetle.
Melandryidae	†Cicindelopsis	†C. eophilus Cockerell, 1921	Lake Unita		A false darkling beetle.
Mordellidae	†Petrimordella	†P. priscula (Cockerell, 1924)		1 specimen	A tumbling flower beetle morphospecies. First named as Mordella priscula (1925).
Mordellidae	†Petrimordella	†P. rasnitsyni (Odnosum & Perkovsky, 2016)		1 specimen	A tumbling flower beetle morphospecies. First named as Isotrilophus rasnitsyni (2016).
Nemonychidae	†Eocaenonemonyx	†E. kuscheli Legalov, 2013	Lake Unita		An Eocaenonemonychini tribe cretonemonychine pine flower weevil.
	†"Eugnamptus"	†"E". decemsatus Scudder, 1878	Lake Gosiute	1 specimen	An Eocaenonemonychini tribe cretonemonychine pine flower weevil. In need of redescription per Legalov (2013).
	†"Eugnamptus"	†"E". grandaevus (Scudder, 1876)	Lake Gosiute	3 specimens	An Eocaenonemonychini tribe cretonemonychine pine flower weevil. First named as Sitona grandaevus (1876) In need of redescription per Legalov (2013).
Peltidae	"Phenolia"	†"P." incapax Scudder, 1876	Lake Gosiute	1 specimen	A possible sheild beetle in need of redescription. First placed as a Nitidulid sap beetle.

Polyphaga - Scarabaeiformia

Family	Genus	Species	Member	Abundance	Notes	Image
Scarabaeidae	Aegialia	†A. rupta Scudder, 1890	Lake Gosiute	1 specimen	An aegialiine scaraboid beetle.
Scarabaeidae	Phyllophaga	†P. avus (Cockerell), Wickham, 1927	Lake Unita	1 specimen	A melolonthine scaraboid beetle. First described as Melolonthites avus (1921)

Polyphaga - Staphyliniformia

Family	Genus	Species	Member	Abundance	Notes
Hydrophilidae	Berosus	†B. sexstriatus Scudder, 1878	Lake Gosiute	1 specimen	A Berosini tribe hydrophiline water scavenger beetle.
	Berosus	†B. tenuis Scudder, 1878	Lake Gosiute	1 specimen	A Berosini tribe hydrophiline water scavenger beetle.
	Enochrus	†E. primaevus (Scudder, 1878)	Lake Gosiute	1 specimen	An enochrine water scavenger beetle. First named as Philhydrus primaevus (1878).
	Hydrobius	†H. confixus Scudder, 1890	Lake Gosiute	1 specimen	A Hydrobiusini tribe hydrophiline water scavenger beetle.
	Hydrobius	†H. decineratus Scudder, 1878	Lake Gosiute	1 specimen	A Hydrobiusini tribe hydrophiline water scavenger beetle.
	Hydrochus	†H. relictus Scudder, 1890	Lake Gosiute	1 specimen	A hydrochine water scavenger beetle.
	Laccobius	†L. elongatus Scudder, 1878	Lake Gosiute	1 specimen	A Laccobiini tribe hydrophiline water scavenger beetle.
	Tropisternus	†T. saxialis Scudder, 1878	Lake Gosiute	1 specimen	A Hydrophilini tribe hydrophiline water scavenger beetle.
	Tropisternus	†T. sculptilis Scudder, 1878	Lake Gosiute	1 specimen	A Hydrophilini tribe hydrophiline water scavenger beetle.
Staphylinidae	Bledius	†B. adamus Scudder, 1878	Lake Gosiute	1 specimen	A Blediini tribe oxyteline rove beetle.
	Bolitogyrus	Undescribed	Lake Unita	1 specimen	A Staphylinini tribe staphylinine rove beetle.
	Gyrophaena	†G. saxicola Scudder, 1876	Lake Unita		A Homalotini tribe aleocharine rove beetle.
	Homalota	†H. recisa Scudder, 1890	Lake Gosiute	2 specimens	A Homalotini tribe aleocharine rove beetle.
	Lathrobium	†L. abscessum Scudder, 1876	Lake Gosiute	7 specimens	A Lathrobiini tribe paederine rove beetle.
	Leistotrophus	†L. patriarchicus Scudder, 1876	Lake Unita		A Staphylinini tribe staphylinine rove beetle.
	Oxytelus	†O. pristinus Scudder, 1876	Lake Unita		A Oxytelini tribe oxyteline rove beetle.
	†Staphylinites	†S. obsoletum Scudder, 1876	Lake Gosiute	1 specimen	A rove beetle of uncertain subfamily affiliations.

Dictyoptera

Family	Genus	Species	Member	Abundance	Notes
Alienopteridae	†Chimaeroblattina	†C. brevipes Barna, 2018	Lake Unita		An alienopterid Dictyopteran.
Alienopteridae	†Grant	†G. viridifluvius Aristov, 2018	Lake Unita		An alienopterid Dictyopteran.
Blattidae	Periplaneta	†P.? smithiae Barna, Zelagin & Šmídová, 2023	Lake Uinta (Parachute Creek)	1 specimen	A possible periplanetean blattid cockroach.
Blaberidae	Diploptera	†D. gemini Barna, 2016	Lake Unita		A Diplopterine beetle cockroache.
		†D. savba Šmídová, 2016	Lake Unita		A Diplopterine beetle cockroache.
		†D. vladimir Vršanský, 2016	Lake Unita		A Diplopterine beetle cockroache.
Ectobiidae	Cariblattoides	†C. labandeirai Vršanský et al., 2012	Lake Uinta		A Pseudophyllodromiine wood cockroach.
Ectobiidae	Ectobius	†E. kohlsi Vršanský et al., 2014	Lake Unita		An ectobiine Ectobiid cockroach.

Diptera

Family	Genus	Species	Member	Abundance	Notes
Anisopodidae	Mycetobia	†"M." terricola (Scudder, 1878)	Lake Gosiute	1 specimen	A wood gnat. First described as Diadocidia? terricola Considered nomen dubium in (2019)
	†Olbiogaster	†O. simplex (Cockerell, 1921)	Lake Unita		A olbiogastrine wood gnat. First described as Eothereva simplex (1921)
	Sylvicola	†S. cadaver Scudder, 1890	Lake Gosiute	1 specimen	A anisopodine wood gnat.
Asilidae	Asilus	†A. palaeolestes Cockerell, 1921	Lake Unita		An Asilini tribe asiline robber fly.
Asilidae	†Stenocinclis	†S. anomala Scudder, 1878	Lake Gosiute	1 specimen	A dasypogonine robber fly.
Anthomyiidae	Anthomyia	†"A." winchesteri Cockerell, 1921	Lake Unita		First described as an anthomyiid fly. Deemed nomen dubium without evidence in 1996.
Bibionidae	Plecia	†P. akerionana Fitzgerald, 1999	Lake Unita (Parachute Creek Member)	1 specimen	A bibonid marchfly. The smallest species from the formation.
		†P. dejecta Scudder, 1890	Lake Gosiute	1 specimen	A bibonid marchfly.
		†P. pealei Scudder, 1890	Fossil Lake (Fossil Butte Member)	Most common insect in Fossil Lake	A bibonid marchfly.
		†P. rhodopterina Cockerell, 1925	Lake Unita		A bibonid marchfly.
		†P. winchesteri Cockerell, 1917	Lake Unita		A bibonid marchfly.
		†P. woodruffi Cockerell, 1916	Lake Unita		A bibonid marchfly.
Blephariceridae	†Philorites	†P. pallescens Cockerell, 1921	Lake Unita		A net-winged midge of uncertain subfamily affinities.
Cecidomyiidae	Lasioptera	†L. recessa Scudder, 1877	Lake Unita	1 specimen	A Lasiopterini tribe cecidomyiine gall midge.
Cecidomyiidae	†Lithomyza	†L. condita Scudder, 1877	Lake Unita	1 specimen	A Lestremiini tribe Lestremiine gall midge.
Chaoboridae	"Corethra"?	†"C".? exita Scudder, 1877	Lake Unita	1 specimen	A punitive phantom midge.
Chironomidae	Chironomus	†C. depletus Scudder, 1877	Lake Unita	1 specimen	A Chironomini tribe chironomine nonbiting midge.
		†C. patens Scudder, 1877	Lake Unita	1 specimen	A Chironomini tribe chironomine nonbiting midge.
		†C. septus Scudder, 1890	Lake Gosiute	1 specimen	A Chironomini tribe chironomine nonbiting midge.
	Undescribed	Undescribed	Lake Unita	Numerous	undescribed midge larvae.
Culicidae	Culex	†C. damnatorum Scudder, 1890	Lake Gosiute	3 specimens	A Culicini tribe culicine mosquito.
Culicidae	Culex	†C. winchesteri Cockerell, 1919	Lake Unita		A Culicini tribe culicine mosquito.
Cylindrotomidae	Cylindrotoma	†C. veterana Cockerell, 1921	Lake Unita		A cylindrotomine long bodied cranefly.
	†Cyttaromyia	†C. fenestrata Scudder, 1877	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
		†C. freiwaldi Cockerell, 1925	Lake Unita	2 specimens	A cylindrotomine long bodied cranefly species.
		†C. fuscula (Cockerell, 1921)	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species. First described as Asilopsis fusculus (1921).
		†C. gelhausi Kania-Kłosok et al., 2021	Lake Unita	3 specimens	A cylindrotomine long bodied cranefly species.
		†C. obdurescens Cockerell, 1925	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
		†C. reclusa Cockerell, 1924	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
Dolichopodidae	Dolichopus	indeterminate	Lake Gosiute	1 specimen	An indeterminate dolichopodine long-legged fly.
Empididae	†Protoedalea	†P. brachystoma Cockerell, 1921	Lake Uinta		A balloon fly of uncertain subfamily affinity.
Empididae	Rhamphomyia	†R. enena Cockerell, 1921	Lake Unita		An Empidini tribe empidine balloon fly.
Heleomyzidae	Heteromyza	†H. detecta Scudder, 1877	Lake Unita	1 specimen	A heleomyzine Heleomyzid fly.
Hippoboscidae	†Eornithoica	†E. grimaldii Nel, Garrouste & Engel, 2023	Fossil Lake (Fossil Butte Member)	1 specimen	A Hippoboscoidea fly Pupipara.
Keroplatidae	Palaeoplatyura	†P. eocenica Cockerell, 1921	Lake Uinta		An Orfeliini tribe keroplatine predatory fungus gnat.
Limoniidae	Dicranomyia	†D. primitiva Scudder, 1877	Lake Unita	4 specimens	A limoniine limoniid cranefly.
		†D. rostrata Scudder, 1877	Lake Unita	2 specimens	A limoniine limoniid cranefly. Includes Tipula decrepita (1877).
		†D. stigmosa Scudder, 1877	Lake Unita	6 specimens	A limoniine limoniid cranefly. Includes Tipula tecta (1877).
	†Pronophlebia	†P. rediviva Scudder, 1877	Lake Unita	1 specimen	A limoniid cranefly of uncertain subfamily placement.
	Rhabdomastix	†R. scudderi (Cockerell, 1921)	Lake Unita		A chioneine limoniid cranefly. First named as Gonomyia scudderi (1921).
Mycetophilidae	Anatella	†A. tacita Scudder, 1890	Lake Gosiute	1 specimen	An Exechiini tribe mycetophiline fungus gnat.
	Boletina	†B. paludivaga Scudder, 1890	Lake Gosiute	1 specimen	A gnoristine fungus gnat.
	Boletina	†B. umbratica Scudder, 1890	Lake Gosiute	1 specimen	A gnoristine fungus gnat.
	Gnoriste	†G. dentoni Scudder, 1877	Lake Unita	1 specimen	A gnoristine fungus gnat.
	Leptomorphus	†L. palaeospilus (Cockerell, 1921)	Lake Unita		A sciophiline fungus gnat. First named as Diomonus palaeospilus.
	Mycetophila	†M. occultata Scudder, 1877	Lake Unita	1 specimen	A Mycetophilini tribe Mycetophiline fungus gnat.
	Rymosia	†R. strangulata Scudder, 1890	Lake Gosiute	1 specimen	An Exechiini tribe mycetophiline fungus gnat.
	†Sackenia	†S. arcuata Scudder, 1877	Lake Unita	2 specimens	A Mycetophiline fungus gnat.
	†Sackenia	†S. gibbosa Cockerell, 1907	Lake Gosiute		A Mycetophiline fungus gnat.
	Sciophila	S. hyattii Scudder, 1890	Lake Gosiute	1 specimen	A sciophiline fungus gnat.
	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Mythicomyiidae	Acrocera	†A. hirsuta Scudder, 1877	Lake Unita	1 specimen	An acrocerine mythicomyiid fly. Possibly nomen dubium
Mythicomyiidae	Reissa	†R. kohlsi Evenhuis, 2019			A mythicomyiine mythicomyiid fly
Oestridae	Cuterebra	†C. ascarides (Scudder, 1877)	Lake Unita	Numerous	A cuterebrine bot fly species. First named as Musca ascarides (1877) Moved from Lithohypoderma ascarides (1916)
	Cuterebra	†C. bibosa (Scudder, 1877)	Lake Unita	Numerous	A cuterebrine bot fly species. First named as Musca bibosa (1877)
	Dermatobia	†D. hydropica (Scudder, 1877)	Lake Unita	3 specimens	A cuterebrine bot fly.
Platypezidae	Callomyia	†C. torporata Scudder, 1890	Lake Gosiute	1 specimen	A flat footed fly.
Stratiomyidae	†Lithophysa	†L. tumulta Scudder, 1890	Lake Gosiute	1 specimen	A soldier fly.
	Nemotelus	†N. eocenicus Cockerell, 1921	Lake Unita		A soldier fly.
	Odontomyia	Undescribed	Lake Unita	Numerous	undescribed soldier fly larvae.
	Sargus	†S. vetus Cockerell, 1921	Lake Unita		A soldier fly.
Psychodidae	"Culex"	†"C." proavitus Scudder, 1877	Lake Untia	1 specimen	A psychodid fly in need of redescription. First described as a mosquito (1877).
Sciaridae	Sciara	†S. deperdita Scudder, 1890	Lake Gosiute	1 specimen	A sciarine dark-winged fungus gnat.
Sciomyzidae	†Sciomyza?	†S.? manca Scudder, 1878	Lake Gosiute Lake Unita	Numerous	A Sciomyzini tribe sciomyzine marsh fly.
Sciomyzidae	†Sciomyza?	†S.? disjecta Scudder, 1878	Lake Gosiute	Numerous	A Sciomyzini tribe sciomyzine marsh fly.
Syrphidae	†Asarkina	†A. quadrata (Scudder, 1878)	Lake Gosiute	1 specimen	A Syrphini tribe syrphine hoverfly. First described as Milesia quadrata (1878)
	Cheilosia	†C. ampla Scudder, 1877	Lake Gosiute	3 specimens	A Rhingiini tribe eristaline hoverfly.
	Eristalis	†E. lapideus Scudder, 1877	Lake Unita	1 specimen	An Eristalini tribe eristaline hoverfly.
	Psilota	†P. tabidosa Scudder, 1890	Lake Gossiute	1 specimen	A Merodontini tribe eristaline hoverfly.
	Syrphus	†S. eocenicus Cockerell, 1925	Lake Uinta		A Syrphini tribe syrphine hoverfly.
	Syrphus	†S. lithaphidis Cockerell, 1921	Lake Unita		A Syrphini tribe syrphine hoverfly.
	Undescribed	Undescribed	Lake Unita	Numerous	undescribed hoverfly larvae.
Tabanidae	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Tachinidae	Tachina	indeterminate	Lake Gosiute	1 specimen	A Tachinini tribe tachinine Tachinid fly.
Tipulidae	†Spiladomyia	†S. simplex (Scudder, 1877)	Lake Unita	1 specimen	A tipulid cranefly of uncertain subfamily affinities.
	Tipula	†T. americana Kania et al., 2018	Lake Unita	1 specimen	A tipuline tipulid cranefly.
		†T. sepulchri Scudder, 1890	Lake Gosiute	1 specimen	A tipuline tipulid cranefly.
		†T. spoliata Scudder, 1890	Lake Gosiute	3 specimens	A tipuline tipulid cranefly.
Incertae sedis	†Acanthomyites	†A. aldrichi Cockerell, 1921	Lake Unita		A brachyceran fly of uncertain family affinities. First treated as a house fly.
	†Poliomyia	†P. recta Scudder, 1878	Lake Gosiute	1 specimen	A schizophoran fly of uncertain affinity.
	†Vinculomusca	†V. vinculata (Scudder, 1877)	Lake Unita	"Several"	A fly pupal case morphospecies of uncertain affinity. First named as Musca vinculata (1877)

Hemiptera

Family	Genus	Species	Member	Abundance	Notes
Aphididae	†?Siphonophoroides	†?S. antiqua Buckton, 1883	Lake Gosiute	1 specimen	A possible drepanosiphine aphid. First identified as possibly Lithaphis diruta (1890)
Cercopidae	Cercopis	†C. astricta Scudder, 1890	Lake Gosiute	1 specimen	A Cercopini tribe cercopine froghopper.
	Cercopis	†C. cephalinus Cockerell, 1921	Lake Unita		A Cercopini tribe cercopine froghopper.
	†Cercopites	†C. calliscens scudder, 1890	Lake Gosiute	1 specimen	A froghopper of uncertain subfamilial affiliation.
	†Cercopites	†C. umbratilis scudder, 1890	Lake Gosiute	1 specimen	A froghopper of uncertain subfamilial affiliation.
	†Palecphora	†P. patefacta Scudder, 1890	Lake Gosiute	1 specimen	A froghopper of uncertain subfamilial affiliation.
Cicadellidae	Aphrodes	†A. adae (SCudder, 1878)	Lake Gosiute	2 specimens	An Aphrodini tribe aphrodine leafhopper. First named as Acocephalus adae (1878).
	Cicadella	†C. parobtecta Dmitriev, 2020	Lake Unita		A Cicadellini tribe cicadelline leafhopper. A replacement name for Tettigonia obtecta (1877 non-Fabricius, 1803)
		†C. priscomarginata (Scudder, 1890)	Lake Gosiute	1 specimen	A Cicadellini tribe cicadelline leafhopper. First named as Tettigonia priscomarginata (1890).
		†C. priscovariegata Scudder, 1890	Lake Gosiute	1 specimen	A Cicadellini tribe cicadelline leafhopper. First named as Tettigonia priscovariegata (1890).
		†C. scudderi Cockerell, 1921	Lake Unita		A Cicadellini tribe cicadelline leafhopper.
	Cicadula	†C. saxosa Scudder, 1890	Lake Gosiute	1 specimen	A Cicadulini tribe deltocephaline leafhopper.
	Coelidia	†C. wyomingensis Scudder, 1890	Fossil Lake (Fossil Butte Member)	1 specimen	A Coelidiini tribe coelidiine leafhopper.
	Iassus	†I. lapidescens (Scudder, 1877)	Lake Uinta	1 specimen	An Iassini tribe iassine leafhopper. First named as Bythoscopus lapidescens (1877)
	Thamnotettix	†T. gannetti Scudder, 1890	Lake Gosiute	2 specimens	An Athysanini tribe deltocephaline leafhopper.
		†T. mutilata Scudder, 1890	Lake Gosiute	1 specimen	An Athysanini tribe deltocephaline leafhopper.
		†T. packardi Cockerell, 1925	Lake Unita		An Athysanini tribe deltocephaline leafhopper. First named as Erythroneura eocenica (1921)
Cixiidae	Cixius	†C.? hesperidum Scudder, 1878	Lake Gosiute	1 specimen	A possible Cixiini tribe cixiine cixiid planthopper. Considered nomen dubium by Swedo and Lefebvre (2004).
Cixiidae	†Oliarites	†O. terrentula (Scudder, 1878)	Lake Gosiute	1 specimen	A cixiine cixiid planthopper. First named as Mnemosyne terrentula (1878) Considered nomen dubium by Swedo and Lefebvre (2004).
Cydnidae	Cyrtomenus	†C. concinnus Scudder, 1878	Lake Gosiute	1 specimen	A Geotomini tribe cydnine burrowing bug.
	†Necrocydnus	†N. gosiutensis Scudder, 1890	Lake Gosiute	1 specimen	A burrowing bug of uncertain subfamilial affiliation.
	†Procydnus	†P. mamillanus (Scudder, 1878)	Lake Gosiute	1 specimen	A cydnine burrowing bug. First named as Cydnus? mamillanus (1878).
	†Stenopelta	↑S. punctulata (Scudder, 1878)	Lake Gosiute	5 specimens	A cydnine burrowing bug. First named as Aethus punctulatus (1878).
Delphacidae	Delphax	†D. veterum Cockerell, 1921	Lake Uinta		A Delphacini tribe delphacine delphacid froghopper
Dinidoridae	†Dinidorites	†D. margiformis Cockerell, 1921	Lake Unita		A dinidorid shield bug of uncertain subfamily affinities.
Fulgoridae	Aphaena	†A. atava Scudder, 1877	Lake Unita	1 specimen	A lanternfly of uncertain subfamily affiliations First named as Aphana atava (1877)
	Aphaena	†A. rotundipennis Scudder, 1878	Lake Gosiute	2 specimens	A lanternfly of uncertain subfamily affiliations First named as Aphana rotundipennis (1878)
	Fulgora	†F. granulosa Scudder, 1878	Lake Gosiute	1 specimen	A Fulgorini tribe fulgorine lanternfly.
	Fulgora	†F. populata Scudder, 1890	Lake Gosiute	1 specimen	A Fulgorini tribe fulgorine lanternfly.
	Lystra?	†L.? leei Scudder, 1878	Lake Gosiute	1 specimen	A Lystrini tribe poiocerine lanternfly.
	Lystra?	†L.? richardsoni Scudder, 1878	Lake Gosiute	11 specimens	A Lystrini tribe poiocerine lanternfly.
Gerridae	†Telmatrechus	†T. parallelus Scudder, 1890	Fossil Lake (Fossil Butte Member) Lake Uinta (Laney Member)	2 specimens	A water strider.
Lophopidae	†Scoparidea	†S. nebulosa Cockerell, 1921	Lake Unita		A lophopid planthopper.
Lyctocoridae	Lyctocoris	†L. terreus (Scudder, 1878)	Lake Gosiute	1 specimen	A Lyctocorini tribe lyctocorine Lyctocorid bug. First named as Rhyparochromus? terreus (1878)
Nogodinidae	†Detyopsis	†D. scudderi Cockerell, 1921	Lake Unita		A nogodinid planthopper.
	†Detyopsis	†D. scudderi Cockerell, 1921	Lake Unita		A nogodinid planthopper.
	†Ficarasites	†F. stigmaticum Scudder, 1890	Lake Gosiute	1 specimen	A nogodinid planthopper.
	†Lithopsis	†L. delicata Cockerell, 1921	Lake Unita		A nogodinid planthopper.
		†L. elongata Scudder, 1890	Lake Gosiute	1 specimen	A nogodinid planthopper.
		†L. fimbriata Scudder, 1878	Lake Gosiute	3 specimens	A nogodinid planthopper.
		†L. simillima Cockerell, 1921	Lake Unita		A nogodinid planthopper.
	"Paralatindia"	†"P." saussurei Scudder, 1890	Lake Gosiute	1 specimen	A nogodinid planthopper. First described as a Paralatindia genus cockroach. Considered nomen dubium by Shcherbakov (2006)
Reduviidae	†Aphelicophontes	†A. danjuddi Swanson et al., 2021	Lake Unita	1 specimen	A Harpactorini tribe Harpactorine Assassin bug.
Rhopalidae	Corizus	†C. guttatus (Scudder, 1878)	Lake Gosiute	2 specimens	A Rhopalini tribe rhopaline scentless plant bug. First named as Reduvius guttatus (1878).
Rhyparochromidae	Aphanus	†A. petrensis Scudder, 1877	Lake Unita	1 specimen	A Gonianotini tribe rhyparochromine seed bug. First named as Pachymerus petrensis (1877).
Rhyparochromidae	Cholula	†C. triguttata Scudder, 1890	Lake Gosiute	1 specimen	A Myodochini tribe rhyparochromine seed bug.
Ricaniidae	†Dilaropsis	†D. ornatus Cockerell, 1921	Lake Unita		A Ricaniid plant hopper.
	†Hammapteryx	†H.? ceryniiformis Cockerell, 1921	Lake Unita		A Ricaniid plant hopper.
		†H.? lepidoides Cockerell, 1921	Lake Unita		A Ricaniid plant hopper.
		†H. reticulata Scudder, 1890	Lake Gosiute	1 specimen	A Ricaniid plant hopper.
Tingidae	†Gyaclavator	†G. kohlsi Wappler et al., 2015	Lake Unita	4 specimens	A lace bug of uncertain subfamily affiliation.
Incertae sedis	"Delphax"	†"D." senilis Scudder, 1877	Lake Unita	1 specimen	First described as an delphacine froghopper. Considered nomen dubium
	†Diaplegma	†D. obdormitum Scudder, 1890	Lake Gosiute	1 specimen	A fulgoroid leafhopper of uncertain familial affinities. First placed as a cixiid planthopper.
	Emiliana	†E. alexandri Shcherbakov, 2006	Lake Unita	1 specimen	A fulgoroid leafhopper of uncertain familial affinities. First placed as a Tropiduchid froghopper
	†Eoliarus	†E. lutensis (Scudder, 1890)	Lake Gosiute	1 specimen	A fulgoroid leafhopper of uncertain familial affinities. First described as Oliarus lutensis (1890)
	†Eoliarus	†E. quadristictus Cockerell, 1925			A fulgoroid leafhopper of uncertain familial affinities.
	†Necygonus	†N. rotundatus Scudder, 1890	Lake Gosiute	1 specimen	A Pentatomomorph sheild bug of uncertain familial affiliation. First placed as a toad bug genus.
	†Protoliarus	†P. humatus Cockerell, 1921	Lake Uinta		A fulgoroid leafhopper of uncertain familial affinities. First described as a lanternfly.

Hymenoptera

Family	Genus	Species	Member	Abundance	Notes
Apidae	†Euglossopteryx	†E. biesmeijeri De Meulemeester, Michez & Engel, 2014	Lake Unita (Parachute Creek Member)		An apid bee.
Bethylidae	Eupsenella	E. sp.			An old world Bethylidae wasp.
Braconidae	Bracon	†B. laminarum Scudder, 1878	Lake Gosiute	1 specimen	A Braconini tribe braconine braconid wasp.
	†Colemanus	†C. keeleyorum Fisher, 2015	Lake Unita (Parachute Creek Member)		A Proteropini tribe ichneutine braconid wasp.
	†Eobracon	†E. cladurus Cockerell, 1921	Lake Unita		A braconid wasp of uncertain subfamily affinities.
	Hormiopterus	†H. petrinus (Scudder, 1877)	Lake Unita		A braconid wasp of uncertain subfamily affinities. First named as Ichneumon petrinus
Eurytomidae	Eurytoma	†E. antiqua (Scudder, 1878)	Lake Gosiute	2 specimens	An eurytomine Eurytomid chalcid wasp. First described as Decatoma antiqua (1878).
Formicidae	†Archimyrmex	†A. rostratus Cockerell, 1923	Lake Unita	Numerous	A myrmeciine ant.
	†Camponotites	†C. vetus (Scudder, 1877)	Green River Formation?	1 specimen	A formicine ant of uncertain affiliation. First described as Camponotus vetus (1877) The type locality was not specified by Scudder, and may Be G.R.F. or White River Formation.
	Dolichoderus	†D. kohlsi Dlussky & Rasnitsyn, 2003	Lake Uinta	Numerous	A Dolichoderus species ant.
	†Eoformica	†E. globularis Dlussky & Rasnitsyn, 2003	Lake Unita	2 specimens	An ant of unidentified affiliation.
		†E. magna Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	An ant of unidentified affiliation.
		†E. pinguis (Scudder, 1877)	Lake Gosiute Lake Uinta	Numerous	An ant of unidentified affiliation. Moved from Liometopum pingue (1877) Includes the jr synonyms Lasius terreus (1878) and Eoformica eocenica (1921)
		Indeterminate	Lake Unita	2 specimen	Two partial specimens that may be referable to Eoformica.
	†Kohlsimyrma	†K. gracilis Dlussky & Rasnitsyn, 2003	Lake Unita	2 specimens	An ant of uncertain affiliation.
		†K. laticeps Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	An ant of uncertain affiliation.
		†K. longiceps Dlussky & Rasnitsyn, 2003	Lake Unita	3 specimens	An ant of uncertain affiliation.
	†Mianeuretus	†M. eocenicus Dlussky & Rasnitsyn, 2003	Lake Unita	2 specimens	An aneuretine ant.
	†Mianeuretus	Indeterminate	Lake Unita	1 specimen	A partial specimen that may belong to Mianeuretus.
	†Myrmecites	†M. rotundiceps Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A myrmicine ant of uncertain affiliation.
	Pachycondyla	†P. labandeirai Dlussky & Rasnitsyn, 2003	Lake Gosiute	1 specimen	A Pachycondyla species ant.
	†Ponerites	†P. coloradensis Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
		†P. eocenicus Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
		†P. hypoponeroides Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
	†Proiridomyrmex	†P. vetulus Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A dolichoderine of uncertain affinity.
	†Proiridomyrmex	Indeterminate	Lake Unita	1 specimen	A partial specimen that may belong to Proiridomyrmex.
	†Solenopsites	†S. minutus Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A Solenopsis like ant of uncertain affiliation.
	†Titanomyrma	†T. lubei Archibald et al., 2011	Fossil Lake (Angelo Member)	3 specimens	A titan ant species.
	Indeterminate	Indeterminate	Lake Unita	1 specimen	A partial ant specimen that may belong to Formica.
	Indeterminate	Indeterminate	Lake Unita	3 specimens	Partial ant specimens of unidentified subfamily placement.
Ichneumonidae	Allomacrus	†A. lanhami (Cockerell, 1941)	Lake Unita		A cylloceriine darwin wasp. First named as Plectiscidea lanhami (1941)
	†Carinibus	†C. molestus Spasojevic et al., 2018	Lake Unita		A darwin wasp of uncertain subfamily placement.
	Eclytus?	†E.? lutatus Scudder, 1890	Lake Gosiute?	1 specimen	A possible Eclytini tribe tryphonine darwin wasp.
	†Eopimpla	†E. grandis Cockerell, 1921	Lake Unita		A darwin wasp of uncertain subfamily placement.
	Glypta?	†G.? transversalis Scudder, 1890	Lake Gosiute?	1 specimen	A Glyptini tribe banchine darwin wasp.
	†Ichninsum	†I. appendicrassum	Lake Unita		A pimpline darwin wasp.
	†Lithotorus	†L. cressoni Scudder, 1890	Lake Gosiute	1 specimen	An Orthocentrine darwin wasp.
	Mesoclistus?	†M.? yamataroti	Lake Unita		A possible Coleocentrini tribe acaenitine darwin wasp.
	Pimpla?	†P.? eocenica Cockerell, 1919	Lake Unita		A possible Pimplini tribe pimpline darwin wasp.
	Phygadeuon	†P. petrifactellus Cockerell, 1921	Lake Uinta		A Phygadeuontini tribe phygadeuontine darwin wasp.
	Rhyssa?	†R.? juvenis Scudder, 1890	Lake Gosiute	1 specimen	A Rhyssini tribe rhyssine darwin wasp.
	Scambus?	†S.? mandibularis	Lake Unita		A possible Ephialtini tribe pimpline darwin wasp.
	Scambus?	†S.? parachuti	Lake Unita		A possible Ephialtini tribe pimpline darwin wasp.
	†Tilgidopsis	†T. haesitans Cockerell, 1921	Lake Unita		A darwin wasp of uncertain subfamily placement.
	†Trymectus	†T. amasidis (Cockerell & LeVeque, 1931)	lake Unita		A darwin wasp of uncertain subfamily placement. First described as Tryphon amasidis (1931)
Siricidae	†Eoteredon	†E. lacoi Archibald, Aase & Nel, 2021	Fossil Lake	1 specimen	A siricid Horntail wood-wasp.
Tenthredinidae	Eutomostethus	†E. karimae Nel, Niu & Wei, 2022	Lake Unita		A blennocampine common sawfly.
Tenthredinidae	Taxonus	†T. nortoni Scudder, 1890	Lake Gosiute	1 specimen	An allantine common sawfly.
Vespidae	Alastor	†A. solidescens (Scudder, 1890)	Lake Gosiute	1 specimen	A eumenine potter wasp. First named as Didineis solidescens (1890).
Xyelidae	†Proxyelia	†P. pankowskii Jouault, Aase & Nel, 2021	Fossil Lake	1 specimen	A xyelid sawfly

Lepidoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Depressariidae	†Hexerites	†H. primalis Cockerell, 1933	Lake Unita	1 specimen	A stenomatine depressariid moth. First placed in the family Thyrididae.

Megaloptera

Family	Genus	Species	Member	Abundance	Notes	Image
Sialidae	Indeterminate	Indeterminate		1 specimen	A probable alderfly larvae

Neuroptera

Family	Genus	Species	Member	Abundance	Notes
Berothidae	†Xenoberotha	†X. angustialata Makarkin, 2017	Lake Unita (Parachute Creek Member)	1 specimen	A berothine beaded lacewing.
Hemerobiidae	Megalomus?	†M.? coloradensis Makarkin, 2024	Lake Unita		A possible Megalomus brown lacewing.
Hemerobiidae	†Pseudonotherobius	†P. kohlsi Makarkin, 2024	Lake Unita		A brown lacewing.
Mantispidae	†Protonolima	†P. mantispinoformis Makarkin, 2019	Lake Unita	1 specimen	A calomantispine mantidfly.
Myrmeleontidae	†Epignopholeon	†E. sophiae Makarkin, 2017	Lake Unita (Parachute Creek Member)	1 specimen	A probable Gnopholeontini tribe myrmeleontine antlion.
Osmylidae	†Osmylidia	†O. taliae Makarkin, Archibald, & Mathewes, 2021	Lake Unita	1 specimen	A protosmyline Osmylid lance lacewing
Incertae sedis	†Callospilopteron	†C. ocellatum Cockerell, 1921			A lacewing of uncertain affiliation. First identified as an hemipteran planthopper.

Raphidioptera

Family	Genus	Species	Member	Abundance	Notes	Image
Raphidiidae	†Archiinocellia	†A. protomaculata (Engel, 2011)	Lake Unita (Parachute Creek Member)	ca. 18 specimens	A raphidiid snakefly. Moved from Agulla protomaculata (2011)

Odonata

Primarily based on Bechly et al (2020):

Family	Genus	Species	Member	Abundance	Notes
Amphipterygidae	†Protamphipteryx	P. basalis Cockerell, 1921	Fossil Lake		An amphipterygine amphipterygid damselfly.
†Carleidae	†Carlea	C. eocenica Bechly et al., 2020	Fossil Lake	1 specimen	A carleid damselfly.
Cordulibellulidae	†Cordulibellula	C. inopinata Sroka, Howells & Nel, 2023	Lake Uinta (Parachute Creek)		A cordulibellulid dragonfly the last member of an ancient lineage.
Corduliidae	†Stenogomphus	†S. carletoni Scudder, 1892	Lake Unita		A green-eyed skimmer.
Corduliidae	†Stenogomphus	†"S." scudderi Cockerell, 1921	Lake Unita		First described as a green-eyed skimmer. Deemed nomen dubium in 1994.
Dysagrionidae	†Dysagrion	D. fredericii Scudder, 1878	Lake Gosiute (Laney Member)	16 specimens	A dysagrionine dysagrionid damselfly.
		D. integrum Bechly et al., 2020	Fossil Lake (Fossil Butte Member)	1 specimen
		D. lakesii Scudder, 1890	Lake Gosiute (Laney Member)	1 specimen
		D. packardii Scudder, 1885	Lake Gosiute (Laney Member)	1 specimen
	†Petrolestes	†P. hendersoni Cockerell, 1927	Lake Uinta (Parachute Creek Member)	1 specimen	A petrolestine dysagrionid damselfly.
	†Petrolestes	†P. inexpectatus Nel, 2024	Lake Uinta	1 specimen	A petrolestine dysagrionid damselfly.
	†Tenebragrion	†T. tynskyi Nel, Palazzo & Aase, 2023	Fossil Lake		A dysagrionid damselfly.
	†Tenebragrion	†T. shermani Bechly et al., 2020	Fossil Lake		A dysagrionid damselfly.
Calopterygidae	†Eocalopteryx	E. atavina Cockerell, 1920	Lake Gosiute	1 specimen	A calopterygid damsefly.
Euphaeidae	†Eodysphaea	E. magnifica Bechly et al., 2020	Lake Uinta (Parachute Creek)		A euphaeid damselfly.
	†Epallagites	E. avus Cockerell, 1925	Lake Unita
	†Labandeiraia	L. americaborealis Petrulevičius et al., 2007	Lake Uinta (Parachute Creek)	1 specimen
	†Labandeiraia	L. riveri Bechly et al., 2020	Fossil Lake	1 specimen
	†Litheuphaea	L. coloradensis Petrulevičius et al., 2007	Lake Uinta (Parachute Creek)	1 specimen
†Eolestidae	†Eolestes	E. syntheticus Cockerell, 1940	Lake Unita		An eolestid damselfly.
Megapodagrionidae	†Viridiflumineagrion	V. aasei Nel, 2022	Lake Uinta (Parachute Creek Member)	1 specimen	A "megapodagrionid" damselfly.
Thaumatoneuridae	†Gusagrion	†G. coloratum Nel, 2024	Lake Uinta	1 specimen	An eodysagrionine thaumatoneurid damselfly.
	†Oreodysagrion	O. tenebris Bechly et al., 2020	Fossil Lake		A thaumatoneurid damselfly.
	†Tynskysagrion	T. brookeae Bechly et al., 2020	Fossil Lake		A thaumatoneurid damselfly.
†Urolibellulidae	†Kohlslibellula	K. lini Nel, 2020	Lake Uinta (Parachute Creek)		A urolibellulid dragonfly
†Urolibellulidae	†Urolibellula	U. eocaenica Zeiri et al., 2015	Lake Uinta (Parachute Creek)		A urolibellulid dragonfly
Zacallitidae	†Zacallites	Z. balli Cockerell, 1928			A zacallitid damselfly.
Zacallitidae	†Zacallites	Z. cockerelli Bechly et al., 2020	Fossil Lake		A zacallitid damselfly.
incertae sedis	†Eopodagrion	"E." abortivum (Scudder, 1878)	Lake Gosiute (Laney Member)	1 specimen	A damselfly of uncertain affinities. First named as Podagrion abortivum (1878) Deemed nomen dubium by Nel (2022).
incertae sedis	†Eopodagrion	E. scudderi Cockerell, 1921	Lake Gosiute (Laney Member)	3 specimens	A damselfly of uncertain affinities.

Orthoptera

Family	Genus	Species	Member	Abundance	Notes
Eumastacidae	†Eoerianthus	†E. eocaenicus Gorochov, 2012	Lake Unita		A monkey grasshopper.
Eumastacidae	†Eoerianthus	†E. multispinosus (Scudder, 1890	Lake Gosiute		A monkey grasshopper. First described as Tyrbula multispinosa (1890.
Gryllidae	Eogryllus	†E. unicolor Gorochov, 2012	Lake Unita		A Gryllini tribe grylline true cricket.
	Eogryllus?	†E.? elongatus Gorochov, 2012	Lake Unita		A Gryllini tribe grylline true cricket. Tentatively placed in Eogryllus
	†Pronemobius	†P. induratus Scudder, 1890	Lake Gosiute		A Gryllini tribe grylline true cricket.
		†P. ornatipes Cockerell, 1921	Lake Unita		A Gryllini tribe grylline true cricket.
		†P. tertiarius (Scudder, 1878)	Lake Gosiute Lake Unita		A Gryllini tribe grylline true cricket. Tentatively placed in Pronemobius First described as Nemobius tertiarius (1878).
Gryllotalpidae	†Pterotriamescaptor?	†P.? americanus Gorochov, 2012	Lake Unita		A gryllotalpine mole cricket.
Mogoplistidae	†Eomogoplistes	†E. longipennis Gorochov, 2012	Lake Unita		A Mogoplistini tribe Mogoplistine scaly cricket.
	†Pteromogoplistes	†P. grandis Gorochov, 2012	Lake Unita		A Mogoplistini tribe Mogoplistine scaly cricket.
		Indeterminate	Lake Unita		Indeterminate scaly cricket fossils.
		†P.? smithii (Scudder, 1890)	Lake Unita		A Mogoplistini tribe Mogoplistine scaly cricket. Tentatively moved from Pronemobius smithii (1890)
Phalangopsidae	Eotrella	†E. mira Gorochov, 2012	Lake Unita		A phalangopsine spider cricket.
	Eozacla	†E. arachnomorpha Gorochov, 2012	Lake Unita		A phalangopsine spider cricket.
	Eozacla	†E. problematica Gorochov, 2012	Lake Unita		A phalangopsine spider cricket.
Tetrigidae	Eotetrix	†E. unicornis Gorochov, 2012	Lake Unita		A batrachideine pygmy grasshopper.
Tettigoniidae	Arethaea	†A. solterae Heads, Thomas & Hedlund, 2023	Lake Uinta (Parachute Creek)	1 specimen	A roundhead katydid.
Trigonidiidae	Indeterminate	Indeterminate	Lake Unita		Indetermiante trigonidiine sword-tail cricket remains.
Incertae sedis	†Tettoraptor	†T. maculatus Gorochov, 2012	Lake Unita		A Tettigoniid grasshopper of uncertain affiliations.

Pocodae

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Paropsocus	†P. disjunctus Scudder, 1890	Lake Unita		A louse of uncertain family affinities.

Strepsiptera

Family	Genus	Species	Member	Abundance	Notes	Image
Myrmecolacidae	Caenocholax	†C. barkleyi Antell & Kathirithamby, 2016	Lake Unita (Parachute Creek Member)	1 specimen	A male twisted-winged insect.
Myrmecolacidae	Caenocholax	†C. palusaxus Antell & Kathirithamby, 2016	Lake Unita (Parachute Creek Member)	1 specimen	A male twisted-winged insect.

Thysanoptera

Family	Genus	Species	Member	Abundance	Notes
Aeolothripidae	†Lithadothrips	†L. vetustus Scudder, 1875	Lake Uinta	2 specimens	An Aeolothripid thrips. First named as Lithadothrips vetusta
Aeolothripidae	†Palaeothrips	†P. fossilis Scudder, 1867	Lake Unita	1 specimen	An Aeolothripid thrips.
Melanthripidae	Melanothrips	†M. extincta Scudder, 1875	Lake Unita	1 specimen	A melanthripid thrips.

Trichoptera

Family	Genus	Species	Member	Abundance	Notes
Hydropsychidae	Hydropsyche	†H. operta (Scudder, 1877)	Lake Unita		A hydropsychine net-spinning caddisfly. First named as Phryganea operta (1877)
Hydroptilidae	Hydroptila	†H. phileos Cockerell, 1921	Lake Unita		A Hydroptilini tribe hydroptiline purse-case caddisfly.
Limnephilidae	†Indusia	†I. calculosa Scudder, 1878	Lake Gosiute?		A northern caddisfly larval case morphospecies.
Limnephilidae	Limnephilus	†L. eocenicus Cockerell, 1921	Lake Unita	1 specimen	A northern caddisfly species.
Incertae sedis	†Litholimnephilops	†L. yinani Robinson, Thomas & Heads, 2018	Lake Unita		A caddis fly of uncertain affiliations.

Mollusks

Based on Grande (1984):

Bivalvia

Family	Genus	Species	Member	Notes	Image
Sphaeriidae	Sphaerium	S. sp.	Lake Gosiute	A fingernail clam.	S. rhomboideum, a modern species
Unionidae	Plesielliptio	P. priscus		A unionid river mussel.
Unionidae	Plesielliptio	P. sp. "A"		A unionid river mussel.

Gastropoda

Family	Genus	Species	Member	Notes
Bulimulidae	Oreoconus	O. sp.	?	An undescribed bulimulid land snail.
Gastrocoptidae	Gastrocopta	G. sp.		An undescribed gastrocoptid land snail.
Hydrobiidae	Hydrobia	†H. aff. utahensis C. A. White, 1876	Lake Gosiute (Laney, Tipton Shale, Wilkins Peak)	An undescribed mud snail. Affiliated to Hydrobia utahensis
Hydrobiidae	Hydrobia	H. sp.	Lake Gosiute (Tipton Shale)	An undescribed mud snail.
Lymnaeidae	Lymnaea	L. similis Meek, 1860	Lake Gosiute (Laney)	A pond snail.
	Lymnaea	L. sp.	Lake Gosiute (Tipton Shale)	A undescribed pond snail.
	†Pleurolimnaea	P. tenuicosta (Meek & Hayden, 1856)	Lake Gosiute (Tipton Shale?)	A pond snail. First named Limnaea tenuicosta (1856)
Planorbidae	Biomphalaria	B. aequalis (C. A. White, 1880)	Lake Gosiute (Laney, Tipton Shale)	A ramshorn snail.
	Biomphalaria	B. storchi (L. S. Russell, 1931)	Lake Gosiute (Laney?, Tipton Shale)	A ramshorn snail.
	†Headonia	†H. pseudoammonius (Schlotheim, 1820)	Lake Gosiute (Laney, Tipton Shale)	A ramshorn snail First reported as Biomphalaria pseudoammonius (1820)
	?Drepanotrema	?D. sp.	Lake Gosiute (Laney)	An undescribed ramshorn snail.
	Gyraulus	G. militaris (C.A. White, 1880)	Lake Gosiute (Tipton Shale)	A ramshorn snail.
	Omalodiscus	†O. cirrus (C. A. White, 1879)	Lake Gosiute (Tipton Shale)	A ramshorn snail.
Physidae	Physa	†?P. longiuscula Meek & Hayden, 1856	Lake Gosiute (Tipton Shale)	An bladder snail.
	Physa	P. sp.	Lake Gosiute (Laney)	An undescribed bladder snail.
	Physella	†P. bridgerensis (Meek, 1873)	Lake Gosiute (Laney, Tipton Shale)	A bladder snail. First named Physa bridgerensis (1873)
	Physella	†P. pleromatis (C. A. White, 1877)	Lake Gosiute (Tipton Shale)	A bladder snail. First named Physa pleromatis (1877)
Pleuroceridae	Elimia	†E. tenera (J. Hall, 1845)	Lake Gosiute (Laney, Tipton Shale, Wilkins Peak)	A pleurocerid snail. the most common snail in the formation The source of the famous "Turritella agate".
Urocoptidae	Holospira	H. sp.		An undescribed urocoptid land snail.
Valvatidae	Valvata	†V. subumbilicata (Meek & Hayden, 1856)	Lake Gosiute (Laney, Tipton Shale)	A valve snail.
Valvatidae	Valvata	Cf. †V. filosa Whiteaves, 1885	Lake Gosiute (Laney, Tipton Shale)	An undescribed valve snail. Similar to Valvata filosa
Viviparidae	Viviparus	†V. meeki Wenz, 1930	Lake Gosiute (Laney, Tipton Shale)	A river snail. First named Paludina trochiformis (1856 non-von Zieten, 1830)
Viviparidae	Viviparus	†V. paludinaeformis (J. Hall, 1845)	Lake Gosiute (Tipton Shale)	A river snail.

Fungi

Genus	Species	Member	Abundance	Notes
†Caenomyces	C. eucalypti Knowlton, 1923	Lake Unita	1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a Syzygioides americana leaf. Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
	†C. planerae R.W. Brown, 1929		1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a Cedrelospermum nervosum leaf. Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
	†C. sapindicola Knowlton, 1923		1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on the Toxicodendron winchesteri holotype leaf. Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
†Eoglobella	†E. longipes Bradley, 1931	Lake Unita	1 specimen	A Staurosporaen fungal spore of uncertain affiliation.
Entophlyctis	†E. willoughbyi Bradley, 1967	Lake Gosiute (Wilkins Peak Member)		An aquatic chytridialean fungi.
Pleotrachelus	†P. askaulos Bradley, 1967	Lake Gosiute (Wilkins Peak Member)		An aquatic chytridialean fungi.
†Sphaerites	†S. myricae (Lesquereux) Meschinelli, 1892	Lake Gosiute	1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a "Rhus" nigricans leaf. First described as Sphaeria myricae (1878) Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.

Unicellular microbiota

Cyanobacteria

Family	Genus	Species	Member	Notes
Chroococcaceae	Chroococcus	?Cf. C. dispersus (Von Keissler) Lemmerman	Lake Unita	An aquatic chroococcacean cyanobacteria.
Chroococcaceae	Chroococcus	Cf. C. westii (W. West) P. Boye, 1929	Lake Unita	An aquatic chroococcacean cyanobacteria.
Coleofasciculaceae	Stigonema	†S. anchistina Bradley, 1970	Lake Gosiute (Wilkins Peak Member)	An aquatic coleofasciculacean cyanobacteria.
Coleofasciculaceae	Symploca	†S. hedraia Bradley, 1970	Lake Gosiute (Wilkins Peak Member)	An aquatic coleofasciculacean cyanobacteria.
Gomontiellaceae	Crinalium	?Cf. C. endophyticum Crow	Lake Unita	An aquatic gomontiellaceous cyanobacteria.
Gomontiellaceae	Crinalium	C. sp.	Lake Unita	An aquatic gomontiellaceous cyanobacteria. First identified as Spirulina sp. (1916)
Hapalosiphonaceae	Hapalosiphon	H. confervaceus Borzi	Lake Unita	An aquatic hapalosiphonaceous cyanobacteria.

Algae

Family	Genus	Species	Member	Abundance	Notes
Chaetophoraceae	Stigeoclonium?	Cf. S.? lubricum (Dillwyn) Kützing	Lake Unita	1 specimen	A possible chaetophoraceous chlorophyte green algae.
Chlorellaceae	†Chlorellopsis	†C. coloniata Reis, 1923			A unicellular stromatolite forming green algae.
Chlorellaceae	†Confervites	†C. mantiensis Bradley, 1929	Lake Uinta (Manti beds)		A filamentous stromatolite forming green algae.
Hydrodictyaceae	Tetraedron	Cf. T. regulare var. torsum (Turner) Brunnthaler	Lake Unita	1 specimen	A hydrodictyaceous chlorophyte green algae.
Microsporaceae	Microspora	Cf. M. pachyderma (Wille) Lagerheim	Lake Unita	1 specimen	A sphaeroplealean chlorophyte green algae.
Oocystaceae	Cf. Eremosphaera	E. sp.	Lake Unita		An oocystacean chlorophyte green algae.
Scenedesmaceae	Coelastrum	Cf. C. verrucosum Retnsch	Lake Unita		A sphaeroplealean chlorophyte green algae.
Schizochlamydaceae	Schizochlamys	†S. haywellensis Bradley, 1970	Lake Gosiute (Wilkins Peak Member)		A sphaeroplealean chlorophyte green algae.

Charophytes

Family	Genus	Species	Member	Abundance	Notes	Image
Zygnemataceae	Spirogyra	†S. wyomingia Bradley, 1970	Lake Gosiute (Wilkins Peak Member)		A zygnematacean charophyte green algae.

Euglenozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Phacaceae	Phacus	Cf. P. caudata Hübner, 1886		1 specimen	A Euglenozoan algal flagellate.

Amoebozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Difflugiidae	Difflugia	†D. calcifera Bradley, 931	Lake Unita	1 specimen	A difflugiid lobose testate amoeba.
Quadrulidae	Quadrula	†Q. minuta Bradley, 1931	Lake Unita	1 specimen	A quadrulid lobose testate amoeba.

Cercozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Euglyphidae	Euglypha	†E. robusticornis Bradley, 1931	Lake Unita		A Cercozoan eukaryote.

Plants

Liverworts

Family	Genus	Species	Member	Abundance	Notes	Image
Jungermanniaceae	†Jungermannites	†J. eophilus (Cockerell) Steere, 1947	Lake Unita	1 specimen	A leafy liverwort morphospecies. First described as Lejeunea eophila (1925).

Lycophytes

Family	Genus	Species	Member	Abundance	Notes	Image
Isoetaceae	†Isoetites	†I. horridus (Dawson) R.W. Brown, 1939	Lake Unita	1 specimen	An isoetaceous quillwort morphospecies. First described as Carpolithes horridus (1884).

Ferns

Family	Genus	Species	Member	Abundance	Notes
Aspleniaceae	Asplenium	†A. delicatula (R.W. Brown) MacGinitie, 1969			A spleenwort fern leaf morphospecies. First described as Aneimia delicatula (1929) Includes Phyllites furcivena (1934) Jr homonym of Asplenium delicatulum C. Presl, 1825
Aspleniaceae	Asplenium	†A. eoligniticum Berry, 1916	Lake Unita	2 specimens	A spleenwort fern leaf morphospecies.
Equisetaceae	Equisetum	†E. haydenii Lesquereux, 1872	Not Green River Formation		A scouring rush morphospecies. First ascribed to the formation Excluded by Knowlton (1923) as a Washakie Formation species.
		†E. winchesteri Brown, 1929	Lake Gosiute	1 specimen	A scouring rush morphospecies.
		†E. wyomingense Lesquereux, 1873	Lake Gosiute Lake Unita		A scouring rush morphospecies.
Lygodiaceae	Lygodium	†L. dentoni Lesquereux, 1875	Rejected	1? specimen	A climbing fern species. Type specimens missing by 1923 Brown (1934) rejected per Knowlton and dubious locality.
Lygodiaceae	Lygodium	†L. kaulfussi Heer, 1861	Fossil Lake Lake Gosiute Lake Unita	Numerous	A climbing fern species. First described as Lygodium neuropteroides (1870)
Marattiaceae	Danaea	†D. coloradensis Knowlton, 1923	Lake Unita	1 specimen	A possible marattiaceous fern leaf morphospecies. The genus affiliation was questioned by MacGinitie (1969).
Marsileaceae	Marsilea	†M. sprungerorum Hermsen, 2019	Lake Uinta (Parachute Creek Member)	Numerous	A water clover fern.
Osmundaceae	Osmunda?	†Q.? sepulta (Newberry) Knowlton, 1923	Lake Gosiute	2 specimens	A possible osmundaceous fern leaf morphospecies. First described as Pecopteris sepulta (1883)
Salviniaceae	Azolla	†A. berryi Brown, 1934	Lake Unita	1 specimen	A mosquito fern morphospecies.
Pteridaceae	Acrostichum	†A. hesperium Newberry, 1883	Lake Gosiute	3 specimens	A parkerioid leather fern leaf morphospecies.

Cycadalean palynomorphs

Genus	Species	Member	Abundance	Notes	Image
†Cycadopites	†C. sp.	Lake Uinta (Parachute Creek Member)	Numerous	Various Cycas - Zamia Cycad palynomorphs.
†Dioonipites	†D. sp.	Lake Uinta (Parachute Creek Member)	5 specimens	A Dioon like Cycad palynomorph.

Conifers

Family	Genus	Species	Member	Abundance	Notes
Cupressaceae	Sequoia	Cf. †S. affinis Lesquereux, 1876	Lake Unita		A coast redwood relative.
Pinaceae	Picea	†P. pinifructus Brown, 1929	Lake Unita	1 specimen	First identified as a spruce seed. Deemed a jr synonym of Cercidiphyllum arcticum by Brown (1939).
	Pinus	†P. balli Brown, 1934	Lake Unita	1 specimen	A 3-needle pine morphospecies.
	Pinus	†P. florissanti Lesquereux, 1885	Lake Unita	1 specimen	A pine morphospecies.
incertae sedis	†Taxites	†T. eocenica Brown, 1929	Lake Uinta	1 specimen	Isolated conifer needles of uncertain affiliation. Similar to some Cupressaceae, Pinaceae, and Taxaceae genera.

Conifer palynomorphs

Genus	Species	Member	Abundance	Notes
Abies	†A. concolipites Wodehouse, 1933	Lake Uinta (Parachute Creek Member)	2 specimens	A true fir palynomorph similar to Abies concolor.
†Abietipites	†A. antiquus Wodehouse, 1933		2 specimens	A generally abietine-like palyonomorph.
†Cedripites	†C. eocenicus Wodehouse, 1933		1 specimen	A Cedrus libani cedar-like palyonomorph.
†Inaperturopollenites	†I. concedipites (Wodehouse) Krutzsch, 1971		6 specimens	A Cunninghamia-like palyonomorph. First described as Cunninghamia concedipites (1933).
†Inaperturopollenites	†I. vacuipites (Wodehouse) Krutzsch, 1971		1 specimen	A Chinese swamp cypress-like palyonomorph. First described as Glyptostrobus vacuipites (1933).
Picea	†P. grandivescipites Wodehouse, 1933		6 specimens	A spruce palyonomorph.
Pinus	†P. scopulipites Wodehouse, 1933		Numerous	A pine palyonomorph. Less common than P. strobipites.
	†P. strobipites Wodehouse, 1933		Numerous	A pine palyonomorph.
	†P. tuberculipites Wodehouse, 1933		6 specimens	A pine palyonomorph.
Taxodium	†T. hiatipites Wodehouse, 1933		Numerous	A bald cypress palyonomorph.
†Tsugaepollenites	†T. viridifluminipites (Wodehouse) R. Potonié, 1958		1 specimen	A hemlock-like palyonomorph. First described as Tsuga viridifluminipites (1933) Moved from Zonalapollenites viridifluminipites (1953).

Gnetalean palynomorphs

Genus	Species	Member	Abundance	Notes	Image
†Gnetaceae-pollenites	†G. eocenipites (Wodehouse) R. Potonié, 1958	Lake Uinta (Parachute Creek Member)	12 specimens	An Ephedra-like palyonomorph. First described as Ephedra eocenipites (1933).

Magnoliids

Family	Genus	Species	Member	Abundance	Notes
Aristolochiaceae	Aristolochia	†A. mortua Cockerell, 1908			An pipevine leaf morphospecies.
Hernandiaceae	Illigera	†I. eocenica Manchester & O'Leary, 2010	Lake Unita		An Illigera seed morphospecies.
Lauraceae	Beilschmiedia	†B. eocenica (R.W. Brown) MacGinitie, 1969 (non-Reid & Chandler, 1933)			A Beilschmiedia leaf morphospecies. Name a jr homonym of Beilschmiedia eocenica E. Reid & M. Chandler, 1933
	Lindera	†L. allardi MacGinitie, 1969			A spicebush leaf morphospecies.
	Lindera	†L. varifolia MacGinitie, 1969	Lake Unita		A spicebush leaf morphospecies.
	Ocotea	†O. coloradensis (R.W. Brown) LaMotte, 1952)	Lake Unita	2 specimens	An Ocotea leaf morphospecies. First described as Oreodaphne coloradensis (1934)
	Ocotea	†O. lancifolia (Lesquereux) LaMotte, 1952	Lake Gosiute Lake Unita	2 specimens	An Ocotea leaf morphospecies. First described as Sapindus lancifolius (1883) Moved from Oreodaphne lancifolia (1934) Also includes Oreodaphne knowltoni (1929)
	Persea	†P. coriacea MacGinitie, 1969 (non-Engelhardt, 1895)			A Persea leaf morphospecies. Name a jr homonym of Persea coriacea Engelhardt, 1895
Magnoliaceae	Talauma	†T. multiperforata Kruse, 1954	Lake Gosiute (Eden Valley)		An magnolia wood morphospecies.

Magnoliid palynomorphs

Genus	Species	Member	Abundance	Notes	Image
Liriodendron	†L. psilopites Wodehouse, 1933	Lake Uinta (Parachute Creek Member)	1 specimen	A Liriodendron tulipifera-like tulip tree palyonomorph.

Monocots

Family	Genus	Species	Member	Abundance	Notes
Arecaceae	†Palmacites	†P. florissanti (Lesquereux) Read & Hickey, 1972	Fossil Lake Lake Uinta		A palmately veined palm leaf morphospecies. Moved from Sabalites florissanti (1930) First described as Flabellaria florissanti (1885).
	†Palmoxylon	†P. colei Tidwell, 1973	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
		†P. contortum Tidwell, 1973	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
		†P. edenense Tidwell, 1973	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
		†P. macginitiei Tidwell et al., 1971	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
	†Phoenicites	†P. haydenii (Newberry) Read & Hickey, 1972		1 specimen	A pinnately veined palm leaf morphospecies. Moved from Geonomites haydenii (1923) First described as Manicaria haydenii (1883).
	Phoenix	†P. windmillis	Fossil Lake (Fossil Butte member) Lake Gosiute (Laney Member) Lake Unita (Parachute Creek Member)	numerous	A Phoenix flower morphospecies.
	†Sabalites	†S. powellii (Newberry) Berry, 1930	Fossil Lake Lake Uinta		A palmate veined palm leaf morphospecies. First described as Sabal powellii (1883).
Cypreaceae?	†Cyperacites	†C.? filiferus	Lake Unita	A number of specimens	Likely rhizomes of monocots?. Oldest specimen first described as Diatryma? filifera, a Gastornis feather. Description of the species as a bird feather was questioned by Alexander Wetmore (1930) Brown (1934) moves the species to Carpolithus filiferus A second group of fossils is described as a cactus, Eopuntia douglassi (1944) Brown redescribes both as cypreaceous rhizomes, and synonymized both species as Cyperus filiferus (1959) Herman Becker (1963) agrees with the synonymy, but moves the species to the Cyperacites morphogenus as Cyperacites? filiferus.
	†Cyperacites	†C. haydenii (Lesquereux) Knowlton, 1898	Lake Gosiute	Several specimens	A sedge-like leaf morphospecies. First described as Cyperites haydenii (1885)
	Cyperus	†C. chavannesi Heer, 1855	Rejected	1 specimen	A sedge leaf morphospecies. Rejected by Knowlton (1923) due to dubious location.
Dioscoreaceae	Dioscorea	†D. lindgrenii Herrera & Manchester	Fossil Lake (Fossil Butte Member)	2 specimens	sweet potato fruits
Dioscoreaceae	Dioscorea	†D. shermanii Herrera & Manchester	Fossil Lake (Fossil Butte Member)	1 specimen	sweet potato fruits
Juncaceae	Juncus	Indeterminate	Rejected	?	Scrappy material tentatively identified as rush leaves. Discredited by Knowlton (1923) due to missing specimens
Poaceae	Arundo	†A. reperta Lesquereux, 1876	Lake Uinta	Several specimens	A giant cane grass morphospecies. The species validity of the fossils was questioned by Knowlton (1923) The species was retained by Brown (1934).
Potamogetonaceae	Potamogeton	†P. rubus R.W. Brown, 1934	Lake Unita		A pondweed morphotype.
Typhaceae	Sparganium	†S. antiquum (Newberry) Berry, 1924	Lake Gosiute Lake Uinta		A bur-reed leaf morphospecies. Moved from Brasenia? antiqua (1883) includes Pontederites hesperia (1923)
	Sparganium	†S. eocenicum R.W. Brown, 1929	Lake Gosiute	1 specimen	A bur-reed leaf morphospecies.
	Typha	†T. lesquereuxi Cockerell, 1906	Lake Unita		A cattail morphospecies.
Zingiberaceae	†Musophyllum	†M. complicatum Lesquereux, 1873	Lake Gosiute	5 specimens	A zingiberaceous monocot leaf morphotaxon.

Monocot palynomorphs

Genus	Species	Member	Abundance	Notes
†Arecipites	†A. punctatus Wodehouse, 1933	Lake Uinta (Parachute Creek Member)	1 specimen	A Phoenix dactylifera-like palm palyonomorph.
†Arecipites	†A. rugosus Wodehouse, 1933		1 specimen	A palm palyonomorph.
†Peltandripites	†P. davisii Wodehouse, 1933		1 specimen	A tuckahoe-like arrow-arum palyonomorph.
Potamogeton	†P. hollickipites Wodehouse, 1933		Numerous	A Potamogeton palyonomorph.
†Smilacipites	†S. echinatus Wodehouse, 1933		2 specimen	A Smilax-like palyonomorph.
	†S. herbaceoides Wodehouse, 1933		1 specimen	A Smilax-like palyonomorph.
	†S. molloides Wodehouse, 1933		1 specimen	A Smilax mollis-like palyonomorph.

Ceratophyllales

Family	Genus	Species	Member	Abundance	Notes	Image
Ceratophyllaceae	Ceratophyllum	†C. muricatum incertum (Brown) Herendeen, Les, & Dilcher, 1990	Fossil Lake	19 Specimens	A prickly hornwort subspecies. First described as a full species by Berry (1930). Moved to subspecies by Herendeen, Les, & Dilcher (1990)

Eudicots

Basal Eudicots

Family	Genus	Species	Member	Abundance	Notes
Berberidaceae	"Mahonia"	†"M." eocenica (R.W. Brown) C.A. Arnold, 1936	Lake Uinta	1 Specimen	An oregon grape leaf morphospecies. First described as Odostemon eocenica (1934)
Menispermaceae	†Menispermites	†M. limacioides MacGinitie, 1969			A moon-seed leaf morphospecies.
Menispermaceae	Stephania	†S. wilfii	Lake Uinta (Wilkins Peak Member)	Numerous	A moon-seed fruit.
Nelumbonaceae	†Nelumbo	Undescribed	Lake Gosiute		An undescribed Lotus leaf species.
Platanaceae	†Macginitiea	†M. wyomingensis (Knowlton & Cockerell) Manchester, 1986			A sycamore relative species. First named Aralia macrophylla Newberry, 1882 (non-Lindley, 1844) Renamed to Aralia wyomingensis (1919) Moved from Platanus wyomingensis (1969)
Proteaceae	Lomatia	†L.? microphylla Lesquereux, 1876	Rejected	specimens missing	First identified as a guitarplant leaf morphospecies. Rejected by Brown (1934) due to missing specimens & dubious locality.
Trochodendraceae?	†Nordenskioldia	†N. borealis Heer, 1870	Rejected		A basal trochodendroid Rejected by Manchester, Crane & Dilcher (1991) considered Carpites cocculoides like.

Superasterids

Family	Genus	Species	Member	Abundance	Notes
Apocynaceae	†Apocynospermum	†A. coloradensis Brown, 1929	Lake Unita	1 specimen	An apocynaceous seed morphospecies.
Apocynaceae	†Aspidospermoxylon	†A. uniseriatum Kruse, 1954	Lake Gosiute		An apocynaceous wood morphospecies.
Araliaceae	†Allopanax	†A. quinus (Macginitie) Doweld	Lake Uinta (Parachute Creek Member)		An Apialean leaf mophogenus. Moved from Araliophyllum quinus (1969).
Asteraceae	†Achaenites	†A. cichorioides Knowlton, 1923	Lake Uinta	1 specimen	An asteraceous seed morphospecies.
Caryophyllaceae	†Alsinites	†A. revelatus Cockerell, 1925	Lake Unita	1 specimen	A caryophyllacous flower morphotaxon.
Ericaceae	Andromeda	†A. delicatula Lesquereux, 1885		2 specimens	A possible bog rosemary ericaceous morphospecies.
Oleaceae	Osmanthus	†O. praemissa (Lesquereux) Cockerell, 1906	Lake Uinta		An Osmanthus leaf morphospecies. Moved from Olea praemissa (1885)
Oleaceae	Osmanthus	†O. viridiflumensis (Knowlton) MacGinitie, 1969)	Lake Uinta	1 specimen	An Osmanthus leaf morphospecies. Moved from Ocotea viridiflumensis (1952) First described as Oreodaphne viridiflumensis (1923)
Polemoniaceae	†Gilisenium	†G. hueberi Lot, Manchester, & Dilcher, 1998	Fossil Lake	1 specimen	A polemoniaceous herb species.
Solanaceae	†Lycianthoides	†L. calycina Deanna et al.	Lake Uinta (Parachute Creek Member)	3 specimens	A nightshade family fruit
Styracaceae	Styrax	†S. transversa MacGinitie, 1969	Lake Unita		A snowbell leaf morphotype.
Symplocaceae	Symplocos	†S. exilis MacGinitie, 1969	Lake Unita		A Symplocaceous leaf morphotype.
Incertae sedis	†Othniophyton	†O. elongatum (MacGinitie) Manchester, Judd, Correa-Narvaez, 2024	Lake Unita		A superastrid plant of possible caryophyllalean affinity. First described as Oreopanax elongatum (1969) O. elongatum synonymy Apocynophyllurn wilcoxense (of Berry, 1929) ?Celastrophyllum lesquereuxii (1929) Ternstroemites viridifluminis (1929)

Superrosids

Family	Genus	Species	Member	Abundance	Notes
Altingiaceae	Liquidambar	†L. callarche Cockerell, 1925	Lake Unita	2 specimens	A sweetgum species.
Anacardiaceae	†Barkleya	†B. schinoloxa (R. W. Brown) Manchester & Judd	Lake Gosiute (Laney Member) Lake Uinta (Parachute Creek Member)	Numerous	An anacardiaceous samara. Moved from Anacardites schinoloxus (1929). Includes Anacardites schinus R. W. Brown, 1934 Possibly the fruits of "Rhus" nigricans.
	†Edenoxylon	†E. parviareolatum Kruse, 1954	Lake Gosiute (Eden Valley)		An anacardiaceous wood morphospecies.
	Rhus	†R. longepetiolata (Lesquereux) R.W. Brown, 1934	Lake Gosiute		An anacardiaceous leaf morphotype. Moved from Ailanthus longepetiolatus (1934)
	Rhus	†"R." nigricans (Lesquereux) Knowlton, 1923	Lake Gosiute (Laney Member) Lake Uinta (Parachute Creek Member)	Numerous	An anacardiaceous leaf morphotype. Moved from Myrica nigricans (1929). Possibly the leaves of Barkleya schinoloxa. "R." nigricans synonymy Comptonia insignis Lesquereux of Brown, 1934 Cupanites nigricans (Lesquereux) Berry, 1924 Koelreuteria nigricans (Lesquereux) Brown, 1946 Koelreuteria viridifluminis (Hollick) Brown, 1934 Myrica alkalina Lesquereux, 1883 Myrica ludwigii Schimper of Lesquereux, 1872 Myrica undulata? Schimper of Lesquereux,1878 (pro part) Myrica zachariensis Saporta of Lesquereux, 1883 (pro part) Planera variabilis Newberry, 1883 (pro part) Rhus myricoides Knowlton, 1923 Rhus variabilis (Newberry) Knowlton, 1923 Staphylea viridiftuminis Hollick, 1929
	Schmaltzia	†S. vexans Cockerell, 1906	Lake Gosiute		A sumac leaf morphospecies. Moved from Rhus vexans (1885) Considered nomen dubium by MacGinitie (1969)
	†Schinoxylon	†S. actinoporosum Kruse, 1954	Lake Gosiute (Eden Valley)		An anacardiaceous wood morphospecies.
	Toxicodendron	†T. winchesteri (Knowlton) MacGinitie, 1969	Lake Gosiute		A poison oak leaf morphospecies. First described as Sapindus winchesteri (1923)
Burseraceae	Bursera	†B. inaequalateralis (Lesquereux) MacGinitie, 1969			A bursera leaf morphotype. First named as Alnites unaequalateralis (1876) B. inaequalateralis synonymy Alnites inaequalateralis Lesquereux, 1878 Alnus inaequalateralis (Lesquereux) Lesquereux, 1883 Comptonia? anomala Knowlton, 1923 Planera inaequalateralis (Lesquereux) Knowlton, 1923 Planera variabilis Newberry, 1883
Cannabaceae	Celtis	†C. mccoshii Lesquereux, 1883	Lake Gosiute Lake Unita	2 specimens	A hackberry leaf morphospecies. Includes Celtis debequensis (1928)
Capparaceae	†Forchhammerioxylon	†F. scleroticum Kruse, 1954	Lake Gosiute (Eden Valley)		A capparaceous wood morphospecies.
Celastraceae	Celastrus	†C. winchesteri (Knowlton) MacGinitie, 1969	Lake Unita	1 specimen	A staff vine leaf morphospecies. Moved from Juglans winchesteri (1923) Includes Juglans alkalina (1875 )
Celastraceae	†Celastrophyllum	†C. emarginatum	Lake Unita		A staff vine relative leaf morphospecies. MacGinitie (1969) considered the species nomen dubium.
Crassulaceae	Sedum?	†S.? hesperium Knowlton, 1923	Lake Uinta	1 specimen	A possible stonecrop leaf morphospecies. Considered nomen dubium by MacGinitie (1969)
Euphorbiaceae	†Heveoxylon	†H. microporosum Kruse, 1954	Lake Gosiute (Eden Valley)		An euphorb wood morphospecies.
Fabaceae	Amorpha	†A. utensis Cockerell, 1925	Lake Unita	1 specimen	A legume leaf morphospecies.
	Arcoa	†A. lindgreni Herendeen & Herrera, 2019	Fossil Lake	3 specimens	A caesalpinioid legume.
	Caesalpinia	†C. flumen-viridensis Herendeen & Dilcher, 1991	Lake Gosiute	2 specimens	A Caesalpinia subg. Mezoneuron seed morphospecies.
	Caesalpinia	†"C." pecorae R.W. Brown, 1956	Lake Unita	2 specimens	First named as a Caesalpinia leaf morphospecies. Considered a Detarieae tribe species by Herendeen & Dilcher (1991).
	†Caesalpinites	†C. falcatus MacGinitie, 1969			A Caesalpinia-like leaf morphospecies.
	Cercis	†C. tenuinervis (Lesquereux) R.W. Brown, 1934	Lake Gosiute	1 specimen	A redbud leaf morphospecies. Considered nomen dubium by MacGinitie (1969).
	Erythrina	†E. roanensis MacGinitie, 1969	Lake Unita		A coral tree leaf morphospecies.
	Gymnocladus	†G. hesperia (R.W. Brown) MacGinitie, 1969	Lake Uinta		A coffeetree leaf morphospecies. Moved from Cassia hesperia (1929) Includes Chrysobalanus lacustris (1929) & Banisteria bradleyi (1929)
	†Leguminosites	†L. alternans Lesquereux, 1875	Rejected		A legume leaf morphospecies. Rejected due to missing specimens by Knowlton (1923) and Brown (1934)
		†L. lesquereuxiana Knowlton, 1898	Lake Unita		A legume leaf morphospecies. L. lesquereuxiana synonymy Bumelia coloradensis Cockerell, 1925 Dalbergia knowltoni Cockerell, 1925 Dalbergia retusa Knowlton, 1923 Leguminosites cassioides Lesquereux, 1878 Sophora coloradensis Knowlton, 1923
		†L. regularis MacGinitie, 1969	Lake Unita		A legume leaf morphospecies.
	↑Parvileguminophyllum	†P. coloradensis (Knowlton) Call & Dilcher, 1994	Lake Uinta		A legume leaf morphospecies. Moved from Mimosites coloradensis (1923). P. coloradensis synonymy Podogonium americanum Lesquereux, 1878 (of Brown 1929) Mimosites debequensis R.W. Brown, 1929 Mimosites falcatus R.W. Brown, 1929
	Swartzia	†S. wardelli MacGinitie, 1969	Lake Uinta		A swartzieous leaf morphotype.
Fagaceae	Quercus	†Q. castaneopsis Lesquereux, 1883	Lake Gosiute Lake Uinta	Several fragmentary specimens	An oak morphospecies. P. cinnamomoides synonymy Quercus castanoides Newberry, 1883 Ilex maculata Lesquereux, 1883 Ilex? affinis Lesquereux, 1872 Considered nomen dubium by MacGinitie (1969).
		†"Q. cuneatus" MacGinitie, 1969 (non-Wangenh. 1787	Lake Unita		An oak leaf morphospecies. Name a jr homonym of Quercus cuneata Wangenheim, 1787
		†Q. drymeja Unger, 1847	Rejected		An oak morphospecies. First described from Europe, Rejected from flora by MacGinitie (1969).
		†Q. petros MacGinitie, 1969	Lake Unita		An oak leaf morphospecies.
		†Q. utensis Cockerell, 1927	Lake Unita		An oak morphospecies. Considered nomen dubium by MacGinitie (1969).
Hamamelidaceae	Distylium	†D. eocenica (R.W. Brown) MacGinitie, 1969	Lake Unita	1 specimen	A winter-hazel leaf morphotype. Moved from Betula coloradea (1934) First named Betula eocenica (1928 non Ettingshausen (1858) Also includes Grewiopsis cissifolius (1929)
Juglandaceae	Juglans	†J. crossii Knowlton, 1898	Lake Gosiute	1 specimen?	A walnut leaf morphospecies.
		†J. bilinica Unger, 1850		1 specimen	A walnut leaf morphospecies. Moved from Hicoria juglandiformis (1898)
		†J. schimperi Lesquereux, 1873	Lake Unita		A walnut leaf morphospecies. Includes Juglans occidentalis (1883) & Ficus ungeri (1872)
	†Palaeocarya	Cf. †P. clarnensis Manchester, 1987	Lake Gosiute?		A Palaeocarya nut morphospecies similar to Palaeocarya clarnensis.
	†Palaeocarya	†P. uintaensis (MacGinitie) Manchester, 1987	Lake Unita		A Palaeocarya nut morphospecies.
	Platycarya	†P. castaneopsis (Lesquereux) Wing & Hickey, 1984	Fossil Lake Lake Uinta		A Platycarya species. Moved from Quercus castaneopsis (1883)
	Pterocarya	†P. macginitii Manchester & Dilcher, 1982	Lake Gosiute		A wingnut seed morphospecies.
	Pterocarya	†P. roanensis MacGinitie, 1969	Lake Unita		A wingnut leaf morphospecies.
Malvaceae	Hibiscus	†H. roanensis R.W. Brown, 1934	Lake Unita		An Hibiscus leaf morphospecies.
	Ochroma	†O. murata R.W. Brown, 1934	Lake Unita	1 specimen	A balsa tree leaf morphospecies.
	Sterculia	†S. coloradensis R.W. Brown, 1929	Lake Unita	1 specimen	A sterculioid leaf morphospecies.
	Triumfetta	T. ovata MacGinitie, 1969 (non-De Candolle, 1824)			A burbark leaf morphospecies. Jr homonym of Triumfetta ovata De Candolle, 1824
Meliaceae	Cedrela	†C. trainii C.A. Arnold, 1936	Lake Unita		A meliaceous leaf morphospecies.
Moraceae	†Welkoetoxylon	†W. multiseriatum Boonchai, Manchester & Wheeler	Lake Gosiute (Tipton Shale Member)	1 specimen	A moraceous wood morphogenus.
Myricaceae	Morella scalariformis	†M. scalariformis (Kruse), Wheeler & Manchester, 2021	Lake Gosiute (Eden Valley)		A myricaceous wood morphospecies. First named as Myrica scalariformis (1954) Moved from Myricoxylon scalariforme (1962)
Myricaceae	Myrica	†M. salicina Unger, 1850	Rejected	sp missing	Identified as a European Myrica species Rejected by Knowlton (1923) due to the specimens being missing.
Myrtaceae	†Syzygioides	†S. americana (Lesquereux) Manchester, Dilcher, & Wing, 1998	Lake Unita	Circa 26 specimens	A myrtaceous species. First described as Eucalyptus americana (1872) Includes Ficus omballi (1929) & Eugenia americana (1969)
Rhamnaceae	†Berchemiopsis	†B. paucidentata MacGinitie, 1969	Lake Unita		A rhamnaceous leaf morphotype.
	Ptelea	†P. cassioides (Lesquereux) MacGinitie, 1969	Rejected		Type specimen leaf from the Florissant Formation Green River Fruits rejected by Call & Dilcher as not Ptelea.
	Ptelea	†P. paliuruoides (R.W. Brown) Manchester & O'Leary, 2010	Lake Unita		A rhamnaceous seed morphospecies. First describe as Carpolithus paliuroides (1934)
	†Rainbowia	†R. rhamnoides Manchester & O'Leary, 2010	Lake Unita		A rhamnaceous seed morphospecies.
Rosaceae	Prunus	†P. stewarti MacGinitie, 1969	Lake Gosiute		A cherry leaf morphospecies. First named Amygdalus gracilis (1883) Moved from jr homonym Prunus gracilis (non Engelm. & Gray, 1845)
Rosaceae	Rosa	†R. hillae Lesquereux, 1885			A rose leaf morphospecies.
Rutaceae	†Amyridoxylon	†A. ordinatum Kruse, 1954	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
	Fagara	†F. biseriata Kruse, 1954	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
	Fagara	†F. monophylloides Kruse, 1954	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
Salicaceae	Populus	†P. cinnamomoides (Lesquereux) MacGinitie, 1969	Lake Uinta	Numerous	An Aspen/cottonwood relative. First described as Ceanothus cinnamomoides (1872) P. cinnamomoides synonymy Carpites newberryana Knowlton, 1926 Cinnamomum scheuchzeri Heer of Lesquereux, 1883 Ficus wyomingiana Lesquereux, 1878 Ilex wyomingensis Lesquereux, 1878 Ilex vyomingensis Lesquereux, 1878 Luehea newberryana (Knowlton) MacGinitie, 1974 Zizyphus cinnamomoides (Lesquereux) Lesquereux, 1878 Zizyphus longifolia Newberry, 1883
		†P. wilmattae Cockerell, 1925	Lake Uinta	Numerous	An Aspen/cottonwood relative.
		†P. tidwellii Manchester, Judd, & Handley, 2006	Lake Uinta	Numerous	An Aspen/cottonwood relative.
	†Pseudosalix	†P. handleyi Boucher, Manchester, & Judd, 2003	Lake Uinta (Parachute Creek Member)		A salicaceous species. Includes Potentilla? byrami (1925), Antholithes polemonioides (1929) & Antholithes sp.6 (1934)
	Salix	†S. cockerelli R.W. Brown, 1934	Lake Unita	4 Specimens	A willow leaf morphospecies.
	Salix	†S. longiacuminata Knowlton ex R.W. Brown, 1934	Lake Gosiute	1 specimen	A willow leaf morphospecies. Includes Banksia cockerelli (1929)
Sapindaceae	Allophylus	†A. flexifolia (Lesquereux) MacGinitie, 1969	Lake Gosiute Lake Uinta	4 Specimens	An Allophylus leaf morphospecies. First named Euonymus flexifolia (1885) A. flexifolia synonymy Fraxinus flexifolia (Lesquereux) Brown, 1934 Fraxinus petiolata Brown, 1929 Odostemon reynoldsii Hollick, 1929
	Athyana	†A. balli (R.W. Brown) MacGinitie, 1969	Lake Unita		A sapindaceous leaf morphotype. Moved from Rhus balli (1934).
	"Cardiospermum"	†"C." coloradensis (Knowlton) MacGinitie, 1969	Lake Uninta	Numerous	A possible ballon vine leaf morphospecies. Moved from Lomatia coloradensis (1929) "C." coloradensis synonymy Lomatia obtusiuscula Cockerell, 1925 Melia coloradensis (Knowlton) Cockerell, 1925 Phyllites coloradensis Knowlton, 1923 Pimelea spatulata Knowlton, 1923
	Koelreuteria	†K. allenii (Lesquereux) W.N. Edwards, 1929	Fossil Lake (Fossil Butte Member) Lake Unita (Parachute Creek Member)		A golden rain tree fruit morphospecies. Includes Koelreuteria uiridifluminis seeds
	Sapindus	†S. dentoni Lesquereux, 1876	Lake Unita		A soapberry leaf morphotype. Includes Salix media (1872) & Myrica torryi (1929)
	Sapindus	†S. obtusifolius Lesquereux, 1874	Lake Unita		A soapberry leaf morphotype. Includes Salix media (1872) & Myrica torryi (1929)
	Thouinia	†T. eocenica R.W. Brown, 1929	Lake Unita		A soap berry leaf morphospecies.
Simaroubaceae	Ailanthus	†A. lesquereuxii Cockerell, 1927			A tree of heaven fruit morphospecies
Surianaceae	Suriana	†S. inordinata Kruse, 1954	Lake Gosiute (Eden Valley)		A surianaceous wood morphospecies.
Tetramelaceae	†Parvaspicula	†P. lepidioides (Cockerell) Deanna et al.	Lake Uinta (Parachute Creek Member)	Numerous	A tetramelaceous fruit morphotype. The type species is Clethra (?) lepidioides Cockerell (1925) Also includes Antholithes pendula (1929)
Tetramelaceae	†Punctaphyllum	†P. glandulosa (Brown) Manchester, Judd, & Kodrul, 2023	Lake Uinta (Parachute Creek Member)	Numerous	A tetramelaceous leaf morphotype. The type species is Cucurbita glandulosa (1929) Includes Aleurites glandulosa (1969) & Dendropanax latens (1974) Also includes the Ficus mississippiensis of Brown (1928)
Ulmaceae	†Cedrelospermum	†C. nervosum (Newberry) Manchester, 1989	Lake Uinta	Numerous	An ulmaceous species. First named as Planera nervosa (1883) C. nervosum synonymy Banksites lineatulus Cockerell, 1925 Dalbergia viridiflumensis Knowlton, 1923 Lomatia lineatulus (Cockerell) MacGinitie, 1969 Myrica praedrymeja Knowlton, 1923 Myrica minuta Knowlton, 1923 Rhus myricoides (Knowlton) Brown, 1929 Salix lineara Knowlton, 1923 Zelkova nervosa (Newberry) Brown, 1929
Vitaceae	Cissus	†C. parrottiaefolia Lesquereux, 1876	Rejected	3 specimens	A grape relative leaf morphospecies. Rejected from formation by Knowlton (1923) due to distinctly different matrix.

Eudicot palynomorphs

Genus	Species	Member	Abundance	Notes
†Ailanthipites	†A. berryi Wodehouse, 1933	Lake Uinta (Parachute Creek Member)		An Ailanthus glandulosa-like soapberry family palynomorph.
†Alni-pollenites	†A. speciipites (Wodehouse) Raatz, 1938		Numerous	An alder-like palynomorph. first described as Alnus speciipites (1933)
Betula	†B. claripites Wodehouse, 1933		3 specimens	A Betula lenta-like birch palynomorph.
†Caprifoliipites	†C. viridi-fluminis Wodehouse, 1933		1 specimen	A caprifoliaceous-like palynomorph.
Carpinus	†C. ancipites Wodehouse, 1933			A Carpinus-like palynomorph.
Carya	†C. juxtaporipites (Wodehouse) LaMotte, 1952		1 specimen	A hickory-like palynomorph. First described as Hicoria juxtaporipites (1933)
Carya	†C. viridi-fluminipitis (Wodehouse) Wilson & Webster, 1944		1 specimen	A Carya myristiciformis-like hickory palynomorph. First described as Hicoria viridi-fluminipites (1933)
Ericipites	†E. brevisulcatus Wodehouse, 1933		1 specimen	A Calluna vulgaris-like heath family palynomorph.
Ericipites	†E. longisulcatus Wodehouse, 1933		2 specimens	A heath family-like palynomorph.
†Juglanspollenites	†J. nigripites (Wodehouse) Wingate & Nichols, 2001		1 specimen	A walnut-like palynomorph. First described as Juglans nigripites (1933)
†Momipites	†M. corylipites (Wodehouse) D.J. Nichols, 1973		2 specimens	An Engelhardia-like palynomorph. First described as Engelhardia corylipites (1933).
†Momipites	†M. coryloides Wodehouse, 1933		Numerous	A Celtis iguanaea-like hackberry palynomorph.
†Myricipites	†M. dubius Wodehouse, 1933		1 specimen	A bayberry-like palynomorph.
Myriophyllum	†M. ambiguipites Wodehouse, 1933		1 specimen	A Myriophyllum spicatum-like watermillfoil family palynomorph.
†Rhoipites	†R. bradleyi Wodehouse, 1933		4 specimens	A Rhus typhina-like cashew family palynomorph.
Salix	†S. discoloripites Wodehouse, 1933		Numerous	A Salix discolor and S. fragilis-like willow palynomorph.
†Talisiipites	†T. fischeri Wodehouse, 1933		Numerous	A Talisia depressa-like soapberry family palynomorph.
Tilia	†T. crassipites Wodehouse, 1933		1 specimen	A Tilia americana-like linden palynomorph.
	†T. tetraforaminipites Wodehouse, 1933		3 specimens	A Tilia americana-like linden palynomorph.
	†T. vescipites Wodehouse, 1933		1 specimens	A Tilia americana-like linden palynomorph.
†Vitipites	†V. dubius Wodehouse, 1933		1 specimen	A Vitis vinifera-like grape family palynomorph.

Angiosperms of uncertain affiliation

Genus	Species	Member	Abundance	Notes
†Achaenites	†A. cichorioides Knowlton, 1923			A achene like structure of unidentified affiliation.
†Antholithes	†A. dubia R.W. Brown, 1929	Lake Unita	1 specimen	An 8-lobed/petaled flower of unidentified affiliation.
	†A. improbus Lesquereux, 1883		2 specimens	A flowering structure of unidentified affiliation.
	†A. vitaciflora R.W. Brown, 1929	Lake Unita	1 specimen	A cluster of possible vitaceous flowers-fruits.
	†A. winchesteri (Knowlton) MacGinitie, 1969	Lake Unita	1 specimen	A flowering structure of unidentified affiliation. First described as Sambucus? winchesteri (1923)
	†A. sp 1 R.W. Brown, 1934	Fossil Lake	1 specimen	A possibly cruciferous or orrograceous flower of unidentified affiliation.
	†A. sp 2 R.W. Brown, 1934	Lake Unita	1 specimen	A flower with many stamens of unidentified affiliation.
	†A. sp 3 R.W. Brown, 1934	Lake Unita	1 specimen	A malvaceous looking flower of unidentified affiliation.
	†A. sp 4 R.W. Brown, 1934	Lake Unita	1 specimen	A flower with a superior ovary surrounded by the five-toothed calyx.
	†A. sp 5 R.W. Brown, 1934	Lake Unita	1 specimen	A flower with tubular receptacle surmounted by a four-parted calyx.
	†A. sp 7 R.W. Brown, 1934	Fossil Lake	1 specimen	A large flower or a badly distorted leaf of a Nelumbo?
†Bonanzacarpum	†B. sprungerorum Manchester & Lott, 2019	Lake Uinta (Parachute Creek Member)	Numerous	A eudicot fruit of uncertain phylogenetic placement.
†Carpites	†C. inquirenda Knowlton, 1923	Lake Unita	1 specimen	A nutlet or fruit of unidentified affiliation.
	†C. newberryanus Knowlton, 1923		3 specimens	A nutlet of unidentified affiliation. First described as Juglans occidentalis nuts (1898). First spelled Carpites newberryana
	†C. viburni Lesquereux, 1878		1 specimen	A seed or nutlet of unidentified affiliation. Includes Carpolithes capsularis (1929)
†Carpolithus	†C. alabaccus R.W. Brown, 1934	Lake Unita	1 specimen	A winged or membranous fruit of unidentified affiliation.
	†C. caryophylloides Knowlton, 1923	Lake Unita		A dry capsular fruit of unidentified affiliation.
	†C. cassioides R.W. Brown, 1929	Lake Unita	2 specimens?	A legume like pod of unidentified affiliation.
	†C. insectoides R.W. Brown, 1934	Lake Unita	1 specimen	A winged fruit of unidentified affiliation.
	†C. palrnites R.W. Brown, 1929	Lake Unita	1 specimen	A palm fruit or husk like fossil of unidentified affiliation.
	†C. prunoides R.W. Brown, 1934	Lake Unita	1 specimen	A flattened fruit of unidentified affiliation. First named Carpolithus ellipticus Brown, 1929 (non-Sternberg, 1825)
	†C. serratifolius R.W. Brown, 1929	Lake Unita	1 specimen	Branchlet with leaves and fruits of possible salicaceous affinity.
	†C. vitaceus R.W. Brown, 1934	Lake Unita	1 specimen	A possibly vitaceous seed of unidentified affiliation.
†Caulinites	†C. acanthus R.W. Brown, 1929	Lake Unita	1 specimen	A thorny stem of unidentified affiliation.
†Caulinites	†C. prehensus R.W. Brown, 1929	Lake Unita	1 specimen	A climbing plant tendril of unidentified affiliation.
†Chaneya	†C. tenuis (Lesquereux) Wang & Manchester, 2000	Lake Gosiute Lake Unita		A sapindalean flower of uncertain affiliations. Includes Astronium truncatum (1953).
†Dicotylophyllum	†D. lesquereuxii (Knowlton) J.A. Wolfe & Tanai, 1987		1 specimen	A leaf of uncertain affiliation. First described as Acer indivisum Lesquereux, 1885 (non-Weber, 1852) Moved from Acer lesquereuxii Knowlton, 1898
†Palibinia	†P. comptonifolia (Brown) Manchester, Judd, & Kodrul, 2023	Lake Uinta (Parachute Creek Member)		A pentapetalean eudicot of uncertain affiliation. Moved from Vauquelinia comptonifolia (1969) Originally named Banksia comptonifolia (1934)
†Phyllites	†P. ericoides R.W. Brown, 1934	Lake Unita	1 specimen	A thick leaf of unidentified affiliation.
	†P. fremonti Unger, 1850	Rejected	1 specimen	A leaf of unidentified affiliation. R.W. Brown (1934) rejected due to doubtful locality and lost specimen.
	†P. glandiferus R.W. Brown, 1934	Lake Unita	1 specimen	An odd-pinnately compound leaf of possible fabaceous affinity.
	†P. juncoides R.W. Brown, 1929	Lake Unita	1 specimen	A leaf of unidentified affiliation.
	†P. ouratea R.W. Brown, 1934	Lake Unita	1 specimen	A possibly herbaceous leaf of unidentified affinity.
	†P. winchesteri Knowlton, 1923	Lake Unita	1 specimen	A leaf of unidentified affiliation.

Plants of uncertain affiliation

Genus	Species	Member	Abundance	Notes	Image
†Rhizocaulon	†R. dichotomum R.W. Brown, 1929	Lake Unita	1 specimen	A root of unidentified affiliation. Has a distinct branched fork.
†Rhizocaulon	†R. natans R.W. Brown, 1929	Lake Unita	1 specimen	A rootlet of unidentified affiliation. small with Trapa natans-like rhizoids.

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^ Andrej V. Gorochov; Conrad C. Labandeira (2012). "Eocene Orthoptera from Green River Formation of Wyoming (USA)". Russian Entomological Journal. 21 (4): 357–370. doi:10.15298/rusentj.21.4.02.
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^ Antell, G.S.; Kathirithamby, J. (2016). "The First Twisted-Wing Parasitoids (Insecta: Strepsiptera) from the Early Eocene Green River Formation of Colorado". Bulletin of the Peabody Museum of Natural History. 57 (2): 165–174. doi:10.3374/014.057.0204. S2CID 89188291.
^ Scudder, S.H. (1875). "The Tertiary Physopoda of Colorado". Bulletin of the United States Geological and Geographical Survey of the Territories. 2 (1): 221–223.
Scudder, S.H. (1867). "The results of an examination of a small collection of fossil insects obtained by Prof. William Denton in the Tertiary Beds of Green River, Colorado". Proceedings of the Boston Society of Natural History. 11: 117–118.
Jason L. Robinson; M. Jared Thomas; Sam W. Heads (2018). "A fossil caddisfly (Insecta: Trichoptera) from the Eocene of Colorado". Zootaxa. 4403 (2): 389–394. doi:10.11646/zootaxa.4403.2.10. PMID 29690240.
^ "Abstract: NONMARINE MOLLUSKS OF THE GREEN RIVER FORMATION (EARLY EOCENE), WESTERN INTERIOR, USA (2002 Denver Annual Meeting (October 27-30, 2002))". gsa.confex.com. Retrieved 2024-09-03.
^ Knowlton, F.H. (1923). "Chapter F". Revision of the flora of the Green River Formation, with descriptions of new species (Report). Professional Papers. Vol. 131. United States Geological Survey. pp. 133–182. doi:10.3133/pp131F.
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^ Brown, R.W. (1929). "Chapter J". Additions to the flora of the Green River Formation (Report). Professional Papers. Vol. 154. United States Geological Survey. pp. 279–292. doi:10.3133/pp154J.
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^ Bradley, W. H. (1967). "Two aquatic fungi (Chytridiales) of Eocene age from the Green River Formation of Wyoming". American Journal of Botany. 54 (5): 577–582. doi:10.1002/j.1537-2197.1967.tb10679.x.
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^ Bradley, W. H. (1970). "Eocene algae and plant hairs from the Green River formation of Wyoming". American Journal of Botany. 57 (7): 782–785. doi:10.1002/j.1537-2197.1970.tb09872.x.
^ Bradley, W.H. (1929). "Chapter G". Algae reefs and oolites of the Green River Formation (Report). Professional Papers. Vol. 154. United States Geological Survey. pp. 203–223. doi:10.3133/pp154G.
Steere, W.C. (1947). "Cenozoic and Mesozoic bryophytes of North America". The American Midland Naturalist. 36 (2): 298–324. doi:10.2307/2421507. JSTOR 2421507.
Brown, R.W. (1939). "Some American fossil plants belonging to the Isoetales". Journal of the Washington Academy of Sciences. 29 (6): 261–269. JSTOR 24530230.
^ MacGinitie, H.D. (1969). "The Eocene green River flora of northwestern Colorado and northeastern Utah". University of California Publications in Geological Sciences. 83 (116): 1–202.
^ LaMotte, R.S. (1952). Catalogue of the Cenozoic plants of North America through 1950. Geological Society of America Memoirs. Vol. 51. Geological Society of America. doi:10.1130/MEM51. ISBN 978-0-8137-1051-8.
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^ Wodehouse, R.P. (1933). "Tertiary pollen-II The oil shales of the Eocene Green River Formation". Bulletin of the Torrey Botanical Club. 60 (7): 479–524. doi:10.2307/2480586. JSTOR 2480586.
Brown, R. (1939). "Fossil Leaves, Fruits, and Seeds of Cercidiphyllum". Journal of Paleontology. 13 (5): 485–499. JSTOR 1298516.
"Inaperturopollenites concedipites". IFPNI. Retrieved October 2, 2024.
"Inaperturopollenites vacuipites". IFPNI. Retrieved October 2, 2024.
"Tsugaepollenites viridifluminipites". IFPNI. Retrieved October 2, 2024.
"Zonalapollenites viridifluminipites". IFPNI. Retrieved October 2, 2024.
"Gnetaceae-pollenites eocenipites". IFPNI. Retrieved October 2, 2024.
^ Manchester, S.R.; O’Leary, E.L. (2010). "Phylogenetic distribution and identification of fin-winged fruits". The Botanical Review. 76 (1): 1–82. Bibcode:2010BotRv..76....1M. doi:10.1007/s12229-010-9041-0.
^ Kruse, H.O. "Some Eocene dicotyledonous woods from Eden Valley, Wyoming" (PDF). Ohio Journal of Science. 54 (4): 243–268.
^ Read, R.W.; Hickey, L.J. (1972). "A revised classification of fossil palm and palm-like leaves". Taxon. 21 (1): 129–137. doi:10.2307/1219237. JSTOR 1219237.
^ Tidwell, W.D.; Medlyn, D.A.; Thayn, G.F. (1973). "Three new species of Palmoxylon from the Eocene Green River formation, Wyoming". The Great Basin Naturalist. 33 (2): 61–76.
Allen, S.E. (2015). "Fossil palm flowers from the Eocene of the Rocky Mountain region with affinities to Phoenix L. (Arecaceae: Coryphoideae)". International Journal of Plant Sciences. 176 (6): 586–596. doi:10.1086/681605.
Cockerell, T. D. A. (1923). "The supposed plumage of the Eocene bird Diatryma" (PDF). American Museum Novitates (62): 1–4.
Wetmore, A. (1930). "The Supposed Plumage of the Eocene Diatryma" (PDF). Auk. 47 (4): 579–580. doi:10.2307/4075897. JSTOR 4075897.
Chaney, R.W. (1944). "A fossil cactus from the Eocene of Utah". American Journal of Botany. 31 (8): 507–528. doi:10.1002/j.1537-2197.1944.tb08065.x. JSTOR 2437414.
Brown, R.W. (1959). "Some Paleobotanical Problematica". Journal of Paleontology. 33 (1): 120–124. JSTOR 1300816.
Becker, H. F. (1962). "Reassignment of Eopuntia to Cyperacites". Bulletin of the Torrey Botanical Club. 89 (5): 319–330. doi:10.2307/2482937. JSTOR 2482937.
Herrera, F.; Manchester, S. R. (2024). "Earliest Dioscorea fruits (Dioscoreaceae) from North America". International Journal of Plant Sciences. 185 (5): 474–481. doi:10.1086/729607. S2CID 267284371.
Herendeen, P.S.; Les, D.H.; Dilcher, D.L. (1990). "Fossil Ceratophyllum (Ceratophyllaceae) from the Tertiary of North America". American Journal of Botany. 77 (1): 7–16. doi:10.1002/j.1537-2197.1990.tb13522.x.
Arnold, C.A. (1936). "Some fossil species of Mahonia from the Tertiary of Eastern and Southeastern Oregon". Contributions from the Museum of Paleontology, University of Michigan. 5 (4): 63.
Han, M.; Manchester, S.; Fu, Q.-Y.; Jin, J.-H.; Quan, C. (2018). "Paleogene fossil fruits of Stephania (Menispermaceae) from North America and East Asia". Journal of Systematics and Evolution. 56 (2): 81–91. doi:10.1111/jse.12288. S2CID 90749898.
Manchester, S. R. (1986). "Vegetative and reproductive morphology of an extinct plane tree (Platanaceae) from the Eocene of western North America". Botanical Gazette. 147 (2): 200–226. doi:10.1086/337587. S2CID 83715341.
Manchester, S. R.; Crane, P. R.; Dilcher, D. L. (1991). "Nordenskioldia and Trochodendron fruits (Trochodendraceae) from the Miocene of northwestern North America". Botanical Gazette. 152: 357–368. doi:10.1086/337898. S2CID 84230472.
Alexander B. Doweld (2017). "Allopanax and Boreopanax, New Names of Fossil Araliales". Annales Botanici Fennici. 54 (4–6): 263–266. doi:10.5735/085.054.0608. S2CID 90295525.
Lott, T.A.; Manchester, S.R.; Dilcher, D.L. (1998). "A unique and complete polemoniaceous plant from the middle Eocene of Utah, USA". Review of Palaeobotany and Palynology. 104 (1): 39–49. Bibcode:1998RPaPa.104...39L. doi:10.1016/S0034-6667(98)00048-7.
Deanna, R.; Martínez, C.; Manchester, S.; Wilf, P.; Campos, A.; Knapp, S.; Chiarini, F. E.; Barboza, G. E.; Bernardello, G.; Sauquet, H.; Dean, E.; Orejuela, A.; Smith, S. D. (2023). "Fossil berries reveal global radiation of the nightshade family by the early Cenozoic". New Phytologist. 238 (6): 2685–2697. Bibcode:2023NewPh.238.2685D. doi:10.1111/nph.18904. PMID 36960534. S2CID 257715632.
Manchester, S.; Judd, W.; Correa-Narvaez, J. (2024). "Vegetative and reproductive morphology of Othniophyton elongatum (MacGinitie) gen. et comb. nov., an extinct angiosperm of possible caryophyllalean affinity from the Eocene of Colorado and Utah, USA". Annals of Botany. Accepted manuscript. doi:10.1093/aob/mcae196. PMID 39520137.
^ Manchester, S. R.; Judd, W. S. (2022). "Extinct Anacardiaceous Samaras and Sumac-Like Leaves from the Eocene of Western North America". International Journal of Plant Sciences. 183 (5): 357–366. doi:10.1086/719948. S2CID 247868997.
Herendeen, P.S.; Herrera, F. (2019). "Eocene fossil legume leaves referable to the extant genus Arcoa (Caesalpinioideae, Leguminosae)". International Journal of Plant Sciences. 180 (3): 220–231. doi:10.1086/701468. S2CID 92041312.
^ Herendeen, P. S.; Dilcher, D. L. (1991). "Caesalpinia subgenus Mezoneuron (Leguminosae, Caesalpinioideae) from the Tertiary of North America". American Journal of Botany. 78 (1): 1–12. doi:10.1002/j.1537-2197.1991.tb12566.x. JSTOR 2445223.
Call, V.B.; Dilcher, D.L. (1994). "Parvileguminophyllum coloradensis, a new combination for Mimosites coloradensis Knowlton, Green River Formation of Utah and Colorado". Review of Palaeobotany and Palynology. 80 (3–4): 306. Bibcode:1994RPaPa..80..305C. doi:10.1016/0034-6667(94)90007-8.
^ Manchester, S. (1987). "Fossil History of the Juglandaceae". Annals of the Missouri Botanical Garden. 21: 1–137.
Wing, S. L.; Hickey, L. J. (1984). "The Platycarya perplex and the evolution of the Juglandaceae". American Journal of Botany. 71 (3): 388–411. doi:10.1002/j.1537-2197.1984.tb12525.x.
^ Manchester, S.R.; Dilcher, D.L. (1982). "Pterocaryoid fruits (Juglandaceae) in the Paleogene of North America and their evolutionary and biogeographic significance". American Journal of Botany. 69 (2): 275–286. doi:10.2307/2443015. JSTOR 2443015.
Nareerat Boonchai; Steven R. Manchester; Elisabeth A. Wheeler (2015). "Welkoetoxylon multiseriatum: fossil moraceous wood from the Eocene Green River Formation, Wyoming, U.S.A.". IAWA Journal. 36 (2): 158–166. doi:10.1163/22941932-00000093.
Manchester, S.R.; Dilcher, D.L.; Wing, S.L. (1998). "Attached leaves and fruits of myrtaceous affinity from the Middle Eocene of Colorado". Review of Palaeobotany and Palynology. 102 (3–4): 153–163. Bibcode:1998RPaPa.102..153M. doi:10.1016/S0034-6667(98)80002-X.
Call, V.B.; Dilcher, D.L. (1995). "Fossil Ptelea samaras (Rutaceae) in North America". American Journal of Botany. 82 (8): 1069–1073. doi:10.1002/j.1537-2197.1995.tb11572.x.
^ Manchester, S.R.; Judd, W.S.; Handley, B. (2006). "Foliage and fruits of early poplars (Salicaceae: Populus) from the Eocene of Utah, Colorado, and Wyoming". International Journal of Plant Sciences. 167 (4): 897–908. doi:10.1086/503918.
^ Boucher, L.D.; Manchester, S.R.; Judd, W.S. (2003). "An extinct genus of Salicaceae based on twigs with attached flowers, fruits, and foliage from the Eocene Green River Formation of Utah and Colorado, USA". American Journal of Botany. 90 (9): 1389–1399. doi:10.3732/ajb.90.9.1389. PMID 21659238.
Wang, Q.; Manchester, S. R.; Gregor, H. J.; Shen, S.; Li, Z. Y. (2013). "Fruits of Koelreuteria (Sapindaceae) from the Cenozoic throughout the northern hemisphere: their ecological, evolutionary, and biogeographic implications". American Journal of Botany. 100 (2): 422–449. doi:10.3732/ajb.1200415. PMID 23360930.
^ Correa Narvaez, J. E.; Allen, S. E.; Huegele, I. B.; Manchester, S. R. (2023). "Fossil leaves and fruits of Tetramelaceae (Curcurbitales) from the Eocene of the Rocky Mountain region, USA, and their biogeographic significance". International Journal of Plant Sciences. 184 (3): 177–200. doi:10.1086/724018. S2CID 256185427.
Manchester, S.R. (1989). "Attached reproductive and vegetative remains of the extinct American-European genus Cedrelospermum (Ulmaceae) from the Early Tertiary of Utah and Colorado". American Journal of Botany. 76 (2): 256–276. doi:10.1002/j.1537-2197.1989.tb11309.x.
"Alni-pollenites speciipites". IFPNI. Retrieved October 2, 2024.
"Carya juxtaporipites". IFPNI. Retrieved October 2, 2024.
"Carya viridi-fluminipitis". IFPNI. Retrieved October 2, 2024.
"Juglanspollenites nigripites". IFPNI. Retrieved October 2, 2024.
"Momipites corylipites". IFPNI. Retrieved October 2, 2024.
Steven Manchester; Terry A. Lott (2019). "Bonanzacarpum sprungerorum sp. nov. – a bizarre fruit from the Eocene Green River Formation in Utah, USA" (PDF). Fossil Imprint. 75 (2): 281–288. doi:10.2478/if-2019-0016. S2CID 209378482.
Wang, Y.; Manchester, S. R. (2000). "Chaneya, a new genus of winged fruit from the Tertiary of North America and eastern Asia". International Journal of Plant Sciences. 161 (1): 167–178. doi:10.1086/314227. PMID 10648207. S2CID 45052368.
Wolfe, J.A.; Tanai, T. (1987). "Systematics, Phylogeny, and Distribution of Acer (maples) in the Cenozoic of Western North America". Journal of the Faculty of Science, Hokkaido University. Series 4, Geology and Mineralogy. 22 (1): 1–246.
Manchester, S. R.; Judd, W. S.; Kodrul, T. (2023). "First recognition of the extinct eudicot genus Palibinia in North America: Leaves and fruits of Palibinia comptonifolia (R.W.Br.) comb. nov. from the Eocene of Utah and Colorado, USA". Journal of Systematics and Evolution. 62: 149–163. doi:10.1111/jse.13011.

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