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Tetraceratops Temporal range: Early Permian, 275 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Holotype
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Clade:	Pantherapsida
Genus:	†Tetraceratops Matthew, 1908
Type species
†Tetraceratops insignis Matthew, 1908

Tetraceratops insignis ("four-horned face emblem") is an extinct synapsid from the Early Permian that was formerly considered the earliest known representative of Therapsida, a group that includes mammals and their close extinct relatives. It is known from a single 90-millimetre-long (3.5 in) skull, discovered in Texas in 1908. According to a 2020 study, it should be classified as a primitive non-therapsid sphenacodont rather than a genuine basal therapsid.

Description

Tetraceratops is known from a single 90-millimetre-long (3.5 in) skull discovered in Texas in the early 1900s. Contrary to its genus name, Tetraceratops actually has six horns, one pair being on the premaxilla bones, one pair on the prefrontal bones, and one pair on the angular processes of the mandible. When it was discovered and described in 1908, the skull was still embedded in a matrix, and only the premaxilla and prefrontal pairs were visible. In life, thus, it would have resembled a large lizard with four horns on its snout, and a pair of large spines emanating from the corners of its jaw.

In addition to horns, Tetraceratops also had an impressive set of teeth. The second pair of teeth on the maxillary bones were large and fang-like. Likewise the first teeth in the upper jaw were long and dagger-like.

Classification

Tetraceratops was originally identified as a member of a group called Pelycosauria, an evolutionary grade of synapsids more basal than therapsids. It has been variously grouped in the family Sphenacodontidae, which is closely related to Therapsida, and Eothyrididae, which is more distantly related. Recent phylogenetic studies classify it as either a pelycosaur-grade synapsid or the basalmost therapsid, rendering its exact phylogenetic placement unclear. However, a recent study by Spindler concluded that no convincing morphological evidence could be made for a therapsid placement of Tetraceratops and that this genus was better placed as a basal sphenacodontian.

See also

• List of synapsids
• Evolution of mammals

References

1. Laurin, M.; Reisz. R. R. (1996). "The osteology and relationships of Tetraceratops insignis, the oldest known therapsid". Journal of Vertebrate Paleontology. 16 (1): 95–102. doi:10.1080/02724634.1996.10011287.
2. D. W. Dilkes, R. R. Reisz (1996). "First Record of a Basal Synapsid ('Mammal-Like Reptile') in Gondwana". Proceedings of the Royal Society B. 263 (1374): 1168. doi:10.1098/rspb.1996.0170.
3. ^ Amson, E.; Laurin M. (2011). "On the affinities of Tetraceratops insignis, an Early Permian synapsid". Acta Palaeontologica Polonica. 56 (2): 301–312. doi:10.4202/app.2010.0063.
4. ^ Spindler, Frederik (2020). "The skull of Tetraceratops insignis (Synapsida, Sphenacodontia)" (PDF). Palaeovertebrata. 43 (1): e1. doi:10.18563/pv.43.1.e1.
5. Sidor, C. A.; Hopson, J. A. (1998). "Ghost lineages and "mammalness": assessing the temporal pattern of character acquisition in the Synapsida". Paleobiology. 24 (2): 254–273. doi:10.1666/0094-8373(1998)024[0254:GLAATT]2.3.CO;2. JSTOR 2401242.
6. Conrad, J.; Sidor, C. A. (2001). "Re-evaluation of Tetraceratops insignis (Synapsida: Sphenacodontia)". Journal of Vertebrate Paleontology. 21: 42A. doi:10.1080/02724634.2001.10010852. S2CID 220414868.
7. Liu, J.; Rubidge, B; Li, J. (2009). "New basal synapsid supports Laurasian origin for therapsids". Acta Palaeontologica Polonica. 54 (3): 393–400. doi:10.4202/app.2008.0071.

External links

• Palaeos.com's article on therapsids
• kheper.net

v t e Synapsida
Kingdom: Animalia Phylum: Chordata Superclass: Tetrapoda Clade: Amniota Tetrapodomorpha see Tetrapodomorpha Reptiliomorpha see Reptiliomorpha Synapsida see below↓
Synapsida Synapsida †Diadectomorpha? †Caseasauria? Eothyrididae Asaphestera? Eothyris Oedaleops Vaughnictis Caseidae Alierasaurus Angelosaurus Arisierpeton Callibrachion Casea Caseoides Caseopsis Cotylorhynchus Datheosaurus Ennatosaurus Eocasea Euromycter Lalieudorhynchus Martensius Oromycter Phreatophasma Ruthenosaurus Trichasaurus Eupelycosauria †Varanopidae? Apsisaurus Archaeovenator Ascendonanus Eoscansor Dendromaia Nitosaurus? Pyozia Thrausmosaurus Mesenosaurinae Anningia Cabarzia Elliotsmithia Heleosaurus Mesenosaurus Microvaranops Varanodontinae Aerosaurus Mycterosaurus? Ruthiromia Tambacarnifex Varanops Varanodon Watongia Metopophora see below↓ Incertae sedis †Dimacrodon †Driveria †Gorgodon †Knoxosaurus †Mastersonia †Protoclepsydrops?
Metopophora †Ophiacodontidae Archaeothyris Baldwinonus Clepsydrops Echinerpeton Ophiacodon Stereophallodon Stereorhachis Varanosaurus Haptodontiformes †Ianthodon †Milosaurus †Edaphosauridae Bohemiclavulus Edaphosaurus Glaucosaurus Gordodon Ianthasaurus Lupeosaurus Melanedaphodon Ramodendron Remigiomontanus Xyrospondylus Sphenacodontia †Cutleria †Haptodus †Hypselohaptodus †Kenomagnathus †Tetraceratops †Palaeohatteriidae Palaeohatteria Pantelosaurus Sphenacodontoidea †Shashajaia †Sphenacodontidae Cryptovenator Ctenorhachis Ctenospondylus Cutleria? Dimetrodon Eosyodon Macromerion Neosaurus Secodontosaurus Sphenacodon Steppesaurus? Tappenosaurus? Tetraceratops? Therapsida see below↓
Therapsida Therapsida †Raranimus †Biarmosuchia see Biarmosuchia †Dinocephalia see Dinocephalia †Anomodontia see Anomodontia includes dicynodonts †Gorgonopsia see Gorgonopsia Eutheriodontia †Therocephalia see Therocephalia Cynodontia see Cynodontia includes mammals Incertae sedis †Dinosaurus †Kamagorgon †Niaftasuchus
See also: Pelycosauria Phthinosuchia Theriodontia Category

• Sphenacodonts
• Cisuralian synapsids of North America
• Fossil taxa described in 1908
• Taxa named by William Diller Matthew
• Kungurian genus first appearances
• Kungurian genus extinctions