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Harrytoombsia Temporal range: Late Devonian: Frasnian, 382.7–372.2 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Model at the Naturhistoriska Riksmuseet in Stockholm
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	†Placodermi
Order:	†Arthrodira
Suborder:	†Brachythoraci
Clade:	†Eubrachythoraci
Clade:	†Coccosteomorphi
Superfamily:	†Incisoscutoidea
Genus:	†Harrytoombsia Miles and Dennis, 1979
Species
Harrytoombsia elegans Miles and Dennis, 1979 (type)

Harrytoombsia is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found from Kimberley, Australia. It was a fast-swimming predator, and it had sharp blades of enamel-like material around the outer edge of the jaw for catching prey.

Phylogeny

Harrytoombsia is a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Harrytoombsia:

References

1. "Fossilworks: Harrytoombsia". fossilworks.org. Archived from the original on 2021-12-12. Retrieved 2021-12-17.
2. ^ Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society. 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082.
3. "Visar vilket djur 1 - NyaUtställningar". utstallning.nrm.se.
4. You-An Zhu; Min Zhu (2013). "A redescription of Kiangyousteus yohii (Arthrodira: Eubrachythoraci) from the Middle Devonian of China, with remarks on the systematics of the Eubrachythoraci". Zoological Journal of the Linnean Society. 169 (4): 798–819. doi:10.1111/zoj12089.

v t e Arthrodira
Kingdom: Animalia Phylum: Chordata Subphylum: Vertebrata Infraphylum: Gnathostomata Class: "Placodermi" Gnathostomata see Gnathostomata Placodermi see Placodermi Arthrodira see below↓
Arthrodira Arthrodira Aethaspis Aleosteus Anarthraspis Antarctaspis Baringaspis Bryantolepis Erikaspis Eskimaspis Heightingtonaspis Kujdanowiaspis Lataspis Lehmanosteus Mediaspis Proaethaspis Qataraspis Sigaspis Simblaspis Yujiangolepis Wuttagoonaspidae Wuttagoonaspis Yiminaspis Phyllolepida Gavinaspis Phyllolepididae Austrophyllolepis Cobandrahlepis Cowralepis Phyllolepis Placolepis Yurammia Actinolepidae Actinolepis Ailuracantha Bollandaspis Szelepis Phlyctaenioidei Dicksonosteus Phlyctaeniidae Aggeraspis Arctaspis Barwickosteus Cartieraspis Diadsomaspis Elegantaspis Gaspeaspis Grifftaylor Heterogaspis Huginaspis Kolpaspis Kueichowlepis Lyhoalepis Neophlyctaenius Pageauaspis Phlyctaenius Prosphymaspis Stipatosteus Arctolepididae Arctolepis Heintzosteus Huginaspis? Kueichowlepis? Groenlandaspididae Africanaspis Boomeraspis Cosmacanthus Groenlandaspis Mithakaspis Mulgaspis Tiaraspis Turrisaspis Williamsaspididae Elvaspis Williamsaspis Brachythoraci Antineosteus Dhanguura? Gemuendenaspis Maideria Xiangshuiosteus? Holonematidae Artesonema Belemnacanthus Bimbianga Deirosteus Deveonema Gyroplacosteus Holonema Megaloplax Rhenonema Tropidosteus Tropinema Homostiidae Angarichthys Cathlesichthys Dhanguura? Euleptaspis Euleptaspis Luetkeichthys Migmatocephala Antineosteus Atlantidosteus Homosteus Tityosteus Buchanosteidae Arenipiscis Buchanosteus Burrinjucosteus Errolosteus Exutaspis Goodradigbeeon Narrominaspis Parabuchanosteus Toombsosteus Taemasosteus Uralosteus Eubrachythoraci see below↓
Eubrachythoraci Coccosteomorphi Coccosteoidea Coccosteidae Belgiosteus Clarkeosteus Coccosteus Dickosteus Jiuchengia Livosteus Millerosteus Protitanichthys Watsonosteus Panxiosteidae Janiosteus Kimberleyichthys? Panxiosteus Plourdosteus? Incisoscutoidea Belosteus? Compagopiscis Cyrtosteus? Harrytoombsia? Mcnamaraspis? Parabelosteus? Torosteus Trematosteus Camuropiscidae Camuropiscis Fallacosteus Incisoscutum Latocamurus Rolfosteus Tubonasus Pachyosteomorphi Leptosteus Rhachiosteus Dunkleosteoidea Dunkleosteus Eastmanosteus Golshanichthys Hussakofia Kiangyousteus Westralichthys Xiangshuiosteus? Heterosteidae Herasmius Heterosteus Yinostius Aspinothoracidi Bruntonichthys Bullerichthys Bungartius Dinichthys Dinomylostoma Erromenosteus Hadrosteus Holdenius Kendrickichthys Mylostoma Squamatognathus Tafilalichthys Tapinosteus Titanichthys Brachydeiridae Brachydeirus Oxyosteus Synauchenia Selenosteidae Alienacanthus Amazichthys Brachyosteus Braunosteus Draconichthys Enseosteus Gorgonichthys Gymnotrachelus Heintzichthys Melanosteus Microsteus Pachyosteus Paramylostoma Phymosteus Rhinosteus Selenosteus Stenosteus Walterosteus

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