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Heightingtonaspis Temporal range: Early - Middle Devonian, 419–382 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	†Placodermi
Order:	†Arthrodira
Genus:	†Heightingtonaspis White, 1969
Species
†Heightingtonaspis anglica (Traquair, 1890) (type) †Heightingtonaspis clarkei (Eastman, 1907) †Heightingtonaspis willsi (White, 1961)
Synonyms
Kujdanowiaspis clarkei Eastman, 1907 Kujdanowiaspis willsi White, 1961 Phlyctaenius anglicus Traquair, 1890

Heightingtonaspis is an extinct genus of primitive arthrodire placoderm fish from the Devonian period in Great Britain, and currently contains three species.

The type species Heightingtonaspis anglica was described by Ramsay Traquair in 1890, and was initially named Phlyctaenius anglicus. The holotype specimen NHMUK PAL PV P 29417 was first discovered on the surface at the Old Red Sandstone supergroup, Shropshire, England and the fossils were radiocarbon dated in situ to 416 million years old. it has also been found in the Senni Beds of Wales.

Heightingtonaspis clarkei was described by Charles R. Eastman in 1907, and was originally named Kujdanowiaspis clarkei. It was later revised and reassigned to the genus Heightingtonaspis by Denison in 1978.

Heightingtonaspis willsi was described by White in 1961, and was originally named Kujdanowiaspis willsi, but was later reassigned to the genus Heightingtonaspis, which was created in 1969 by White. H. willsi was discovered in the same location as H. anglica, in Shropshire, England.

Heightingtonaspis is one of the more basal members of the order Arthrodira, as shown in the cladogram below:

References

1. ^ Robert Denison. (1978). Placodermi. In H.-P. Schultze (ed.), Handbook of Palaeoichthyology 2:1-128
2. ^ Dineley D. L., Metcalfe S. J. (1999). Chapter 4. Early Devonian fossil fishes sites of the Welsh Borders, Fossil Fishes of Great Britain, 2731
3. ^ White, E.I. (1969). The deepest vertebrate fossil and other arctolepid fishes. Biological Journal of the Linnean Society 1(3): 293-310
4. ^ Dupret, Vincent (2010). "Revision of the genus Kujdanowiaspis Stensiö, 1942 (Placodermi, Arthrodira, "Actinolepida") from the Lower Devonian of Podolia (Ukraine)". Geodiversitas. 32 (1): 5–63. doi:10.5252/g2010n1a1. S2CID 129505974.
5. Dupret, V.; Zhu, M. I. N.; Wang, J. N. Q. (2009). "The morphology of Yujiangolepis liujingensis (Placodermi, Arthrodira) from the Pragian of Guangxi (south China) and its phylogenetic significance" (PDF). Zoological Journal of the Linnean Society. 157: 70. doi:10.1111/j.1096-3642.2009.00519.x.

v t e Arthrodira
Kingdom: Animalia Phylum: Chordata Subphylum: Vertebrata Infraphylum: Gnathostomata Class: "Placodermi" Gnathostomata see Gnathostomata Placodermi see Placodermi Arthrodira see below↓
Arthrodira Arthrodira Aethaspis Aleosteus Anarthraspis Antarctaspis Baringaspis Bryantolepis Erikaspis Eskimaspis Heightingtonaspis Kujdanowiaspis Lataspis Lehmanosteus Mediaspis Proaethaspis Qataraspis Sigaspis Simblaspis Yujiangolepis Wuttagoonaspidae Wuttagoonaspis Yiminaspis Phyllolepida Gavinaspis Phyllolepididae Austrophyllolepis Cobandrahlepis Cowralepis Phyllolepis Placolepis Yurammia Actinolepidae Actinolepis Ailuracantha Bollandaspis Szelepis Phlyctaenioidei Dicksonosteus Phlyctaeniidae Aggeraspis Arctaspis Barwickosteus Cartieraspis Diadsomaspis Elegantaspis Gaspeaspis Grifftaylor Heterogaspis Huginaspis Kolpaspis Kueichowlepis Lyhoalepis Neophlyctaenius Pageauaspis Phlyctaenius Prosphymaspis Stipatosteus Arctolepididae Arctolepis Heintzosteus Huginaspis? Kueichowlepis? Groenlandaspididae Africanaspis Boomeraspis Cosmacanthus Groenlandaspis Mithakaspis Mulgaspis Tiaraspis Turrisaspis Williamsaspididae Elvaspis Williamsaspis Brachythoraci Antineosteus Dhanguura? Gemuendenaspis Maideria Xiangshuiosteus? Holonematidae Artesonema Belemnacanthus Bimbianga Deirosteus Deveonema Gyroplacosteus Holonema Megaloplax Rhenonema Tropidosteus Tropinema Homostiidae Angarichthys Cathlesichthys Dhanguura? Euleptaspis Euleptaspis Luetkeichthys Migmatocephala Antineosteus Atlantidosteus Homosteus Tityosteus Buchanosteidae Arenipiscis Buchanosteus Burrinjucosteus Errolosteus Exutaspis Goodradigbeeon Narrominaspis Parabuchanosteus Toombsosteus Taemasosteus Uralosteus Eubrachythoraci see below↓
Eubrachythoraci Coccosteomorphi Coccosteoidea Coccosteidae Belgiosteus Clarkeosteus Coccosteus Dickosteus Jiuchengia Livosteus Millerosteus Protitanichthys Watsonosteus Panxiosteidae Janiosteus Kimberleyichthys? Panxiosteus Plourdosteus? Incisoscutoidea Belosteus? Compagopiscis Cyrtosteus? Harrytoombsia? Mcnamaraspis? Parabelosteus? Torosteus Trematosteus Camuropiscidae Camuropiscis Fallacosteus Incisoscutum Latocamurus Rolfosteus Tubonasus Pachyosteomorphi Leptosteus Rhachiosteus Dunkleosteoidea Dunkleosteus Eastmanosteus Golshanichthys Hussakofia Kiangyousteus Westralichthys Xiangshuiosteus? Heterosteidae Herasmius Heterosteus Yinostius Aspinothoracidi Bruntonichthys Bullerichthys Bungartius Dinichthys Dinomylostoma Erromenosteus Hadrosteus Holdenius Kendrickichthys Mylostoma Squamatognathus Tafilalichthys Tapinosteus Titanichthys Brachydeiridae Brachydeirus Oxyosteus Synauchenia Selenosteidae Alienacanthus Amazichthys Brachyosteus Braunosteus Draconichthys Enseosteus Gorgonichthys Gymnotrachelus Heintzichthys Melanosteus Microsteus Pachyosteus Paramylostoma Phymosteus Rhinosteus Selenosteus Stenosteus Walterosteus

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