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Shantungosuchus Temporal range: 129–110 Ma PreꞒ Ꞓ O S D C P T J K Pg N Early Cretaceous
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Archosauria
Clade:	Pseudosuchia
Clade:	Crocodylomorpha
Clade:	Crocodyliformes
Clade:	Mesoeucrocodylia
Clade:	†Shartegosuchoidea
Genus:	†Shantungosuchus Yang, 1961
Species
†S. chuhsienensis Young, 1961 (type) †S. hangjinensis Wu et al, 1994 †S. brachycephalus Young, 1982

Shantungosuchus is an extinct genus of Early Cretaceous crocodyliform found in China. It includes three species: Shantungosuchus chuhsienensis and S. brachycephalus, which were both described by Yang Zhongjian – usually referred to as "Young" – in 1961 and 1982, and S. hangjinensis, which was described by Xiao-Chun Wu et al in 1994. S. chuhsienensis is the type for this genus.

Etymology

The primary part of Shantungosuchus' name comes from Shan-tung, the Wade-Giles romanization of Shandong (simplified Chinese: 山东; traditional Chinese: 山東; pinyin: Shāndōng; Wade–Giles: Shantung), a province located on the eastern coast of the People's Republic of China, where it was first discovered. The second part, suchus is an Ancient Greek word referring to the Egyptian crocodile deity Sobek that is commonly used as a suffix for crocodylomorph genera and crocodile-like animals in general.

Description

Shantungosuchus chuhsienenis was first described from an articulated skeleton that was preserved as an impression of its ventral surface. Its small size, slender body, and triangular skull made it distinct from other atoposaurids. Its total length is only about 30 cm (12 in).

Shantungosuchus belongs to Protosuchia, a group of early crocodilian relatives that were all rather small in size, about 1 meter in length, and terrestrial rather than aquatic. They are most easily distinguished from other crocodylians by their short premaxilla and maxillas, a transversely broad shelf on the jugal bone, and a pair of posterolaterally divergent ridges on the pterygoid, and two large depressions on the sphenoid bone. The angular shape of the rest of the skull is absent from the posterolateral section of the jaw. The dentaries near the symphysis are superficially asymmetrical. All known forms of Shantungosuchus also have a square-shaped fossa on the jugal, and a leaf-shaped palatine bone.

Taxonomy

Two of the species of Shantungosuchus, including the type, were described by Yang Zhongjian (referenced here as C.C. (Chung Chien) Young) in 1961. Since the description of S. chuhsienensis, there has been uncertainty about the placement of the genus. It was suggested to be an atoposaurid by several different authors from the 1960s to the 1980s,. However, in Xiao-Chun Wu's 1994 description of S. hangjinensis, he noted the bones were more similar to the family Protosuchidae rather than Atoposauridae, and proposed to classify it as such.

Protosuchia has recently been considered a paraphyletic grouping of early crocodyliforms. Protosuchia is now commonly split into two groups: the basal family Protosuchidae, which is a true clade, and a group of basal crocodyliforms more closely related to Hsisosuchus and Mesoeucrocodylia. Clark et al. (2004) recovered Shantungosuchus as closely related to mesoeucrocodylians, and Buscalioni (2017) found the genus to group in a clade with Sichuanosuchus and Shartegosuchidae, which was formally named Shartegosuchoidea by Dollman et al. (2018) in their description of a new Shartegosuchus specimen.

Distribution and habitat

Shantungosuchus chuhsienensis has been found in the Early Cretaceous (Barremian-Aptian) Mengyin Formation of Shandong province. S. hangjinensis, however, is slightly younger, and is found in the lower Cretaceous Luohandong Formation of Outer Mongolia, and S. brachycephalus may also have been found in the same region. The Luohandong formation provides clues to the environment of Shantungosuchus. For instance, S. hangjinensis likely shared its habitat with fish such as Sinamia, the turtle Ordosemys, and another species of crocodile-like reptile called Ikechosaurus, and several types of dinosaurs- though none have been identified to the level of genus.

See also

• List of crurotarsans
• Sichuanosuchus

References

1. ^ Wu, Xiao-Chun. Donald B. Brinkman, and Jun-Chang Lu. A new species Shantungosuchus from the Lower Cretaceous of Inner Mongolia (China) with comments on S. chuhsiensis Young, 1961, and the phylogenetic position of the genus. Journal of Vertebrate Paleontology, 1994, 14(2):210-229.
2. Matsumoto, R.; Evans, S. E. (2010-12-10). "Choristoderes and the freshwater assemblages of Laurasia". Journal of Iberian Geology (in Spanish). 36 (2): 253–274. ISSN 1886-7995.
3. Colbert, Edwin H., Morales, Michael. Minkoff, Eli C., Colbert's Evolution of the Vertebrates: A History of the Backboned Animals through Time. Fifth Edition. 2001. Wiley-Liss. p.183
4. Buffetaut, E. (1982). Radiation évolutive, paléoécologie et biogéographie des Crocodiliens mésosuchiens. Mémoires de la Société Géologique de France, 60, 1-88.
5. Clark, J.M. (1985). A new crocodylomorph from the Late Jurassic Morrison Formation of western Colorado, with a discussion of relationships within the "Mesosuchia". Master's thesis, Univ. of California, Berkeley, 85 pp.
6. Buscaloni , Angela D. and José Luis Sanz. (1988). PHYLOGENETIC RELATIONSHIPS OF THE ATOPOSAURIDAE (ARCHOSAURIA, CROCODYLOMORPHA). Historical Biology, 1988. Vol. 1. pp. 233-250.
7. Pol, D.; Ji, S.; Clark, J.M.; Chiappe, L.M. (2004). "Basal crocodyliforms from the Lower Cretaceous Tugulu Group (Xinjiang, China), and the phylogenetic position of Edentosuchus" (PDF). Cretaceous Research. 25 (4): 603–622. doi:10.1016/j.cretres.2004.05.002.
8. A. D. Buscalioni. 2017. The Gobiosuchidae in the early evolution of Crocodyliformes. Journal of Vertebrate Paleontology e1324459:1-21.
9. Kathleen N. Dollman; James M. Clark; Mark A. Norell; Xu Xing; Jonah N. Choiniere (2018). "Convergent evolution of a eusuchian-type secondary palate within Shartegosuchidae". American Museum Novitates. 3901: 1–23. doi:10.1206/3901.1.
10. Lü, J. & Wu, X-C. 1996 Restudy of Shantungosuchus brachycephalus Young, 1982. Vert. PalAs. 34, 184-201.
11. Lucas, Spencer G., 2001. Chinese Fossil Vertebrates. New York. University of Columbia Press. p. 169.
12. Weishampel, et al. (2004). "Dinosaur distribution." Pp. 517-607.

External links

• Paleobiology database: S. chuhsienensis

v t e Pseudosuchia
Kingdom: Animalia Phylum: Chordata Class: Sauropsida Clade: Archosauria Sauropsida see Sauropsida Archosauromorpha see Archosauromorpha Pseudosuchia see below↓
Pseudosuchia Pseudosuchia †Dasygnathoides †Nundasuchus †Phytosauria? †Ornithosuchidae Aenigmaspina? Dynamosuchus Ornithosuchus Riojasuchus Venaticosuchus Suchia †Heteropelta? †Koilamasuchus? †Mandasuchus †Nundasuchus? †Ticinosuchus †Erpetosuchidae Archeopelta? Dyoplax? Erpetosuchus Pagosvenator Parringtonia Tarjadia †Aetosauriformes Acaenasuchus Euscolosuchus Revueltosaurus Aetosauria see Aetosauria †Gracilisuchidae Gracilisuchus Maehary? Parvosuchus Turfanosuchus Yonghesuchus Paracrocodylomorpha see below↓
Paracrocodylomorpha Paracrocodylomorpha †Mambawakale †Poposauroidea Benggwigwishingasuchus Lotosaurus Schultzsuchus? Qianosuchus Ctenosauriscidae Arizonasaurus Bromsgroveia Bystrowisuchus Ctenosauriscus Hypselorhachis Tsylmosuchus? Xilousuchus Poposauridae Dolichobrachium Poposaurus Shuvosauridae Effigia Shuvosaurus Sillosuchus Loricata †Apatosuchus †Arganasuchus †Batrachotomus †Dagasuchus †Decuriasuchus †Etjosuchus †Fasolasuchus †Heptasuchus †Heteropelta? †Incertovenator? †Luperosuchus †Mandasuchus? †Prestosuchus †Procerosuchus †Saurosuchus †Stagonosuchus †Youngosuchus †Rauisuchidae Arganasuchus? Energosuchus? Jushatyria? Polonosuchus Postosuchus Procerosuchus? Rauisuchus Scolotosuchus Scythosuchus? Teratosaurus Tikisuchus? Vivaron Vjushkovisaurus? Vytshegdosuchus? Crocodylomorpha see below↓
Crocodylomorpha Crocodylomorpha †Carnufex †Dibothrosuchus †Dromicosuchus †Hesperosuchus †Heteropelta? †Junggarsuchus †Kayentasuchus †Parrishia †Phyllodontosuchus †Pseudhesperosuchus †Redondavenator †Sphenosuchus †Syntomiprosopus? †Trialestes †Saltoposuchidae Litargosuchus Saltoposuchus Terrestrisuchus †Hallopodidae Almadasuchus Hallopus Macelognathus †Thalattosuchia see Thalattosuchia Crocodyliformes †Entradasuchus †Eopneumatosuchus †Kemkemia †Microchampsa †Platyognathus †Stegomosuchus †Tagarosuchus †Protosuchidae Coloradisuchus Dianosuchus Edentosuchus Erythrochampsa Hemiprotosuchus Hoplosuchus Orthosuchus? Pedeticosaurus? Protosuchus †Notochampsidae Notochampsa Orthosuchus? †Gobiosuchidae Cassissuchus Gobiosuchus Lisboasaurus? Zaraasuchus Mesoeucrocodylia †Dianchungosaurus †Hsisosuchus †Induszalim †Lusitanisuchus †Neuquensuchus †Pachycheilosuchus? †Titanochampsa †Thalattosuchia? †Shartegosuchoidea Hadongsuchus Shantungosuchus Sichuanosuchus Zosuchus Shartegosuchidae Adzhosuchus Fruitachampsa Kyasuchus Nominosuchus Shartegosuchus Metasuchia †Ziphosuchia see Ziphosuchia includes notosuchians Neosuchia see Neosuchia includes crocodilians
See also: Rauisuchia Prestosuchidae Bathyotica Sphenosuchia Protosuchia Mesosuchia Category

• Crocodyliforms
• Early Cretaceous crocodylomorphs of Asia
• Berriasian life
• Valanginian life
• Cretaceous China
• Fossils of China
• Paleontology in Shandong
• Fossil taxa described in 1961
• Prehistoric pseudosuchian genera