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Bolboceratidae Temporal range: Aptian–Present PreꞒ Ꞓ O S D C P T J K Pg N
An unidentified bolboceratid
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Superfamily:	Scarabaeoidea
Family:	Bolboceratidae Mulsant, 1842
Synonyms
Bolboceratinae

Bolboceratidae is a family of beetle. It was historically treated as a subfamily of the earth-boring dung beetles, or "dor beetles" (family Geotrupidae), but has been considered a separate family by many authors since 1995. Some recent classifications have persisted in treating bolboceratids as a subfamily (e.g.) but these classifications are contradicted by recent phylogenetic studies of relationships indicating that bolboceratids are not closely related to geotrupids (e.g., that bolboceratids are more closely related to Pleocomidae and Passalidae).

Genera

• Athyreus
• Australobolbus
• Blackbolbus
• Blackburnium
• Bolbaffer
• Bolbaffroides
• Bolbelasmus
• Bolbobaineus
• Bolbocaffer
• Bolbocerastes
• Bolboceratex
• Bolboceratops
• Bolbocerodema
• Bolboceroides
• Bolbocerosoma
• Bolbochromus
• Bolbogonium
• Bolbohamatum
• Bolboleaus
• Bolborhachium
• Bolborhinum
• Bolborhombus
• Bolbothyreus
• Bolbotrypes
• Bradycinetulus
• Cretobolbus
• Elephastomus
• Eubolbitus
• Eucanthus
• Gilletinus
• Halffterobolbus
• Meridiobolbus
• Mesoathyreus
• Mimobolbus
• Namibiobolbus
• Namibiotrupes
• Neoathyreus
• Odonteus
• Parabolbapium
• Parathyreus
• Pereirabolbus
• Prototrupes
• Pseudoathyreus
• Socotrabolbus
• Somalobolbus
• Senaspidius
• Zefevazia

Fossil genera

• †Amberathyreus Bai and Zhang 2017 Burmese amber, Myanmar, Cenomanian
• †Cretobolbus Nikolajev 1996, Zaza Formation, Russia, Aptian

References

1. Gomes Gonçalves, Marcos Paulo (December 2017). "Relationship Between Meteorological Conditions and Beetles in Mata de Cocal". Revista Brasileira de Meteorologia. 32 (4): 543–554. doi:10.1590/0102-7786324003. ISSN 0102-7786.
2. Bolboceratidae at TOL
3. Smith, Andrew B. T. (2006). "A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification". The Coleopterists Bulletin. 60 (sp5). Coleopterists Society: 144–204. doi:10.1649/0010-065x(2006)60[144:arotfn]2.0.co;2. ISSN 0010-065X.
4. Bouchard, P.; Bousquet, Y.; Davies, A.; Alonso-Zarazaga, M.; Lawrence, J.; Lyal, C.; Newton, A.; Reid, C.; Schmitt, M.; Ślipiński, A.; Smith, A. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. Bibcode:2011ZooK...88....1B. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.
5. Mckenna, Duane D.; Wild, Alexander L.; Kanda, Kojun; Bellamy, Charles L.; Beutel, Rolf G.; et al. (2015). "The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution". Systematic Entomology. 40 (4): 835–880. Bibcode:2015SysEn..40..835M. doi:10.1111/syen.12132. hdl:10057/11540. ISSN 0307-6970.

External links

• Encyclopedia of Life entry

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• Bolboceratidae
• Beetle families
• Scarabaeoidea stubs