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Hydraenidae Temporal range: Early Jurassic–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Staphyliniformia
Superfamily:	Staphylinoidea
Family:	Hydraenidae Mulsant, 1844
Subfamilies
Hydraeninae Mulsant, 1844 Ochthebiinae C. G. Thomson, 1860 Orchymontiinae Perkins, 1997 Prosthetopinae Perkins, 1994

Hydraenidae is a family of very small aquatic beetles, sometimes called "Minute moss beetles", with a worldwide distribution. They are around 0.8 to 3.3 mm in length. The adults store air on the underside of the body as well as beneath the elytra which allows them to crawl underwater, often on the underside of the water surface tension, though they cannot swim. Some species have gills that effectively allow them to stay underwater indefinitely. Larvae vary from being fully terrestrial, to being aquatic at least in their earliest instars. The diet of hydraenid larvae and adults is thought to consist of algae, spores and other plant matter. At least some hydraenid adults use stridulation to communicate. There are around 1,300 species in 42 genera.

Genera

These 56 genera belong to the family Hydraenidae:

• Adelphydraena Perkins, 1989
• Archaeodraena Jäch & Yamamoto, 2017
• Aulacochthebius Kuwert, 1887
• Calobius Wollaston, 1854
• Cobalius Rey, 1886
• Coelometopon Janssens, 1972
• Davidraena Jäch, 1994
• Decarthrocerus Orchymont, 1948
• Discozantaena Perkins & Balfour-Browne, 1994
• Edaphobates Jäch & Díaz, 2003-01
• Enicocerus Stephens, 1829
• Ginkgoscia Jäch & Díaz, 2004-01
• Gondraena Jäch, 1994
• Gymnanthelius Perkins, 1997
• Gymnochthebius Orchymont, 1943
• Haptaenida Perkins, 1997
• Heptaenida Perkins, 1997
• Homalaena Ordish, 1984
• Hughleechia Perkins, 1981
• Hydraena Kugelann, 1794
• Hydraenida Germain, 1901
• Hydroenida Germain, 1901
• Hymenodes Mulsant, 1844
• Laeliaena Sahlberg, 1900
• Limnebius Leach, 1815
• Madagaster Perkins, 1997
• Menomadraena Perkins, 2017
• Meropathus Enderlein, 1901
• Micragasma Sahlberg, 1900
• Neochthebius Orchymont, 1932
• Nucleotops Perkins & Balfour-Browne, 1994
• Ochtebiites Ponomarenko, 1977
• Ochthebius Leach, 1815
• Ochtheosus Perkins, 1997
• Oomtelecopon Perkins, 2005
• Orchymontia Broun, 1919
• Parasthetops Perkins & Balfour-Browne, 1994-31
• Parhydraena Orchymont, 1937
• Parhydraenopsis Perkins, 2009-16
• Phothydraena Kuwert, 1888
• Pneuminion Perkins, 1997
• Podaena Ordish, 1984
• Prionochthebius Kuwert, 1887
• Prosthetops Waterhouse, 1879
• Protochthebius Perkins, 1997
• Protosthetops Perkins, 1994
• Protozantaena Perkins, 1997
• Pterosthetops Perkins, 1994
• Sebasthetops Jäch, 1998-01
• Sicilicula Balfour-Browne, 1958
• Spanglerina Perkins, 1980
• Trinomadraena Perkins, 2017
• Tympallopatrum Perkins, 1997
• Mesoceration Janssens, 1967
• Parhydraenida Balfour-Browne, 1975
• Tympanogaster Janssens, 1967

Extinct genera

• †Archaeodraena Jäch and Yamamoto 2017, Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
• †Ochtebiites Ponomarenko 1977 Abasheva Formation, Russia, Early Jurassic (Pliensbachian), Ichetuy Formation, Russia, Karabastau Formation, Kazakhstan, Late Jurassic (Oxfordian), Mogotuin Formation, Mongolia, Early Cretaceous (Aptian)

References

1. Jäch, Manfred A., Beutel, Rolf G., Delgado, Juan A. and Díaz, Juan A.. "Hydraenidae Mulsant, 1844: Coleoptera, Beetles". Handbook of Zoology Online, edited by Andreas Schmidt-Rhaesa. Berlin, Boston: De Gruyter, 2016. Accessed 2022-10-02.
2. Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. Bibcode:2011ZooK...88....1B. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
3. "Hydraenidae". GBIF. Retrieved 16 June 2019.
4. "Hydraenidae Report". Integrated Taxonomic Information System. Retrieved 16 June 2019.

External links

• Data related to Hydraenidae at Wikispecies
• Hydraenidae, Identification and Ecology of Australian Freshwater Invertebrates.
• Hydraenidae at Tolweb

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