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Cynarctus Temporal range: Middle Miocene–Late Miocene PreꞒ Ꞓ O S D C P T J K Pg N
Cynarctus Skull
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Subfamily:	†Borophaginae
Tribe:	†Borophagini
Subtribe:	†Cynarctina
Genus:	†Cynarctus Matthew, 1902
Type species
†Cynarctus saxatilis
Species
"C, acridens", Barbour and Cook 1914 C. crucidens, Barbour and Cook 1914 "C. fortidens", Hall "et al." 1962 C. galushai, Wang et al. 1999 C. marylandica, Berry 1938 C. saxatilis, Matthew 1902 C. voorhiesi, Wang wt al. 1999 C. wangi, Jasinski & Wallace 2015

Cynarctus is an extinct genus of the Borophaginae subfamily of canids native to North America. The genus was first founded by W. D. Matthew in 1901, based from a pair of lower jaws, Cynarctus saxitilis, found in the Pawnee Creek Beds of Colorado. It lived during the Middle to Late Miocene 16.0—10.3 mya, existing for approximately 5.7 million years. Fossils have been uncovered in Colorado, California, Maryland, western Nebraska, and Texas. It was likely an omnivore, and lacked the bone-cracking adaptations found in some later borophagines. Newer findings have proved the genus to be described as a large dog-like raccoon, a result from combining characteristics from Canidae with Procyonidae.

Species

Cynarctus crucidens

Found by Mr. A. C. Whitford, from the Department of Geology at The University of Nebraska, and collected for the Nebraska Geological Survey in 1913. This species of Cynarctus is represented by a right lower jaw that was located in beds (Lower Pliocene) in Sioux County, Nebraska. Having an almost complete dentition, it was found to have slight differences in the structure of the mouth/jaw area from the species of C. saxatilis. Findings confirmed that the premolars were more reduced, teeth were narrower, canine smaller, and that the jaw was relatively longer and more slender than species C. saxatilis.

Cynarctus acridens

Collected by Erwin H. Barbour and Harold J. Cook, for the same Nebraska Geological Survey in 1913, as the species C. crucidens was found. This species is also represented by a part of a lower right jaw and is considered to be the smallest that has so far been discovered. It differs from the species C. saxitilis and C. crucidens, whom have a heavy and broad posterior cinguli, by having a comparatively very weak posterior cingulum.

Cynarctus fortidens

A part of a right maxilla (Early Pliocene age) was located 75 feet above stream on the west side of Turkey Creek in Donely County, Texas. The fossil was obtained by W. W. Dalquest, during June 1960. The principal cusps of the teeth are rounded, suggesting that it was mainly frugivorous. This species differs from C. crucidens as it lacks an accessory cusp between the protocone and paracone of the fourth upper premolar.

References

1. ^ Barbour, Erwin; Cook, Harold (1 January 1914). "TWO NEW FOSSIL DOGS OF THE GENUS CYNARCTUS FROM NEBRASKA". Conservation and Survey Division. 4.
2. Paleobiology Database: Cynartus
3. Wang, Xiaoming; Tedford, Richard H. (2008). Dogs, Their Fossil Relatives and Evolutionary History. Columbia. p. 34. ISBN 978-0-231-13528-3.
4. ^ McGrew, Paul O. (1937). "The Genus Cynarctus". Journal of Paleontology. 11 (5): 444–449. ISSN 0022-3360. JSTOR 1298460.
5. Hall, E. Raymond; Dalquest, Walter W. (1962). "A New Doglike Carnivore, Genus Cynarctus, Pliocene, of Texas". University of Kansas Publications Museum of Natural History. 14: 135–138.

• Wang, X., R.H. Tedford, and B.E. Taylor. 1999. Phylogenetic systematics of the Borophaginae (Carnivora, Canidae). Bulletin of the American Museum of Natural History, 243:1-391.

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