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Interatherium Temporal range: Early-Mid Miocene (Colhuehuapian-Mayoan) ~21.0–11.608 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Skeleton of Interatherium rodens in the Field Museum of Natural History, Chicago
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Notoungulata
Family:	†Interatheriidae
Subfamily:	†Interatheriinae
Genus:	†Interatherium Ameghino 1887
Species
I. anguliferum Ameghino 1894 I. brevifrons Ameghino 1894 I. dentatum Ameghino 1894 I. interruptum Ameghino 1894 I. robustum Ameghino 1887 I. rodens (Moreno 1882) I. supernum Ameghino 1894
Synonyms
Tembotherium Moreno 1882

Interatherium is an extinct genus of interatheriid notoungulate from the Early to Middle Miocene (Colhuehuapian-Mayoan). Fossils have been found in the Santa Cruz, Collón Curá and Sarmiento Formations in Argentina.

Description

This animal, of the size similar to that of today's American Mink (it was about 40 centimeters long excluding the tail), was equipped with a rather unusual morphology if related to that of its closest relatives. Contrary to shapes like Protypotherium, Interatherium possessed short and robust legs, and a long body and similar to that of a weasel. The front legs were equipped with four fingers. The ointment phalanges were compressed laterally, and some of these were divided.

The Interatherium skull was equally aberrant: it was in fact much more compact than that of other archaic types, and the muzzle was as if it had been crushed. The jaw, in particular, was very deep and tall, protruded backwards and owned considerable notches for the insertion of powerful masses masses (one of the main muscles capable of chewing in mammals). The Masseter muscle was anchored to a large bony flange placed in the lower part of the jaw, known as the descending process. This process is greatly reduced in related interatheres: Interatherium's well developed process was superficially similar to the ones seen in glyptodonts, a taxon of giant armadillos. Like the other interactors, Interatherium possessed a complete teeth, including 44 teeth, but differ in having a diastema between the incisors and premolars. The space was accentuated by the dimensions of the last incisive and the canine, so small as to be absent in some specimens.

Classification

Interatherium was described for the first time in 1887 by Florentino Ameghino, on the basis of fossil remains found in the province of Santa Cruz, Argentina, in strata dating to the lower Miocene. The type species is Interatherium rodens, which was named on the basis of a right maxilla that was first collected by Francisco P. Moreno in 1876-77. Several other species were described by Ameghino in 1894 (I. anguliferum, I. brevifrons, I. dentatum, I. interruptum, I. rodens, I. supernum, I. excavatus), many of which were then considered synonyms of I. rodens, though many of their holotypes have been lost.

Interatherium is the eponymous genus of the Interatheriidae family, a family that includes numerous medium-sized species from the Miocene of South America; Interatherium was most closely related to another Interathere from Santa Cruz, Cochilius.

Paleobiology

Due to the short legs and the elongated body, some paleontologists believe that Interatherium may have been an animal that lived in underground dens, but the legs do not show particular adaptations to digging. It was certainly a herbivorous animal that fed on leaves, grass and vegetation that grew at the ground level, perhaps close to bodies of water.

References

1. Interatherium at Fossilworks.org
2. ^ Fernández, M., Fernicola, J. C., & Cerdeño Serrano, M. E. (2019). On the type materials of the genera Interatherium Ameghino, 1887 and Icochilus Ameghino, 1889 (Interatheriidae, Notoungulata, Mammalia) from early Miocene of the Santa Cruz Province, Argentina.
3. ^ Fernández, M. (2015). Revisión taxonómica de Interatherium Ameghino 1887 e Icochilus Ameghino 1889 (Interatheriidae, Notoungulata) de la Edad Mamífero Santacrucense (Mioceno Temprano) de la Provincia de Santa Cruz, Argentina. Luján: Departamento de Ciencias Básicas, Universidad Nacional de Luján.
4. Ameghino, F. (1887). Enumeración sistemática de las especies de mamíferos fósiles coleccionados por Carlos Ameghino en los terrenos eocenos de la Patagonia austral y depositados en el Museo de La Plata. Boletín del Museo de la Plata.
5. Ameghino, F. (1894). Enumération synoptique des espèces de mammifères fossiles des formations éocènes de Patagonie. Imp. de PE Coni é hijos.
6. Kramarz, A. G., Vucetich, M. G., Carlini, A. A., Ciancio, M. R., Alejandra, M., Abello, C. M. D., & Gelfo, J. N. (2010). 18 A new mammal fauna at the top of the Gran Barranca sequence and its biochronological significance. The paleontology of Gran Barranca: evolution and environmental change through the middle Cenozoic of Patagonia, 264.

v t e Genera of South American native ungulates
Kingdom: Animalia Phylum: Chordata Class: Mammalia Superorder: Laurasiatheria
Xenungulata, Pyrotheria, Astrapotheria, and other minor groups †Kollpaniidae? Andinodus Molinodus Pascualodus? Pucanodus Simoclaenus Tiuclaenus †Didolodontidae Didolodus Enneoconus Ernestokokenia Escribania? Lamegoia Paulogervaisia Raulvaccia Ricardocifellia Saltaodus Umayodus Xesmodon †Protolipternidae Asmithwoodwardia Miguelsoria Protolipterna †Xenungulata †Carodniidae Carodnia Etayoa Notoetayoa Rodcania †Pyrotheria †Colombitheriidae Colombitherium Proticia †Pyrotheriidae Baguatherium Berracotherium Carolozittelia Griphodon Propyrotherium Pyrotherium †Astrapotheria Eoastrapostylops †Trigonostylopidae Shecenia Tetragonostylops Trigonostylops †Astrapotheriidae Antarctodon Astrapodon Comahuetherium Liarthrus Maddenia Parastrapotherium †Albertogaudryinae Albertogaudrya Astraponotus †Astrapotheriinae Astrapothericulus Astrapotherium Scaglia †Uruguaytheriinae Granastrapotherium Hilarcotherium Uruguaytherium Xenastrapotherium
Litopterna †Eulitopterna †Adianthidae Adianthus Proadiantus Proectocion? Proheptaconus Thadanius Tricoelodus †Macraucheniidae Polymorphis †Cramaucheniinae Caliphrium Coniopternium Cramauchenia Llullataruca Phoenixauchenia Pternoconius Theosodon †Macraucheniinae Cullinia Huayqueriana Macrauchenia Macraucheniopsis Micrauchenia Oxyodontherium Paranauchenia Promacrauchenia Scalabrinitherium Windhausenia Xenorhinotherium †Proterotherioidea †Anisolambdidae †Anisolambdinae Anisolambda Eolicaphrium Guilielmofloweria Lambdaconops Paranisolambda Polyacrodon Protheosodon Wainka †Sparnotheriodontinae Heteroglyphis Notiolofos Phoradiadius Sparnotheriodon Victorlemoinea †Proterotheriidae Mesolicaphrium Neodolodus Olisanophus Promylophis Prothoatherium Pseudobrachytherium Salladolodus †Megadolodinae Bounodus Megadolodus †Proterotheriinae Anisolophus Brachytherium Diadiaphorus Diplasiotherium Eoauchenia Epecuenia Epitherium Lambdaconus Neobrachytherium Neolicaphrium Paramacrauchenia Picturotherium Prolicaphrium Proterotherium Tetramerorhinus Thoatheriopsis Thoatherium Uruguayodon Villarroelia †Notopterna †Amilnedwardsiidae Amilnedwardsia Ernestohaeckelia Rutimeyeria †Indaleciidae Adiantoides Indalecia †Notonychopidae Notonychops Requisia Incertae sedis Kyraodus Proacrodon
Notoungulata incertae sedis Acoelodus Archaeogaia Carolodarwinia Perutherium †Notioprogonia Satshatemnus Seudenius †Henricosborniidae Henricosbornia Nanolophodon Othnielmarshia Peripantostylops Simpsonotus †Notostylopidae Boreastylops Chilestylops Edvardotrouessartia Homalostylops Notostylops Otrhonia Parastylops †Toxodontia †Homalodotheriidae Asmodeus Chasicotherium Homalodotherium Trigonolophodon †Isotemnidae Anisotemnus Coelostylodon Distylophorus Hedralophus Isotemnus Lophocoelus Pampatemnus Periphragnis Pleurocoelodon Pleurostylodon Rhyphodon Thomashuxleya †Eutoxodontia †Leontiniidae Anayatherium Ancylocoelus Colpodon Coquenia Elmerriggsia Gualta Henricofilholia Huilatherium Leontinia Martinmiguelia Purperia? Scarrittia Taubatherium Termastherium †Notohippidae Acoelohyrax Argyrohippus Deuterotherium? Edvardocopeia Eomorphippus Eurygenium Mendozahippus Moqueguahippus Morphippus Nesohippus Notohippus Pampahippus Pascualihippus Patagonhippus Perhippidion Plexotemnus Puelia Purperia? Rhynchippus Rosendo Stilhippus Teushentherium Trimerostephanos †Toxodontidae †Nesodontinae Adinotherium Nesodon Proadinotherium †Toxodontinae Abothrodon Andinotoxodon Calchaquitherium Ceratoxodon Charruatoxodon Dinotoxodon Eutomodus Falcontoxodon Gyrinodon Haplodontherium Hemixotodon Hoffstetterius Hyperoxotodon Mesenodon Mesotoxodon Minitoxodon Mixotoxodon Neoadinotherium Neotoxodon Nesodonopsis Nonotherium Ocnerotherium Palyeidodon Paratrigodon Pericotoxodon Piauhytherium Pisanodon Plesiotoxodon Posnanskytherium Prototrigodon Stenotephanos Stereotoxodon Toxodon Toxodontherium Trigodon Trigodonops Xotodon †Typotheria Campanorco Griphotherion †Archaeopithecidae Archaeopithecus Teratopithecus †Oldfieldthomasiidae Allalmeia Brachystephanus Camargomendesia Colbertia Dolichostylodon Itaboraitherium Kibenikhoria Maxschlosseria Oldfieldthomasia Paginula Suniodon Tsamnichoria Ultrapithecus Xenostephanus †Interatheriidae Antofagastia Ignigena Johnbell Munizia Punapithecus †Interatheriinae Archaeophylus Argyrohyrax Boleatherium Brucemacfaddenia Caenophilus Choichephilum Cochilius Eopachyrucos Federicoanaya Interatherium Juchuysillu Miocochilius Neoicochilus Patriarchus Proargyrohyrax Progaleopithecus Protypotherium Santiagorothia †Notopithecinae Antepithecus Guilielmoscottia Notopithecus Transpithecus †Mesotheriidae Fiandraia? Trachytherus †Mesotheriinae Altitypotherium Caraguatypotherium Eotypotherium Eutypotherium Hypsitherium Mesotherium Microtypotherium Plesiotypotherium Pseudotypotherium Rusconitherium Typotheriopsis †Archaeohyracidae Archaeohyrax Archaeotypotherium Bryanpattersonia Eohyrax Pascualhyrax Protarchaeohyrax Pseudhyrax Punahyrax †Hegetotheriidae †Hegetotheriinae Ethegotherium Hegetotheriopsis Hegetotherium Hemihegetotherium Prohegetotherium Pseudohegetotherium Sallatherium †Pachyrukhinae Medistylus Propachyrucos Prosotherium Tremacyllus †Pachyrukhini Pachyrukhos Paedotherium Tremacyllus

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