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Macraucheniidae Temporal range: Oligocene-Early Holocene (Deseadan-Lujanian) ~30–0.010 Ma PreꞒ Ꞓ O S D C P T J K Pg N (possible Late Eocene records)
Skeleton of Macrauchenia patachonica
Skeleton of Thesodon
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Litopterna
Superfamily:	†Macrauchenioidea
Family:	†Macraucheniidae Gill, 1872
Subfamilies and genera
†Amilnedwardsia †Llullataruca †Polymorphis †Cramaucheniinae †Macraucheniinae †Cullinia †Huayqueriana †Macrauchenia †Macraucheniopsis †Micrauchenia †Oxyodontherium †Paranauchenia †Promacrauchenia †Scalabrinitherium †Windhausenia †Xenorhinotherium

Macraucheniidae is a family in the extinct South American ungulate order Litopterna, that resembled camelids. They had three functional digits on the fore and hind feet, as well as elongate necks. The family is generally divided up into two subfamilies, Cramaucheniinae (which may be paraphyletic) and Macraucheniinae. The family shows retraction of the nasal region, most extremely to the top of the skull in derived macraucheniine taxa like Macrauchenia. which has been interpreted to have supported a probsocis, perhaps like that of a saiga antelope to filter dust, or a moose-like prehensile lip. The earliest unambiguous members of the family date to the late Oligocene around 30 million years ago. Polymorphis from the Eocene has historically been placed as a macraucheniid, but this has been doubted. Most early representatives had a body masses in the range of 80–120 kilograms (180–260 lb), though some like Llullataruca were as small as 35–55 kilograms (77–121 lb), and the last representatives of the family from the Pleistocene like Macrauchenia were over 1,000 kilograms (2,200 lb). The family reached its apex of diversity during the late Miocene around 10-6 million years ago, before declining to only a few species belong to the genera Macrauchenia and Xenorhinotherium by the Late Pleistocene.

Below is a phylogenetic tree of the Macraucheniidae, after McGrath et al. 2018:.

Cladogram of Macraucheniidae after Lobo, Gelfo & Azevedo (2024):

References

1. Püschel, Hans P.; Alarcón-Muñoz, Jhonatan; Soto-Acuña, Sergio; Ugalde, Raúl; Shelley, Sarah L.; Brusatte, Stephen L. (2023-02-25). "Anatomy and phylogeny of a new small macraucheniid (Mammalia: Litopterna) from the Bahía Inglesa Formation (late Miocene), Atacama Region, Northern Chile". Journal of Mammalian Evolution. 30 (2): 415–460. doi:10.1007/s10914-022-09646-0. ISSN 1573-7055.
2. ^ Lobo, Leonardo Souza; Gelfo, Javier N.; de Azevedo, Sergio A. K. (2024-12-31). "The phylogeny of Macraucheniidae (Mammalia, Panperissodactyla, Litopterna) at the genus level". Journal of Systematic Palaeontology. 22 (1). doi:10.1080/14772019.2024.2364201. ISSN 1477-2019.
3. Blanco, R. Ernesto; Jones, Washington W.; Yorio, Lara; Rinderknecht, Andrés (October 2021). "Macrauchenia patachonica Owen, 1838: Limb bones morphology, locomotory biomechanics, and paleobiological inferences". Geobios. 68: 61–70. Bibcode:2021Geobi..68...61B. doi:10.1016/j.geobios.2021.04.006.
4. Croft, Darin A.; Gelfo, Javier N.; López, Guillermo M. (2020-05-30). "Splendid Innovation: The Extinct South American Native Ungulates". Annual Review of Earth and Planetary Sciences. 48 (1): 259–290. Bibcode:2020AREPS..48..259C. doi:10.1146/annurev-earth-072619-060126. ISSN 0084-6597. S2CID 213737574.
5. Andrew J. McGrath; Federico Anaya; Darin A. Croft (2018). "Two new macraucheniids (Mammalia: Litopterna) from the late middle Miocene (Laventan South American Land Mammal Age) of Quebrada Honda, Bolivia". Journal of Vertebrate Paleontology. 38 (3): e1461632. Bibcode:2018JVPal..38E1632M. doi:10.1080/02724634.2018.1461632. S2CID 89881990.

Bibliography

• Forasiepi, Analía M.; MacPhee, Ross D.E.; Hernández del Pino, Santiago; Schmidt, Gabriela I.; Amson, Eli; Grohé, Camille (2016). "Exceptional skull of Huayqueriana (Mammalia, Litopterna, Macraucheniidae) from the Late Miocene of Argentina: anatomy, systematics and paleobiological implications" (PDF). Bulletin of the American Museum of Natural History. 404: 1–76. doi:10.1206/0003-0090-404.1.1. S2CID 89219979. Retrieved 2018-10-01.

External links

• Phylogeny of Macraucheniidae

v t e Genera of South American native ungulates
Kingdom: Animalia Phylum: Chordata Class: Mammalia Superorder: Laurasiatheria
Xenungulata, Pyrotheria, Astrapotheria, and other minor groups †Kollpaniidae? Andinodus Molinodus Pascualodus? Pucanodus Simoclaenus Tiuclaenus †Didolodontidae Didolodus Enneoconus Ernestokokenia Escribania? Lamegoia Paulogervaisia Raulvaccia Ricardocifellia Saltaodus Umayodus Xesmodon †Protolipternidae Asmithwoodwardia Miguelsoria Protolipterna †Xenungulata †Carodniidae Carodnia Etayoa Notoetayoa Rodcania †Pyrotheria †Colombitheriidae Colombitherium Proticia †Pyrotheriidae Baguatherium Berracotherium Carolozittelia Griphodon Propyrotherium Pyrotherium †Astrapotheria Eoastrapostylops †Trigonostylopidae Shecenia Tetragonostylops Trigonostylops †Astrapotheriidae Antarctodon Astrapodon Comahuetherium Liarthrus Maddenia Parastrapotherium †Albertogaudryinae Albertogaudrya Astraponotus †Astrapotheriinae Astrapothericulus Astrapotherium Scaglia †Uruguaytheriinae Granastrapotherium Hilarcotherium Uruguaytherium Xenastrapotherium
Litopterna †Eulitopterna †Adianthidae Adianthus Proadiantus Proectocion? Proheptaconus Thadanius Tricoelodus †Macraucheniidae Polymorphis †Cramaucheniinae Caliphrium Coniopternium Cramauchenia Llullataruca Phoenixauchenia Pternoconius Theosodon †Macraucheniinae Cullinia Huayqueriana Macrauchenia Macraucheniopsis Micrauchenia Oxyodontherium Paranauchenia Promacrauchenia Scalabrinitherium Windhausenia Xenorhinotherium †Proterotherioidea †Anisolambdidae †Anisolambdinae Anisolambda Eolicaphrium Guilielmofloweria Lambdaconops Paranisolambda Polyacrodon Protheosodon Wainka †Sparnotheriodontinae Heteroglyphis Notiolofos Phoradiadius Sparnotheriodon Victorlemoinea †Proterotheriidae Mesolicaphrium Neodolodus Olisanophus Promylophis Prothoatherium Pseudobrachytherium Salladolodus †Megadolodinae Bounodus Megadolodus †Proterotheriinae Anisolophus Brachytherium Diadiaphorus Diplasiotherium Eoauchenia Epecuenia Epitherium Lambdaconus Neobrachytherium Neolicaphrium Paramacrauchenia Picturotherium Prolicaphrium Proterotherium Tetramerorhinus Thoatheriopsis Thoatherium Uruguayodon Villarroelia †Notopterna †Amilnedwardsiidae Amilnedwardsia Ernestohaeckelia Rutimeyeria †Indaleciidae Adiantoides Indalecia †Notonychopidae Notonychops Requisia Incertae sedis Kyraodus Proacrodon
Notoungulata incertae sedis Acoelodus Archaeogaia Carolodarwinia Perutherium †Notioprogonia Satshatemnus Seudenius †Henricosborniidae Henricosbornia Nanolophodon Othnielmarshia Peripantostylops Simpsonotus †Notostylopidae Boreastylops Chilestylops Edvardotrouessartia Homalostylops Notostylops Otrhonia Parastylops †Toxodontia †Homalodotheriidae Asmodeus Chasicotherium Homalodotherium Trigonolophodon †Isotemnidae Anisotemnus Coelostylodon Distylophorus Hedralophus Isotemnus Lophocoelus Pampatemnus Periphragnis Pleurocoelodon Pleurostylodon Rhyphodon Thomashuxleya †Eutoxodontia †Leontiniidae Anayatherium Ancylocoelus Colpodon Coquenia Elmerriggsia Gualta Henricofilholia Huilatherium Leontinia Martinmiguelia Purperia? Scarrittia Taubatherium Termastherium †Notohippidae Acoelohyrax Argyrohippus Deuterotherium? Edvardocopeia Eomorphippus Eurygenium Mendozahippus Moqueguahippus Morphippus Nesohippus Notohippus Pampahippus Pascualihippus Patagonhippus Perhippidion Plexotemnus Puelia Purperia? Rhynchippus Rosendo Stilhippus Teushentherium Trimerostephanos †Toxodontidae †Nesodontinae Adinotherium Nesodon Proadinotherium †Toxodontinae Abothrodon Andinotoxodon Calchaquitherium Ceratoxodon Charruatoxodon Dinotoxodon Eutomodus Falcontoxodon Gyrinodon Haplodontherium Hemixotodon Hoffstetterius Hyperoxotodon Mesenodon Mesotoxodon Minitoxodon Mixotoxodon Neoadinotherium Neotoxodon Nesodonopsis Nonotherium Ocnerotherium Palyeidodon Paratrigodon Pericotoxodon Piauhytherium Pisanodon Plesiotoxodon Posnanskytherium Prototrigodon Stenotephanos Stereotoxodon Toxodon Toxodontherium Trigodon Trigodonops Xotodon †Typotheria Campanorco Griphotherion †Archaeopithecidae Archaeopithecus Teratopithecus †Oldfieldthomasiidae Allalmeia Brachystephanus Camargomendesia Colbertia Dolichostylodon Itaboraitherium Kibenikhoria Maxschlosseria Oldfieldthomasia Paginula Suniodon Tsamnichoria Ultrapithecus Xenostephanus †Interatheriidae Antofagastia Ignigena Johnbell Munizia Punapithecus †Interatheriinae Archaeophylus Argyrohyrax Boleatherium Brucemacfaddenia Caenophilus Choichephilum Cochilius Eopachyrucos Federicoanaya Interatherium Juchuysillu Miocochilius Neoicochilus Patriarchus Proargyrohyrax Progaleopithecus Protypotherium Santiagorothia †Notopithecinae Antepithecus Guilielmoscottia Notopithecus Transpithecus †Mesotheriidae Fiandraia? Trachytherus †Mesotheriinae Altitypotherium Caraguatypotherium Eotypotherium Eutypotherium Hypsitherium Mesotherium Microtypotherium Plesiotypotherium Pseudotypotherium Rusconitherium Typotheriopsis †Archaeohyracidae Archaeohyrax Archaeotypotherium Bryanpattersonia Eohyrax Pascualhyrax Protarchaeohyrax Pseudhyrax Punahyrax †Hegetotheriidae †Hegetotheriinae Ethegotherium Hegetotheriopsis Hegetotherium Hemihegetotherium Prohegetotherium Pseudohegetotherium Sallatherium †Pachyrukhinae Medistylus Propachyrucos Prosotherium Tremacyllus †Pachyrukhini Pachyrukhos Paedotherium Tremacyllus

• Macraucheniids
• Litopterns
• Prehistoric mammal families
• Quaternary mammals of South America
• Neogene mammals of South America
• Paleogene mammals of South America
• Ypresian first appearances
• Holocene extinctions